identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
BC05165FF3C45F5A87646872BED82206.text	BC05165FF3C45F5A87646872BED82206.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Veraphis Casey 1897	<div><p>Genus Veraphis Casey, 1897</p><p>Veraphis Casey, 1897: 509. Type species: Eutheia impressa LeConte, 1879 (designated by Franz in Newton and Franz 1998).</p><p>Diagnosis.</p><p>This genus is distinguished from other genera of Scydmaeninae by the following combination characteristics: body flattened and elongated (Figs 1 A, 2 A); procoxal cavities broadly open (Fig. 1 E); prosternum with a narrow intercoxal carina (Fig. 1 E); each elytron with a setose basal fovea (Figs 1 B, 2 B). For more detailed diagnostic characters and phylogenetic information on this genus, refer to Jałoszyński (2014 b).</p><p>Distribution.</p><p>Northern Hemisphere (China, Japan, Korea, USA, Finland, Sweden, Mongolia, Russia (Far East, East Siberia)).</p><p>Remarks.</p><p>According to Jałoszyński 2024, this genus is primarily found in leaf litter and soil, and all species discovered in China inhabit alpine regions at altitudes above 2000 m. Additionally, species found in Japan are also thought to be distributed in cooler climates. These species are very similar in external morphology, making examination of the aedeagus crucial for species identification. In some species, modifications of the male legs are present, which can be useful for distinguishing species.</p></div>	https://treatment.plazi.org/id/BC05165FF3C45F5A87646872BED82206	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Byeon, Ui-Joung;Park, Jong-Seok	Byeon, Ui-Joung, Park, Jong-Seok (2025): Two new species of the genus Veraphis Casey (Coleoptera, Staphylinidae, Scydmaeninae) from Korea. ZooKeys 1224: 165-174, DOI: 10.3897/zookeys.1224.142859
34D920B8BC1E57EB8786C07B8F6379D4.text	34D920B8BC1E57EB8786C07B8F6379D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Veraphis myeonggiensis Byeon & Park 2025	<div><p>Veraphis myeonggiensis sp. nov.</p><p>Figs 2, 3E – H, 4</p><p>Type material designated.</p><p>Holotype • ♂ NIBR: “ Korea: Gyeonggi Prov. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.4052&amp;materialsCitation.latitude=37.970055" title="Search Plazi for locations around (long 127.4052/lat 37.970055)">Mt. Myeonggi</a>, 520, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.4052&amp;materialsCitation.latitude=37.970055" title="Search Plazi for locations around (long 127.4052/lat 37.970055)">Nonnamgi-gil</a>, Buk-myeon, Gapyeong-gun, 5. X. 2023, 37°58'12.2"N, 127°24'18.7"E, 402 m, sifting leaf litter &amp; soil, J.-W. Kang, J.-I. Shin ”.</p><p>Diagnosis.</p><p>Vertex with two shallow longitudinal grooves extending from posterior margin to posterior 1 / 2; area between grooves relatively flattened and impressed (Fig. 2 B). Protibiae of male with small subapical pin-like projection (Fig. 2 C). Metaventrite with shallow longitudinal impression (Fig. 2 D). Aedeagus (Fig. 3 E – H) strongly elongated and symmetrical, length 0.23 mm. In ventral view, median lobe gradually widening from base to apical 1 / 4, widest at apical 1 / 4, then slightly narrowing; sides from base to middle somewhat parallel; apex rounded; in lateral view, slightly curved near base and subapical region with strong ventral indentation. Endophallus symmetrical, structure complex. Parameres slender, reaching middle of median lobe; each paramere with two short apical setae and one subapical seta.</p><p>Male description.</p><p>Body length 1.14 mm; reddish-brown, appendages lighter; flattened and elongated; surface covered yellow hairs (Fig. 2 A). Head wider than long, widest across eyes; punctures of surface inconspicuous; hairs short and sparse (Fig. 2 B). Temples nearly 1 / 4 length of eyes (Fig. 2 B). Vertex with small pits on medioposterior margin; two shallow longitudinal grooves extending from posterior margin to posterior 1 / 2 present; area between grooves relatively flattened and impressed (Fig. 2 B). Antennae with distinct distal three-segmented club; antennomere 1 strongly elongate, 2 elongate but less so than 1, 3 slightly wider than long, 4–6 as long as wide, 5 slightly larger than 4, 6 slightly smaller than 5, 7 subpentagonal, 8 distinctly wider than long, 9–11 forming a club (Fig. 2 A). Pronotum distinctly wider than head, as long as wide, widest near middle; anterior margin somewhat rounded, lateral margins strongly rounded in anterior 1 / 3, somewhat parallel in posterior 1 / 3, posterior angles somewhat right-angled, posterior margin weakly sinuate; pronotal base with shallow median pits and transverse impression, lateral pits distinct; punctures of surface inconspicuous; hairs short and sparse (Fig. 2 B). Elytra slightly wider than pronotum, distinctly longer than wide, widest near middle; lateral margins and posterior margin relatively rounded; punctures of surface inconspicuous; hairs short and sparse; each elytron with distinct humeral denticle (Fig. 2 A, B). Hind wings well-developed. Metaventrite with shallow longitudinal impression (Fig. 2 D). Legs moderately long and slender. Protibiae with small subapical pin-like projection (Fig. 2 C). Aedeagus (Fig. 3 E – H) strongly elongated and symmetrical, length 0.23 mm. In ventral view, median lobe gradually widening from base to apical 1 / 4, widest at apical 1 / 4, then slightly narrowing; sides from base to middle somewhat parallel; apex rounded; in lateral view, slightly curved near base and subapical region with strong ventral indentation. Endophallus symmetrical, complex structured. Parameres slender, reaching middle of median lobe; each paramere with two short apical setae and one subapical seta.</p><p>Sexual dimorphism.</p><p>Unknown.</p><p>Distribution.</p><p>South Korea (Fig. 4).</p><p>Etymology.</p><p>The specific epithet is an adjective derived from the type locality ‘ Mt. Myeonggiʼ.</p><p>Habitat.</p><p>This species was collected from soil and leaf litter in mixed forest at relatively low altitudes, unlike other previously known species.</p><p>Remarks.</p><p>Veraphis myeonggiensis can be classified within the japonicus species group based on the characteristics of the male leg, antennae, and aedeagus (Jałoszyński and Hoshina 2005). The aedeagus of this species is similar to that of V. tottoriensis Jałoszyński &amp; Hoshina from Japan. However, the apex of the median lobe is more strongly curved ventrally, and the central structure of the endophallus is absent. Additionally, the parameres are slightly shorter. It also shares a similar aedeagus with V. modestus Jałoszyński but differs in the apex of the median lobe and the structure of the endophallus. Moreover, the overall morphology in lateral view is clearly distinct. Also, this species is externally very similar to E. odaesanensis, but it is smaller in size and clearly differed in the aedeagus.</p></div>	https://treatment.plazi.org/id/34D920B8BC1E57EB8786C07B8F6379D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Byeon, Ui-Joung;Park, Jong-Seok	Byeon, Ui-Joung, Park, Jong-Seok (2025): Two new species of the genus Veraphis Casey (Coleoptera, Staphylinidae, Scydmaeninae) from Korea. ZooKeys 1224: 165-174, DOI: 10.3897/zookeys.1224.142859
375C78297D085017B1EA315EB6007720.text	375C78297D085017B1EA315EB6007720.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Veraphis odaesanensis Byeon & Park 2025	<div><p>Veraphis odaesanensis sp. nov.</p><p>Figs 1, 3 A – D, 4</p><p>Type material designated.</p><p>Holotype • ♂ NIBR: “ Korea: Gangwon Prov. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.56561&amp;materialsCitation.latitude=37.787277" title="Search Plazi for locations around (long 128.56561/lat 37.787277)">Mt. Odae</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.56561&amp;materialsCitation.latitude=37.787277" title="Search Plazi for locations around (long 128.56561/lat 37.787277)">Dongsan-ri</a>, Jinbu-myeon, Pyeongchang-gun, 17. VIII. 2022, 37°47'14.2"N, 128°33'56.2"E, 897 m, sifting leaf &amp; soil litter, J.-W. Seo, J.-I. Shin ” Paratypes Korea: Gangwon Prov. • 1 ♂ 1 ♀ (1 ♂ genitalia dissected; CBNUIC), Pyeongchang-gun, Jinbu-myeon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.56561&amp;materialsCitation.latitude=37.787277" title="Search Plazi for locations around (long 128.56561/lat 37.787277)">Dongsan-ri</a>, 21. IX – 02. XI. 2022, 37°47'14.2"N, 128°33'56.2"E, 897 m, F. I. T, J.-W. Kang, J-I. Shin • 1 ♀ (CBNUIC), 02. XI. 2022, 37°47'14.2"N, 128°33'56.2"E, 897 m, F. I. T, J.-W. Kang, J-I. Shin • 1 ♀ (CBNUIC), 17. VIII. 2022, 37°47'14.2"N, 128°33'56.2"E, 897 m, sifting soil &amp; leaf litter, J.-W. Seo, J-I. Shin • 8 ♀♀ (7 ♀♀ 95 % EtOH in tube; CBNUIC), Odaesan-ro 1211-14, Mt. Odae, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.56561&amp;materialsCitation.latitude=37.787277" title="Search Plazi for locations around (long 128.56561/lat 37.787277)">Sangwonsa</a>, 17. VIII – 21. IX. 2022, 37°47'14.2"N, 128°33'56.2"E, 897 m, F. I. T, J-I. Shin U.-J. Hwang .</p><p>Diagnosis.</p><p>Vertex with two shallow longitudinal grooves extending from posterior margin to posterior 1 / 2; area between grooves relatively flattened and impressed (Fig. 1 B). Protibiae of male with small subapical pin-like projection (Fig. 1 D). Metaventrite with shallow longitudinal impression (Fig. 1 E). Aedeagus (Fig. 3 A – D) strongly elongated and symmetrical, length 0.24 mm. In ventral view, lateral margins of median lobe somewhat parallel, middle of apex weakly protruding, with small shallow punctures in subapical region in ventral view; in lateral view, slightly curved near base and subapical region with strong ventral indentation. Endophallus symmetrical, U-shaped structured. Parameres slender, reaching middle of median lobe; each paramere with two short apical setae and one subapical seta.</p><p>Male description.</p><p>Body length 1.27–1.32 mm; reddish-brown, appendages lighter; flattened and elongated; surface covered yellow hairs (Fig. 1 A). Head wider than long, widest across eyes; punctures of surface inconspicuous; hairs short and sparse (Fig. 1 B). Temples nearly 1 / 4 length of eyes, with shorter hairs than those on surface (Fig. 1 B). Vertex with small pits on medioposterior margin; two shallow longitudinal grooves extending from posterior margin to posterior 1 / 2 present; area between grooves relatively flattened and impressed (Fig. 1 B). Antennae with distinct distal three-segmented club; antennomere 1 strongly elongate, 2 elongate but weaker than 1, 3 slightly wider than long, 4–6 as long as wide, 5 slightly larger than 4, 6 slightly smaller than 5, 7 subpentagonal, 8 distinctly wider than long, 9–11 forming club (Fig. 1 A). Pronotum distinctly wider than head, as long as wide, widest near middle; anterior margin somewhat rounded, lateral margins strongly rounded in anterior 1 / 3, somewhat parallel in posterior 1 / 3, posterior angles somewhat right-angled, posterior margin weakly sinuate; pronotal base with shallow median pits and transverse impression, lateral pits distinct; punctures of surface inconspicuous; hairs short and sparse (Fig. 1 B). Elytra slightly wider than pronotum, distinctly longer than wide, widest near middle; lateral margins and posterior margin relatively rounded; punctures of surface inconspicuous; hairs short and sparse; each elytron with distinct humeral denticle (Fig. 1 A, B). Hind wings well-developed. Metaventrite with shallow longitudinal impression (Fig. 1 E). Legs moderately long and slender. Protibiae with small subapical pin-like projection (Fig. 1 C). Aedeagus (Fig. 3 A – D) strongly elongated and symmetrical, length 0.24 mm. In ventral view, lateral margins of median lobe somewhat parallel, middle of apex weakly protruding, with small shallow punctures in subapical region in ventral view; in lateral view, slightly curved near base and subapical region with strong ventral indentation. Endophallus symmetrical, U-shaped. Parameres slender, reaching middle of median lobe; each paramere with two short apical setae and one subapical seta.</p><p>Sexual dimorphism.</p><p>Protibiae without subapical pin-like projection (Fig. 1 D).</p><p>Distribution.</p><p>South Korea (Fig. 4).</p><p>Etymology.</p><p>The specific epithet is an adjective derived from the type locality ‘ Mt. Odaeʼ.</p><p>Habitat.</p><p>This species was collected from relatively high-altitude mountains in South Korea, at elevations above 800 m. It was frequently captured using flight intercept traps, indicating its ability to fly with well-developed wings. Additionally, it was also collected from soil and leaf litter.</p><p>Remarks.</p><p>Veraphis odaesanensis can be classified within the japonicus species group based on the characteristics of the male leg, antennae, and aedeagus (Jałoszyński and Hoshina 2005). This species shows clear differences in the aedeagus from the Japanese species. In China, it shares a similar aedeagus with V. assingi Jałoszyński, but the apex of the median lobe is more pointed, and there are significant differences in the morphology of the endophallus and the median lobe in lateral view.</p></div>	https://treatment.plazi.org/id/375C78297D085017B1EA315EB6007720	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Byeon, Ui-Joung;Park, Jong-Seok	Byeon, Ui-Joung, Park, Jong-Seok (2025): Two new species of the genus Veraphis Casey (Coleoptera, Staphylinidae, Scydmaeninae) from Korea. ZooKeys 1224: 165-174, DOI: 10.3897/zookeys.1224.142859
