taxonID	type	description	language	source
3B6987A0FFBF806EFE6FFEF8FEB6FBE0.taxon	type_taxon	Type species Kamchatkapora o zhgibesovi sp. n.	en	Taylor, Paul D., Grischenko, Andrei V. (2024): A new bryozoan genus from the Sea of Okhotsk and the taxonomy and geological history of ‘ stratocormidial’ cyclostome bryozoans. Journal of Natural History (J. Nat. Hist.) 58 (41 - 44): 1917-1935, DOI: 10.1080/00222933.2024.2396995, URL: http://dx.doi.org/10.1080/00222933.2024.2396995
3B6987A0FFBF806EFE6FFEF8FEB6FBE0.taxon	etymology	Etymology In reference to the type locality off the Kamchatka Peninsula.	en	Taylor, Paul D., Grischenko, Andrei V. (2024): A new bryozoan genus from the Sea of Okhotsk and the taxonomy and geological history of ‘ stratocormidial’ cyclostome bryozoans. Journal of Natural History (J. Nat. Hist.) 58 (41 - 44): 1917-1935, DOI: 10.1080/00222933.2024.2396995, URL: http://dx.doi.org/10.1080/00222933.2024.2396995
3B6987A0FFBF806EFE6FFEF8FEB6FBE0.taxon	diagnosis	Diagnosis Colony massive, probably hemispherical when complete, multilamellar, each layer formed of numerous cup-shaped subcolonies overarching earlier layers joined at their outer edges; autozooids free-walled, apertures clustered into small groups often elongated radially from subcolony centres, minimally differentiated from kenozooids but very slightly larger and with walls prolonged into low spines at aperture corners; kenozooids free-walled, more numerous than autozooids, occupying centres of subcolonies and surrounding autozooids elsewhere; gonozooids fixed-walled, the densely pseudoporous roof sutured, lobate, surrounding autozooid clusters, the lobes extending between autozooid clusters, ooeciopore elliptical.	en	Taylor, Paul D., Grischenko, Andrei V. (2024): A new bryozoan genus from the Sea of Okhotsk and the taxonomy and geological history of ‘ stratocormidial’ cyclostome bryozoans. Journal of Natural History (J. Nat. Hist.) 58 (41 - 44): 1917-1935, DOI: 10.1080/00222933.2024.2396995, URL: http://dx.doi.org/10.1080/00222933.2024.2396995
3B6987A0FFBF806EFE6FFEF8FEB6FBE0.taxon	discussion	Remarks Among existing stratocormidial cyclostome genera, Kamchatkapora resembles Multifascigera d’Orbigny, 1853 in having autozooidal apertures grouped into clusters that usually radiate from the centres of the subcolonies. However, in Multifascigera the clusters are fascicles bounded by exterior wall calcification, lacking in Kamchatkapora, and the colony surface between clusters is formed of exterior wall and not open kenozooids as in Kamchatkapora. The new genus also shows similarities to Semimulticavea d’Orbigny, 1853 with respect to the fixed-walled gonozooids and free-walled autozooids and kenozooids, as well as the general growth pattern. However, autozooidal apertures are arranged in well-defined uniserial rows in Semimulticavea, unlike the irregular clusters seen in Kamchatkapora. Further comparisons with established stratocormidial cyclostomes can be found in the Discussion section below. Among extant cyclostome genera, the fixed-walled gonozooids and free-walled autozooids and kenozooids of Kamchatkapora together with the overall lamellar colony-form prompts comparison with the extant genus Favosipora MacGillivray, 1885, which was revised by Gordon and Taylor (2001). Despite sometimes forming multilamellar colonies, as in F. ainui Taylor and Grischenko, 2015 from northern Japan, Favosipora is not known to construct large and massive colonies, nor does it possess well-defined subcolonies on the same plane (cf. the vertically staked subcolonies of F. bathyalis Gordon and Taylor, 2010). The similarity in skeletal organisation and configuration of the autozooids and gonozooids suggests a close relationship between Kamchatkapora and Favosipora notwithstanding the stratocormidial colony-form of Kamchatkapora. For this reason, Kamchatkapora is provisionally assigned to the same family – Densiporidae – as Favosipora until molecular data allows its affinities to be more firmly established. The fusiform colonies of Densipora, the type genus of Densiporidae, have screw-like ridges on their surface and the autozooidal apertures are surrounded by about 6 – 9 rounded spines processes (Bock et al. 2018), neither feature apparent in Kamchatkopora. Composition Monospecific.	en	Taylor, Paul D., Grischenko, Andrei V. (2024): A new bryozoan genus from the Sea of Okhotsk and the taxonomy and geological history of ‘ stratocormidial’ cyclostome bryozoans. Journal of Natural History (J. Nat. Hist.) 58 (41 - 44): 1917-1935, DOI: 10.1080/00222933.2024.2396995, URL: http://dx.doi.org/10.1080/00222933.2024.2396995
3B6987A0FFB98063FE91FB75FD7FF9CE.taxon	description	(Figures 2 – 5)	en	Taylor, Paul D., Grischenko, Andrei V. (2024): A new bryozoan genus from the Sea of Okhotsk and the taxonomy and geological history of ‘ stratocormidial’ cyclostome bryozoans. Journal of Natural History (J. Nat. Hist.) 58 (41 - 44): 1917-1935, DOI: 10.1080/00222933.2024.2396995, URL: http://dx.doi.org/10.1080/00222933.2024.2396995
3B6987A0FFB98063FE91FB75FD7FF9CE.taxon	materials_examined	Type material Holotype: NHMUK 2017.7.11.2 (Figure 2 (C, D )), Station 190, Sea of Okhotsk, shelf off western Kamchatka Peninsula, 56 ° 51 ' 1 '' N, 154 ° 52 ' 0 '' E, 114.5 m depth, dredged using a beam trawl from RV Professor Probatov, 6 August 2013, collected by T. B. Morozov. Paratypes: NHMUK 2017.7.11.3 (Figure 2 (E, F )), 2017.7.11.4 (Figure 2 (A, B )); ZIRAS 1 / 53,003, 13 colony fragments.	en	Taylor, Paul D., Grischenko, Andrei V. (2024): A new bryozoan genus from the Sea of Okhotsk and the taxonomy and geological history of ‘ stratocormidial’ cyclostome bryozoans. Journal of Natural History (J. Nat. Hist.) 58 (41 - 44): 1917-1935, DOI: 10.1080/00222933.2024.2396995, URL: http://dx.doi.org/10.1080/00222933.2024.2396995
3B6987A0FFB98063FE91FB75FD7FF9CE.taxon	etymology	Etymology Honorific for Prof. Dr Vladimir P. Ozhgibesov, Head of Regional and Oil Geology Department (1992 – 2003) of the Perm State National Research University, Perm, Russia, in recognition of his contribution to the knowledge of Permian bryozoans of Pre-Urals.	en	Taylor, Paul D., Grischenko, Andrei V. (2024): A new bryozoan genus from the Sea of Okhotsk and the taxonomy and geological history of ‘ stratocormidial’ cyclostome bryozoans. Journal of Natural History (J. Nat. Hist.) 58 (41 - 44): 1917-1935, DOI: 10.1080/00222933.2024.2396995, URL: http://dx.doi.org/10.1080/00222933.2024.2396995
3B6987A0FFB98063FE91FB75FD7FF9CE.taxon	description	Description Colony massive, largest fragment measuring 5.5 cm high by 7 cm wide and having 19 layers (Figure 2 (D )), colonies probably hemispherical when complete; multilamellar, each layer formed of numerous cup-shaped subcolonies overarching earlier layers, vertical distance between layers averaging 2.42 mm (range 1.92 – 3.0 mm, SD 0.39 mm, N = 10); subcolonies either fused at their outer edges to form compound structures of 2 – 8 subcolonies (Figure 3), or irregularly abutting one another; fractured specimens reveal vertical stacks of subcolonies, each new subcolony generally positioned directly over the centre of a subcolony in the underlying layer (Figure 2 (D )). New subcolonies apparently originate through upward growth of kenozooids at subcolony centres and between clusters of autozooids; subcolony undersides formed on basal exterior walls, slightly rugose transversely to growth direction, lacking pseudopores. Colony surface covered in hummocks (monticules) centred on the subcolonies. Early astogeny unknown. Subcolonies circular to elliptical in surface view, averaging 6.6 mm (range 5.0 – 9.7 mm, SD 1.5 mm, N = 10) in maximum width by 4.8 mm (range 3.8 – 7.0 mm, SD 1.0 mm, N = 10) in minimum width; central, autozooid-bearing area of subcolonies averaging 3.9 mm (range 2.8 – 5.4 mm, SD 0.9 mm, N = 10) in maximum width by 3.2 mm (range 2.4 – 4.3 mm, SD 0.5 mm, N = 10). Autozooids free-walled, apertures rounded polygonal, clustered into small groups often elongated radially from subcolony centres (Figure 4 (B )), minimally differentiated from kenozooids but very slightly larger and with walls prolonged into low spines 20 – 30 µm high at aperture corners (Figure 4 (C )). Aperture minimum diameter averaging 162 µm (range 143 – 202 µm, SD 19.8 µm, N = 10). Kenozooids free-walled, more numerous than autozooids, occupying centres and peripheries of subcolonies and surrounding autozooids elsewhere (Figure 3 (A )); of similar size to autozooids but more variable; apertures rarely closed by a diaphragm, rounded polygonal, minimum diameter averaging 134 µm (range 66 – 192 µm, SD 36.7 µm, N = 10). Gonozooids fixed-walled (Figure 5 (A, D )), the densely pseudoporous roof sutured (Figure 5 (C )), lobate, surrounding autozooid clusters, the lobes spreading out between autozooid clusters, measuring about 1.5 – 2.5 mm in maximum width by 0.9 – 2.0 mm in minimum width. Ooeciopore elliptical, 242 µm by 11 µm in single measured example, collapsed slightly inwards of the brood chamber roof (Figure 5 (B )). Oeciostome short, funnel-shaped. Interior walls close to apertures exhibiting a distally foliated ultrastructural fabric (Figure 4 (D )), the tabular fibrous crystallites averaging in width 3.69 µm (range 1.17 – 6.98 µm, SD 1.8 µm, N = 10), pierced by irregularly spaced interzooidal pores. Mural spines (Figure 4 (E, F )) developed widely on surfaces of interior walls, length averaging 18.1 µm (range 11.3 – 27.1 µm, SD 4.2 µm, N = 10), shaft moderately straight, ending in a head 10 – 16 µm in diameter bearing spike-like spinelets each up to 5 µm in length. Exterior wall roofing gonozooids with circular or elliptical (Figure 5 (C )), diameter averaging 4.04 µm (range 3.01 – 4.99 µm, SD 0.53 µm, N = 10), average centre-to-centre spacing 10.26 µm (range 8.79 – 12.11 µm, SD 1.09 µm, N = 10).	en	Taylor, Paul D., Grischenko, Andrei V. (2024): A new bryozoan genus from the Sea of Okhotsk and the taxonomy and geological history of ‘ stratocormidial’ cyclostome bryozoans. Journal of Natural History (J. Nat. Hist.) 58 (41 - 44): 1917-1935, DOI: 10.1080/00222933.2024.2396995, URL: http://dx.doi.org/10.1080/00222933.2024.2396995
3B6987A0FFB98063FE91FB75FD7FF9CE.taxon	discussion	Remarks All of the material available is fragmentary. Some of the fragments fit together and it is thought likely that all are pieces of a single large colony broken up during trawling. If so, the intact colony may have been more than 15 cm across. Details of the growth pattern and zooidal budding in Kamchatkapora await study using serial thin sectioning or, preferably, Computerised Tomographic (CT) scanning (see Key and Wyse Jackson 2022 for a review of this application in bryozoology). Preliminary observations suggest that new subcolonies were generated mostly by upward growth from groups of kenozooids at the centres of earlier subcolonies. Intrazooidal fission like that described from some other stratocormidial cyclostomes (eg Hillmer et al. 1975) has not been observed and may not play a role in the formation of new subcolonies.	en	Taylor, Paul D., Grischenko, Andrei V. (2024): A new bryozoan genus from the Sea of Okhotsk and the taxonomy and geological history of ‘ stratocormidial’ cyclostome bryozoans. Journal of Natural History (J. Nat. Hist.) 58 (41 - 44): 1917-1935, DOI: 10.1080/00222933.2024.2396995, URL: http://dx.doi.org/10.1080/00222933.2024.2396995
3B6987A0FFB98063FE91FB75FD7FF9CE.taxon	distribution	Distribution Sea of Okhotsk, off Kamchatka, north-eastern Pacific.	en	Taylor, Paul D., Grischenko, Andrei V. (2024): A new bryozoan genus from the Sea of Okhotsk and the taxonomy and geological history of ‘ stratocormidial’ cyclostome bryozoans. Journal of Natural History (J. Nat. Hist.) 58 (41 - 44): 1917-1935, DOI: 10.1080/00222933.2024.2396995, URL: http://dx.doi.org/10.1080/00222933.2024.2396995
