taxonID	type	description	language	source
CF0DD0F6EE6F5C12969321DF5523A284.taxon	description	Fig. 1 A ‒ D	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
CF0DD0F6EE6F5C12969321DF5523A284.taxon	description	Description. Male. Head: Eye bridge four facets wide. Prefrons with many setae and clypeus bare. Scape brown with 3 anterior setae, pedicel brown with many setae; flagellomeres brown; 4 th flagellomere 2.6 times as long as wide, neck portion about 1 / 8 of its whole length (Fig. 1 A), hairs yellow and almost as long as its width. Palpus yellowish brown, 2 - segmented, 1 st segment with 4 setae on its outside and with distinct brownish sensory area bearing minute sensilla, 2 nd segment with 6 setae (Fig. 1 B). Thorax: Color predominantly brown with dorsocentral and dorsolateral setae on mesonotum; scutellum with many setae; posterior pronotum with setae. Legs predominantly yellowish brown; fore tibial spur about 1.2 times as long as width of tibial apex. Wing membrane bare; distal 2 / 3 of R 5 with both dorsal and ventral macrotrichia, bM bare, r-m with several dorsal macrotrichia, M 1, M 2, and CuA 1 with dorsal macrotrichia, stem of M bare; length of R 1 = 1.3 R, c = 0.8 w, r-m = 1.6 bM; stem of CuA very short, almost absent; R 1 ending at same level of branching point of M 1 and M 2; wing length 4.1 mm. Halter brown, knob and distal half of stalk with setae. Abdomen: Tergites and sternites brown with dark brownish setae. Gonocoxite slightly wider than long (Fig. 1 C); gonostylus nearly semicircular in shape with 3 apical spines (Fig. 1 D); tegmen trapezoidal with darkened border (Fig. 1 C); aedeagus distinct (Fig. 1 C). Body size: 3.4 mm. Female. Unknown.	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
2740551B27015012A1B804FC5C1605A6.taxon	description	Fig. 2 A, B	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
7BA809D661C55503A72F2F240F3D3E51.taxon	description	Fig. 6 A ‒ D	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
7BA809D661C55503A72F2F240F3D3E51.taxon	description	Description. Male. Head: Eye bridge 3 ‒ 4 facets wide. Prefrons with setae and clypeus bare. Scape brown with about 5 setae anteriorly, pedicel brown with about 8 setae; flagellomeres dark brown except for yellowish brown base of 1 st flagellomere; 4 th flagellomere about 3.0 ‒ 3.5 times as long as wide, neck portion about 1 / 10 of its whole length (Fig. 6 A), hairs yellowish brown and almost as long as width of flagellomere. Palpus brown, 3 - segmented with length ratio 1.5: 1.0: 1.6, 1 st segment with 5 ‒ 7 setae mainly on its outside and without distinct sensory area bearing minute sensilla, 2 nd segment with 6 ‒ 8 setae, 3 rd segment slender with 5 ‒ 8 setae (Fig. 6 B). Thorax: Color predominantly dark brown; dorsocentral setae on mesonotum short, dorsolateral setae on mesonotum and setae on scutellum consist of longer and shorter ones; posterior pronotum with setae. Legs brown; tibiae and basitarsi of all legs with differentiated spine-like short setae in ground setosity; fore tibial spur about 1.1 ‒ 1.2 times as long as width of tibial apex. Wing with brownish anterior veins and light brownish posterior veins; wing membrane bare; R 5 with both dorsal and ventral macrotrichia, bM bare, r-m, M 1, M 2, stem of M, CuA 1, and CuA 2 with dorsal macrotrichia; length of R 1 = 0.7 ‒ 0.8 R, c = 0.7 w, r-m = 1.0 ‒ 1.7 bM, stem of CuA very short, almost absent; R 1 ending at same level of or slightly beyond branching point of M 1 and M 2; wing length 2.9 ‒ 3.5 mm. Halter brown, knob with setae. Abdomen: Tergites and sternites brown with dark brownish setae, membrane between tergites and sternites yellowish brown. Gonocoxite wider than long with narrow base (Fig. 6 C); gonostylus longer than wide, its inside moderately swollen on which many spines are present (Fig. 6 D); upper half of tegmen rounded with two pocket-like structures (Fig. 6 C); aedeagus nearly Y-shaped (Fig. 6 C). Body size: 3.6 ‒ 4.4 mm. Female. Similar to male, but hairs of 4 th flagellomere shorter than its width. Fore tibial spur about 1.3 ‒ 1.4 times as long as width of tibial apex. Wing length 3.0 ‒ 3.8 mm. Body size 4.5 ‒ 5.2 mm.	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
4B14AFA610D1540993F36250C658DC90.taxon	description	Fig. 9 A ‒ C	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
4B14AFA610D1540993F36250C658DC90.taxon	description	Description. Male. Head: Eye bridge 3 ‒ 4 facets wide. Prefrons with setae and clypeus bare. Scape brown with a seta, pedicel brown with about 12 setae; flagellomeres brown; 4 th flagellomere 2.1 times as long as wide, neck portion about 1 / 9 of its whole length (Fig. 9 A), hairs yellowish brown and shorter than width of flagellomere. Palpus yellowish brown and 3 - segmented; 1 st segment with 3 ‒ 4 setae mainly on its outside and without distinct brownish sensory area, 2 nd segment with about 10 setae, 3 rd segment slender with 10 ‒ 12 setae. Length ratio of segments of palpus is unclear because it is preserved obliquely. Thorax: Color predominantly dark brown; setae on mesonotum and scutellum mostly shed off in the holotype; posterior pronotum with setae. Coxae, femora, and tibiae predominantly yellowish brown, tarsi brown; tibiae and basitarsi with differentiated spine-like short setae in ground setosity; fore tibial spur about 1.3 times as long as width of tibial apex. Wing with brownish anterior veins and light brownish posterior veins; wing membrane without macrotrichia; R 5 with both dorsal and ventral macrotrichia, bM bare, r-m with 8 dorsal macrotrichia, M 1, M 2, CuA 1, and CuA 2 with dorsal macrotrichia, stem of M with about 20 dorsal macrotrichia; length of R 1 nearly equal to that of R, c = 0.7 w, r-m = 1.5 bM, stem of CuA very short, almost absent; R 1 ending slightly beyond level of branching point of M 1 and M 2; wing length 3.1 mm. Halter yellowish brown, knob with setae. Abdomen: Tergites and sternites brown with dark brownish setae, membrane between tergites and sternites yellowish brown. Gonocoxite wider than long (Fig. 9 B); gonostylus nearly ovoid in shape with 5 spines and with dense setae above uppermost spine (Fig. 9 C), two lowest spines overlap at same level and look like one spine from ventral view; tegmen nearly triangular with minute denticles on lower part (Fig. 9 B); aedeagus Y-shaped with dark brownish base (Fig. 9 B). Body size: 3.6 mm. Female. Unknown.	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
15D2FCA4F48859E297F289BD0E1C1E24.taxon	description	Fig. 7 A ‒ E	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
15D2FCA4F48859E297F289BD0E1C1E24.taxon	description	Description. Male. See Rudzinski (2005) for a morphological description of the male of this species. Wing length and body size of the specimens examined in this study are 2.5 ‒ 3.1 mm and 2.6 ‒ 3.1 mm respectively. Female. Similar to male, but flagellum of antenna about 0.8 times shorter than that of male. Wing length 2.8 ‒ 3.2 mm. Body size 3.6 ‒ 4.0 mm. Final-instar larva. Head (Fig. 7 D, E): Well-chitinized, dark brown. Frontal plate (fp) with 4 pairs of sensory pits (p 1 ‒ p 4), of these, p 3 and p 4 with very short, minute sensory hair. Gena (ge) with 5 pairs of dorsal sensory pits along marginal suture (p 5 ‒ p 9). Besides them, each gena with 3 dorsolateral, 1 ventrolateral, and 1 ventral sensory pits, of these, anterior dorsolaeral pit with minute sensory hair like p 3 and p 4. Anterior part of head dorsally with labrum (lb), clypeus (cl), and antennal plate (ap), laterally with mandible (md), and ventrally with maxilla and hypopharynx (hp). Maxilla is subdivided into three sclerites, galeolacinia (gl), stipes (st), and cardo (cd); galeolacinia with serrated margin, fused basally with stipes. Thorax and abdomen (Fig. 7 C): Body comprises 3 thoracic and 9 abdominal segments. Creamy white in color, but yellowish when alive. Surface smooth without setae, locomotory pad slightly developed ventrally on abdominal segments. Prothoracic and 1 st to 7 th abdominal segments with spiracles; diameter of prothoracic spiracles larger than that of abdominal ones. Body size: 5.5 ‒ 9.0 mm. Pupa. Similar to pupa of Trichosia pilosa (currently known as Leptosciarella pilosa) figured in Plachter (1979) in having a remarkable process on vertex. The pupa of T. caudata also has this characteristic process on vertex (Tuomikoski 1957). Body size: 3.4 ‒ 4.1 mm.	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
D18EE45DCBE9520C9274BD633C7DB951.taxon	description	Figs 4 A, B, 5 A ‒ F	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
D18EE45DCBE9520C9274BD633C7DB951.taxon	description	Description. Male. Head: Eye bridge 3 ‒ 4 facets wide. Prefrons with setae and clypeus bare. Scape and pedicel brown each with about 6 ‒ 10 setae; flagellomeres brown except for yellowish base of 1 st flagellomere; 4 th flagellomere about 3.5 ‒ 4.0 times as long as wide, neck portion about 1 / 9 of its whole length (Fig. 5 A), hairs yellowish brown and slightly longer than width of flagellomere. Palpus yellowish brown, 3 - segmented with length ratio 1.3: 1.0: 1.5, 1 st segment with 2 ‒ 5 setae mainly on its outside and without distinct sensory area bearing minute sensilla, 2 nd segment with 4 ‒ 7 setae, 3 rd segment slender with 4 ‒ 8 setae (Fig. 5 B). Thorax: Color predominantly dark brown (Fig. 4 A); setae on mesonotum and scutellum consist of longer and shorter ones; posterior pronotum bare. Coxae, femora, and tibiae predominantly yellowish brown, tarsi brown (Fig. 4 A); middle tibia, hind tibia, and basitarsi of all legs with differentiated spine-like short setae in ground setosity; fore tibial spur about 1.1 ‒ 1.2 times as long as width of tibial apex (Fig. 5 E). Wing with brownish anterior veins and light brownish posterior veins (Fig. 4 B); wing membrane fumous, with numerous macrotrichia and minute pits in which base of macrotrichia are embedded (Figs 4 B, 5 F); R 5 with both dorsal and ventral macrotrichia almost along its whole length, bM bare, r-m, M 1, M 2, stem of M, CuA 1, and CuA 2 with dorsal macrotrichia; length of R 1 = 0.9 ‒ 1.0 R, c = 0.7 ‒ 0.8 w, r-m = 0.9 ‒ 1.8 bM (x), stem of CuA very short, almost absent; R 1 ending slightly beyond branching point of M 1 and M 2; wing length 3.4 ‒ 3.7 mm. Halter brown, knob and distal 1 / 4 ‒ 1 / 3 of stalk with setae. Abdomen: Tergites and sternites dark brown with dark brownish setae, abdominal segments slender (Fig. 4 A). Gonocoxite wider than long (Fig. 5 C); gonostylus longer than wide, its inside with a wide lobe on which some spines are present (Fig. 5 D); upper half of tegmen nearly quadrate with two pocket-like structures (Fig. 5 C), upper margin of tegmen fringed with fine teeth; aedeagus nearly Y-shaped (Fig. 5 C). Body size: 3.5 ‒ 3.8 mm. Female. Similar to male, but 4 th flagellomere about 3.0 times as long as wide, hairs almost as long as width of flagellomere. Wing length 3.5 ‒ 3.8 mm. Body size 3.6 ‒ 4.8 mm.	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
823EA8B21A505719A07E79ABB4487CC2.taxon	description	Fig. 10 A ‒ D	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
823EA8B21A505719A07E79ABB4487CC2.taxon	description	Description. Male. Head: Eye bridge 4 ‒ 5 facets wide. Prefrons with setae and clypeus bare. Scape brown with 3 setae, pedicel brown with about 8 setae; flagellomeres brown; 4 th flagellomere 3.5 times as long as wide, neck portion about 1 / 13 of its whole length (Fig. 10 A), hairs yellowish brown and slightly shorter than width of flagellomere. Palpus brown, 3 - segmented with length ratio 1.5: 1.0: 2.3, 1 st segment with 7 setae and with indistinct sensory area bearing minute sensilla, 2 nd segment with 9 setae, 3 rd segment slender with 8 setae (Fig. 10 B). Thorax: Color predominantly dark brown; setae on mesonotum and scutellum consist of longer and shorter ones, scutellum with 5 long setae; posterior pronotum with setae. Legs predominantly brown; tibiae and basitarsi with differentiated spine-like short setae in ground setosity; fore tibial spur about 1.3 times as long as width of tibial apex. Wing with brownish anterior veins and light brownish posterior veins; wing membrane without macrotrichia but a macrotrichium is seen between M 1 and M 2 of left wing; R 5 and M 1 with both dorsal and ventral macrotrichia, bM with 3 dorsal macrotrichia, r-m with 7 dorsal macrotrichia, M 2, CuA 1, and CuA 2 with dorsal macrotrichia, stem of M with 14 ‒ 15 dorsal macrotrichia; length of R 1 = 1.2 R, c = 0.7 w, r-m = 1.0 bM, stem of CuA very short, almost absent; R 1 ending slightly beyond level of branching point of M 1 and M 2; wing length 4.0 mm. Halter brown, knob and distal 1 / 3 of stalk with setae. Abdomen: Tergites and sternites dark brown with dense dark brownish setae. Gonocoxite wider than long with U-shaped cleft in base (Fig. 10 C); gonostylus pyriform, left gonostylus with 6 spines and right gonostylus with 7 spines inside, 2 spines directed downward compared to the other ones (Fig. 10 D); tegmen nearly triangular but with arcuate base, and with rounded membranous appendage on upper portion (Fig. 10 C); aedeagus located beneath tegmen (Fig. 10 C). Body size: 3.9 mm. Female. Unknown.	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
0A0F1F35E184527596F7513DE0F5AF59.taxon	description	Fig. 8 A ‒ D	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
0A0F1F35E184527596F7513DE0F5AF59.taxon	description	Description. Male. Head: Eye bridge 4 facets wide. Prefrons with setae, clypeus bare. Scape brown with 2 ‒ 5 setae, pedicel brown with about 9 setae; flagellomeres brown; 4 th flagellomere about 2.5 times as long as wide, neck portion about 1 / 6 of its whole length (Fig. 8 A), hairs yellowish brown and slightly shorter than width of flagellomere. Palpus brown, 2 - segmented (holotype) or 3 - segmented (paratypes), 1 st segment with 1 ‒ 4 setae and with indistinct sensory area bearing minute sensilla, 2 nd and 3 rd segments slender with 3 ‒ 6 setae respectively (Fig. 8 B). Thorax: Color predominantly dark brown; mesonotum with both long and short setae; scutellum with some setae, at most 4 of them distinctly longer than the others; posterior pronotum with setae. Legs brown; tibiae and basitarsi with differentiated spine-like short setae in ground setosity; fore tibial spur about 1.3 times as long as width of tibial apex. Wing with brownish anterior veins and light brownish posterior veins; wing membrane bare; distal part of R 5 with both dorsal and ventral macrotrichia, bM and stem of M bare, r-m, M 1, M 2, CuA 1, and CuA 2 with dorsal macrotrichia; length of R 1 = 0.5 ‒ 0.7 R, c = 0.7 w, r-m = 1.4 ‒ 1.6 bM but in a paratype from Tochigi r-m = 0.6 bM, stem of CuA very short, almost absent; R 1 ending slightly before level of branching point of M 1 and M 2; wing length 2.2 ‒ 2.7 mm. Halter brown, knob with setae. Abdomen: Tergites and sternites dark brown with dark brownish setae, membrane between tergites and sternites brown. Gonocoxite wider than long with broad base (Fig. 8 C); gonostylus nearly ovoid in shape with 7 ‒ 9 spines on about upper 2 / 5 of inside (Fig. 8 D); tegmen membranous with sclerotized border, wider than long, minute denticles located on central portion (Fig. 8 C); aedeagus distinct and large (Fig. 8 C). Body size: 2.3 ‒ 2.8 mm. Female. Same as male except for the following characters: Palpus 3 - segmented, but one paratype from Miyake Island (PCMS) asymmetrically with 3 - segmented right palpus and 2 - segmented left palpus; 1 st segment of palpus bare or with a seta. Wing vein r-m = 1.4 ‒ 1.8 bM; wing length 2.3 ‒ 2.5 mm. Body size 2.5 ‒ 3.5 mm.	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
66157C737777572EBB5740FEE8766429.taxon	description	Fig. 3 A ‒ D	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
66157C737777572EBB5740FEE8766429.taxon	description	Description. Male. Head: Eye bridge 2 ‒ 3 facets wide. Prefrons with about 5 setae and clypeus bare or with a seta. Scape brown with about 5 setae anteriorly, pedicel brown with about 10 setae; flagellomeres brown, base of 1 st flagellomere yellow; 4 th flagellomere about 5.0 times as long as wide, neck portion about 1 / 8 of its whole length (Fig. 3 A), hairs yellowish brown and longer than width of flagellomere. Palpus yellowish brown, 3 - segmented with length ratio 1.8: 1.0: 2.1, 1 st segment with 1 ‒ 3 (mostly 2) setae on its outside and with indistinct sensory area bearing minute sensilla, 2 nd segment with 6 ‒ 7 setae, frequently one of them distinctly longer than the others, 3 rd segment slender with about 10 setae (Fig. 3 B). Thorax: Color predominantly brown; mesonotum with both short and long setae; scutellum with at most 4 long setae and some short setae; posterior pronotum bare. Coxae, femora, and tibiae predominantly yellowish brown, tarsi brown; tibiae and basitarsi, especially hindtibia with differentiated spine-like short setae in ground setosity; fore tibial spur about 1.2 times as long as width of tibial apex. Wing with brownish anterior veins and light brownish posterior veins; wing membrane bare; distal 1 / 5 ‒ 1 / 4 of R 5 with both dorsal and ventral macrotrichia, bM, r-m, M 1, M 2, CuA 1, and CuA 2 with dorsal macrotrichia, stem of M bare or with several dorsal macrotrichia; length of R 1 = 0.6 ‒ 0.7 R, c = 0.7 ‒ 0.8 w, r-m = 0.5 ‒ 1.0 bM, stem of CuA = 0.3 ‒ 0.7 bM; R 1 ending slightly before branching point of M 1 and M 2; wing length 1.8 ‒ 2.4 mm. Halter yellowish brown, knob and distal 1 / 4 ‒ 1 / 3 of stalk with setae. Abdomen: Tergites and sternites brown with brownish setae, membrane between tergites and sternites yellowish brown. Gonocoxite wider than long (Fig. 3 C); gonostylus longer than wide with apical spines, and inside of dorsal surface with arcuate concavity where 3 ‒ 5 spines are present, 2 of them arranged parallel (Fig. 3 D); tegmen membranous, wider than long with weakly sclerotized base (Fig. 3 C); aedeagus nearly V-shaped (Fig. 3 C). Body size: 1.9 ‒ 2.9 mm. Female. Unknown.	en	Sutou, Mitsuaki, Menzel, Frank (2025): The genera Mouffetina Frey and Trichosia Winnertz (Diptera, Sciaridae) in Japan with a key for the Japanese species and an updated world checklist. Contributions to Entomology 75 (1): 167-182, DOI: 10.3897/contrib.entomol.75.e145605
