taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
927EBA5465FE5FD7AD4AE4B18288A116.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244339	https://doi.org/10.3897/zookeys.1226.140386.figure3	Figure 3. Wings of two new Santocellus species a S. legrandi sp. nov., holotype, male, Nicaragua, Río San Juan, Bartola (CAS); forewing length, 18.7 mm b S. helene, sp. nov., holotype, male, Bolivia, Cochabamba (CAS); forewing length, 19.4 mm. Arrows highlight interspecific differences in markings.	Figure 3. Wings of two new Santocellus species a S. legrandi sp. nov., holotype, male, Nicaragua, Río San Juan, Bartola (CAS); forewing length, 18.7 mm b S. helene, sp. nov., holotype, male, Bolivia, Cochabamba (CAS); forewing length, 19.4 mm. Arrows highlight interspecific differences in markings.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
927EBA5465FE5FD7AD4AE4B18288A116.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244343	https://doi.org/10.3897/zookeys.1226.140386.figure7	Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm.	Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
927EBA5465FE5FD7AD4AE4B18288A116.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244344	https://doi.org/10.3897/zookeys.1226.140386.figure8	Figure 8. Side-by-side comparisons of male external abdominal features a, c, e Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) b, d, f Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a, b abdominal segments A 6 - terminus, lateral c, d terminus of S 8 + 9, with terminal membrane bearing distinctive patches of gonocristae e, f callus cerci.	Figure 8. Side-by-side comparisons of male external abdominal features a, c, e Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) b, d, f Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a, b abdominal segments A 6 - terminus, lateral c, d terminus of S 8 + 9, with terminal membrane bearing distinctive patches of gonocristae e, f callus cerci.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
927EBA5465FE5FD7AD4AE4B18288A116.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244345	https://doi.org/10.3897/zookeys.1226.140386.figure9	Figure 9. Gonarcal complex of Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a posterolateral b lateral c frontolateral d posteroventral e dorsal. Abbreviations: gc gonocornu (fused mesally, forming rounded plate); gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme, dorsal view; g. br. gonarcal bridge; mu-d mediuncus, dorsal view; mu-f mediuncus, frontal view; rod internal mediuncal apodemes.	Figure 9. Gonarcal complex of Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a posterolateral b lateral c frontolateral d posteroventral e dorsal. Abbreviations: gc gonocornu (fused mesally, forming rounded plate); gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme, dorsal view; g. br. gonarcal bridge; mu-d mediuncus, dorsal view; mu-f mediuncus, frontal view; rod internal mediuncal apodemes.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
927EBA5465FE5FD7AD4AE4B18288A116.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244346	https://doi.org/10.3897/zookeys.1226.140386.figure10	Figure 10. Side-by-side comparison of head and body (dorsum) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Madre de Dios, Peru d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba f sketch of thoracic sclerites (from Séméria 1976).	Figure 10. Side-by-side comparison of head and body (dorsum) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Madre de Dios, Peru d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba f sketch of thoracic sclerites (from Séméria 1976).	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
927EBA5465FE5FD7AD4AE4B18288A116.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244347	https://doi.org/10.3897/zookeys.1226.140386.figure11	Figure 11. Side-by-side comparison of the head (frontal) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba.	Figure 11. Side-by-side comparison of the head (frontal) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
927EBA5465FE5FD7AD4AE4B18288A116.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244348	https://doi.org/10.3897/zookeys.1226.140386.figure12	Figure 12. Side-by-side comparison of the wings of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e Santocellus helene sp. nov., male; Bolivia, Cochabamba. The numbers on the right of each image indicate the maximum length of the forewing recorded for that species.	Figure 12. Side-by-side comparison of the wings of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e Santocellus helene sp. nov., male; Bolivia, Cochabamba. The numbers on the right of each image indicate the maximum length of the forewing recorded for that species.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244337	https://doi.org/10.3897/zookeys.1226.140386.figure1	Figure 1. Head of Santocellus legrandi, sp. nov. Holotype, male, Nicaragua, Río San Juan, Bartola (CAS) a frontal b head, prothorax, dorsal c head, frontolateral d ventral. Width across head (dorsal, including eyes), 1.85 mm.	Figure 1. Head of Santocellus legrandi, sp. nov. Holotype, male, Nicaragua, Río San Juan, Bartola (CAS) a frontal b head, prothorax, dorsal c head, frontolateral d ventral. Width across head (dorsal, including eyes), 1.85 mm.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244338	https://doi.org/10.3897/zookeys.1226.140386.figure2	Figure 2. External features of Santocellus legrandi, sp. nov. Holotype, male, Nicaragua, Río San Juan, Bartola (CAS) a head, thorax, dorsal b body, lateral c abdomen, dorsal d terminus, lateral e protarsal claw, lateral. Width across head (including eyes), 1.85 mm; diameter of callus cerci, ~ 0.2 mm.	Figure 2. External features of Santocellus legrandi, sp. nov. Holotype, male, Nicaragua, Río San Juan, Bartola (CAS) a head, thorax, dorsal b body, lateral c abdomen, dorsal d terminus, lateral e protarsal claw, lateral. Width across head (including eyes), 1.85 mm; diameter of callus cerci, ~ 0.2 mm.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244339	https://doi.org/10.3897/zookeys.1226.140386.figure3	Figure 3. Wings of two new Santocellus species a S. legrandi sp. nov., holotype, male, Nicaragua, Río San Juan, Bartola (CAS); forewing length, 18.7 mm b S. helene, sp. nov., holotype, male, Bolivia, Cochabamba (CAS); forewing length, 19.4 mm. Arrows highlight interspecific differences in markings.	Figure 3. Wings of two new Santocellus species a S. legrandi sp. nov., holotype, male, Nicaragua, Río San Juan, Bartola (CAS); forewing length, 18.7 mm b S. helene, sp. nov., holotype, male, Bolivia, Cochabamba (CAS); forewing length, 19.4 mm. Arrows highlight interspecific differences in markings.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244340	https://doi.org/10.3897/zookeys.1226.140386.figure4	Figure 4. Male abdomen of Santocellus legrandi, sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) a dorsum of abdominal segments A 2 - terminus b lateral view of abdominal segments A 6 - terminus c dorsum of abdominal segments A 7 to terminus with gonarcus everted d lateral view of abdominal terminus. Length of T 8 along dorsal margin, 0.68 mm. Abbreviations: c. c. callus cerci; d. ap. dorsal apodeme of T 9 + ectoproct; gcr field of gonocristae on terminal membrane of S 9; gs gonarcus; S 8 + 9 fused eighth and ninth sternites; T 3, T 7, T 8 third, seventh, and eighth tergites; T 9 + e fused ninth tergite and ectoproct; v. ap. ventral apodeme of S 8 + 9.	Figure 4. Male abdomen of Santocellus legrandi, sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) a dorsum of abdominal segments A 2 - terminus b lateral view of abdominal segments A 6 - terminus c dorsum of abdominal segments A 7 to terminus with gonarcus everted d lateral view of abdominal terminus. Length of T 8 along dorsal margin, 0.68 mm. Abbreviations: c. c. callus cerci; d. ap. dorsal apodeme of T 9 + ectoproct; gcr field of gonocristae on terminal membrane of S 9; gs gonarcus; S 8 + 9 fused eighth and ninth sternites; T 3, T 7, T 8 third, seventh, and eighth tergites; T 9 + e fused ninth tergite and ectoproct; v. ap. ventral apodeme of S 8 + 9.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244341	https://doi.org/10.3897/zookeys.1226.140386.figure5	Figure 5. Male genitalia of Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) a gonarcal complex, dorsal b same, lateral c same, frontal d same, posteroventral. Abbreviations: gcr, field of gonocristae on terminal membrane of ninth sternite; gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme; mu-d mediuncus, dorsal view; mu-f mediuncus.	Figure 5. Male genitalia of Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) a gonarcal complex, dorsal b same, lateral c same, frontal d same, posteroventral. Abbreviations: gcr, field of gonocristae on terminal membrane of ninth sternite; gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme; mu-d mediuncus, dorsal view; mu-f mediuncus.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244342	https://doi.org/10.3897/zookeys.1226.140386.figure6	Figure 6. Cleared abdominal terminus of female Santocellus legrandi sp. nov., allotype, Costa Rica, San José (SDMC) a segments A 7 to terminus, lateral b genitalia, lateral c subgenitale, distal view d spermathecal complex below bursa copulatrix, lateral view. Abbreviations: b. c. bursa copulatrix; c. c. callus cerci; g. l. gonapophysis lateralis; k-sg distal knob of subgenitale; sg-b base of subgenitale; sp spermatheca; sp. d. spermathecal duct; sp. i. spermathecal invagination; S 7 seventh sternite; T 8 eighth tergite; vel, spermathecal velum.	Figure 6. Cleared abdominal terminus of female Santocellus legrandi sp. nov., allotype, Costa Rica, San José (SDMC) a segments A 7 to terminus, lateral b genitalia, lateral c subgenitale, distal view d spermathecal complex below bursa copulatrix, lateral view. Abbreviations: b. c. bursa copulatrix; c. c. callus cerci; g. l. gonapophysis lateralis; k-sg distal knob of subgenitale; sg-b base of subgenitale; sp spermatheca; sp. d. spermathecal duct; sp. i. spermathecal invagination; S 7 seventh sternite; T 8 eighth tergite; vel, spermathecal velum.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244344	https://doi.org/10.3897/zookeys.1226.140386.figure8	Figure 8. Side-by-side comparisons of male external abdominal features a, c, e Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) b, d, f Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a, b abdominal segments A 6 - terminus, lateral c, d terminus of S 8 + 9, with terminal membrane bearing distinctive patches of gonocristae e, f callus cerci.	Figure 8. Side-by-side comparisons of male external abdominal features a, c, e Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) b, d, f Santocellus helene sp. nov., holotype, Bolivia, Cochabamba, Alto Palmar (CAS) a, b abdominal segments A 6 - terminus, lateral c, d terminus of S 8 + 9, with terminal membrane bearing distinctive patches of gonocristae e, f callus cerci.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244346	https://doi.org/10.3897/zookeys.1226.140386.figure10	Figure 10. Side-by-side comparison of head and body (dorsum) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Madre de Dios, Peru d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba f sketch of thoracic sclerites (from Séméria 1976).	Figure 10. Side-by-side comparison of head and body (dorsum) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Madre de Dios, Peru d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba f sketch of thoracic sclerites (from Séméria 1976).	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244347	https://doi.org/10.3897/zookeys.1226.140386.figure11	Figure 11. Side-by-side comparison of the head (frontal) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba.	Figure 11. Side-by-side comparison of the head (frontal) of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce, female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e S. helene sp. nov., male; Bolivia, Cochabamba.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244348	https://doi.org/10.3897/zookeys.1226.140386.figure12	Figure 12. Side-by-side comparison of the wings of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e Santocellus helene sp. nov., male; Bolivia, Cochabamba. The numbers on the right of each image indicate the maximum length of the forewing recorded for that species.	Figure 12. Side-by-side comparison of the wings of currently known Santocellus species a S. atlanticis, male; Brasil, Rio Grande do Sul b S. riodoce female; Brasil, Espírito Santo c S. risi, male; Peru, Madre de Dios d S. legrandi sp. nov., male; Nicaragua, Río San Juan, Bartola e Santocellus helene sp. nov., male; Bolivia, Cochabamba. The numbers on the right of each image indicate the maximum length of the forewing recorded for that species.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
E596F489F95457B7BC3DD5E91CE47B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1244343	https://doi.org/10.3897/zookeys.1226.140386.figure7	Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm.	Figure 7. Head and thoracic region of Santocellus helene sp. nov., holotype, male, Bolivia, Cochabamba, Alto Palmar (CAS) a upper body (head, thorax) dorsal b head, prothorax, lateral c head, frontal d head, ventral e tarsus (dorsum) and tarsal claw (lateral). Width across head (including eyes) 1.82 mm.	2025-02-07	Tauber, Catherine A.		Zenodo	biologists	Tauber, Catherine A.			
