taxonID	type	description	language	source
3A4687F46A6CFF83FF1CFDD9BAFBF74D.taxon	materials_examined	Holotype: KIZ 039283, adult male, collected from the E’ga Pass, Luobenzhuo Village, Lushui County, Nujiang Lisu Autonomous Prefecture, Yunnan Province, China (98.74384 ºE, 26.42762 ºN, elevation 3081 m, GCJ 02) by Zhong-Bin Yu and Shao-Bing Hou 30 October, 2021. Paratypes (six adult males, one adult female): KIZ 038628, 038640, 038644, adult males, KIZ 038641, adult female, from Tingming Lake, Lushui County, Nujiang Lisu Autonomous Prefecture, Yunnan Province, China (98.68500 ºE, 26.05722 ºN, elevation 3546 m, GCJ 02) by Zhong-Bin Yu, Dong An, and Tian-En Chen on 4 August, 2021; KIZ 038629, 038631, adult males, from Pianma, Lushui County, Nujiang Lisu Autonomous Prefecture, Yunnan Province, China (98.63600 ºE, 25.98334 ºN, elevation 2307 m, GCJ 02) by Zhong-Bin Yu, Dong An, and Tian-En Chen on 4 August 2021.	en	Wang, Kai, Yu, Zhongbin, Wu, Yunhe, Hou, Shao-Bing, Wang, De, Xiong, Yun, Ye, Xin-Long, Duan, Jian-Ping, Li, Hai-Jun, Li, Jie, Jin, Jie-Qiong, Yang, Shen- Pin, Che, Jing (2025): Three New Species of the Genus Scutiger (Amphibia: Anura: Megophryidae) from the Gaoligong Mountain Range in China, with An Updated Key to the Chinese Congeners. Zootaxa 5725 (2): 203-230, DOI: 10.11646/zootaxa.5725.2.2, URL: https://doi.org/10.11646/zootaxa.5725.2.2
3A4687F46A6CFF83FF1CFDD9BAFBF74D.taxon	etymology	Etymology: The species name “ pardalotus ” is derived from Greek, which means “ spotted like a leopard ”. It describes the beautiful spotted coloration patterns on the dorsum of the species. The proposed Chinese common name of the new species is “ Ë ỄÊẊḆ ” (pinyin: Bao Ban Chi Tu Chan), and the proposed English name is Leopard Lazy Toad.	en	Wang, Kai, Yu, Zhongbin, Wu, Yunhe, Hou, Shao-Bing, Wang, De, Xiong, Yun, Ye, Xin-Long, Duan, Jian-Ping, Li, Hai-Jun, Li, Jie, Jin, Jie-Qiong, Yang, Shen- Pin, Che, Jing (2025): Three New Species of the Genus Scutiger (Amphibia: Anura: Megophryidae) from the Gaoligong Mountain Range in China, with An Updated Key to the Chinese Congeners. Zootaxa 5725 (2): 203-230, DOI: 10.11646/zootaxa.5725.2.2, URL: https://doi.org/10.11646/zootaxa.5725.2.2
3A4687F46A6CFF83FF1CFDD9BAFBF74D.taxon	diagnosis	Diagnosis: Scutiger pardalotus sp. nov. can be diagnosed from all congeners by a combination of the following morphological characters: (1) body size moderate, SVL 47.5 – 56.4 mm; (2) forearm and hand length moderate, LFHL 43.0 – 49.9 % SVL; (3) posterior head moderate, PHL 34.7 – 42.5 % HL; (4) inter-nare distance long, IND 21.6 – 29.4 % HW; (5) snout long, SEL 37.5 – 45.2 % HL; (6) eyes large, ED 20.5 – 28.4 % HL; (5) single pair of pectoral glands and single pair of axillary glands present in males; (6) fine spines present on pectoral glands, but not on peripheral margin of chin on ventral head nor axillary glands; (7) maxillary teeth present; (8) tongue rounded; (9) nuptial pads present only on first two fingers in males, covered with a few distinctively enlarged black spines; (10) no distinct enlarged, raised tubercles scattered on dorsal and lateral forearms and anterior brachium; (11) no raised tubercles present on dorsal eyelids and supralabial; (12) vocal sacs present; (13) supratympanic fold strongly developed, extend posterior of axillary; (14) toes with distinct, well developed dermal fringes and well developed webbing, reaching tip of toe I and half webbed on toe II and III; (15) parietal head flat; (16) dorsum smooth or with a few relatively flat tubercles; and (17) dorsal surface Pale Pinkish Buff (Color 3) to Light Orange Yellow (Color 7), speckled with Brunt Umber (Color 48), irregular spots and dorsolateral stripes. Comparisons: The new species is morphologically most similar to and phylogenetically sister to S. gongshanensis, but S. pardalotus sp. nov. can be diagnosed from the S. gongshanensis readily by having differential webbing between toes (half-webbed vs. only feebly webbed). From the other new species described above, S. pardalotus sp. nov. differs from S. lisu sp. nov. by having a smaller body size in adults (SVL <56.4 mm vs. ≥ 75.2 mm), longer snout (SEL 36.4 – 45.2 % HL vs. 33.4 – 35.3 %), smaller patches of pectoral glands in males (vs. much larger), different webbing on toes (half webbed vs. feebly webbed), better developed dermal fringes on toes (vs. much weaker), a distinct dorsal coloration (dorsal surface Pale Pinkish Buff [Color 3] to Light Orange Yellow [Color 7], speckled with Brunt Umber [Color 48], irregular spots and dorsolateral stripes vs. dorsal surface near uniformly Grayish Horn Color [Color 268] to Hair Brown [Color 277]), as well as by the presence of vocal sacs in males (vs. absence). For species distributed closely along the Gaoligong Mountain and the adjacent Biluo Snow Mountain, S. pardalotus sp. nov. differs from all by the presence of vocal sacs (vs. absence) and a spotted coloration pattern (vs. uniformly colored or not heavily spotted). Additionally, S. pardalotus sp. nov. differs from S. biluoensis, S. meiliensis, S. tengchongensis by the absence of distinct axillary glands in males (vs. presence) and by a differential webbing between toes (half webbed vs. no webbing or only feebly webbed). For remaining species of the genus, S. pardalotus sp. nov. differs from the S. boulengeri complex (including S. boulengeri and S. bangdaensis, as well as the two synonyms of S. boulengeri, namely Cophophryne alticola and Aelurophryne tainingensis), as well as from S. bhutanensis, S. chintingensis, S. ghunsa, S. glandulatus, S. jiulongensis, S. liupanensis, S. mammatus, S. maculatus, S. nepalensis, S. ningshanensis, S. nyingchiensis, S. pingwuensis, S. sikimmensis S. spinosus, S. tuberculatus, S. wanglangensis, S. wolong, and S. wuguanfui by the absence of nuptial spines on axillary glands in breeding males (vs. presence); from S. muliensis, S. nyingchiensis, S. sikkimensis by the presence of maxillary teeth (vs. absence); from S. adungensis by much weaker toe webbings (half webbed vs. one-fourth or less) and by the absence of distinct round tubercles on dorsum (vs. presence); and from S. chintingensis, S. liupanensis, S. ningshanensis, S. wanglangensis, and S. wolong by the presence of vocal sacs (vs. absence).	en	Wang, Kai, Yu, Zhongbin, Wu, Yunhe, Hou, Shao-Bing, Wang, De, Xiong, Yun, Ye, Xin-Long, Duan, Jian-Ping, Li, Hai-Jun, Li, Jie, Jin, Jie-Qiong, Yang, Shen- Pin, Che, Jing (2025): Three New Species of the Genus Scutiger (Amphibia: Anura: Megophryidae) from the Gaoligong Mountain Range in China, with An Updated Key to the Chinese Congeners. Zootaxa 5725 (2): 203-230, DOI: 10.11646/zootaxa.5725.2.2, URL: https://doi.org/10.11646/zootaxa.5725.2.2
3A4687F46A6CFF83FF1CFDD9BAFBF74D.taxon	description	Description of holotype: Adult male, body moderate, SVL 52.6 mm; head large, dorsally compressed, width about equal to length, HW 99.4 % HL; parietal region slightly concave; snout relatively long, slightly pointy anteriorly, projecting beyond jaw, SEL 41.4 % HL; lateral supralabial convex posterior to nares; nares closer to snout than eyes, facing dorsolaterally, further apart from each other, IN 25.1 % HW; loreal not concave; canthus rostralis distinct; eye raised, large, ED 27.6 % HL, DEW 23.6 % HL; iris ellipse shaped, vertically oriented; tympanum concealed; frontal and temporal flat; supratympanic folds strongly developed, raised, extending posteriorly to posterior jaw only. Limbs sturdy; forearm muscular, FAL 48.1 % SVL, thicken toward elbows, FAW 25.2 % FAL; fingers relative thick, short, free of webbing, finger length I <II <IV <III; a few, distinctively enlarged, raised nuptial spines present on each of first two fingers; metacarpal tubercles and subdigital tubercles indistinct. Hindlimb muscular, strong, relatively short, heels not meeting anteriorly and reaching shoulder with tip of longest toe when adpressed, HLL 127.1 % SVL, FEM 38.5 % SVL, TFL 64.3 % SVL, TIB 36.0 % SVL; foot longer than tibia, FL 122.6 % TIB; toe tips rounded, toe length I <II <III <V <IV; toes with distinct dermal fringes, well developed; webbing well developed, webbing formula I 1 – 2 - II 11 / 2 – 2 + III 2 – 3 + IV 4 - – 21 / 2 V; subdigital tubercles present on inner two toes, indistinct under remining ones; subarticular tubercles absent, subdigital ridges distinct; inner metatarsal tubercle distinct, oval shaped, about same length as first toe, outer metacarpal tubercle absent. Dorsal skins of head, body, and limbs smooth overall, with few flat warts and tubercles on dorsum; fine tubercles covering regions from posterior supralabial to corner of mouth inferior of supratympanic fold; dorsal head smooth; tubercles on dorsum moderate in size, feebly raised; dorsal forelimbs smooth, medial side with few raised tubercles; skin of hindlimbs smooth. Areas near cloaca with few flat tubercles. No spines on ventral lower lip; single pair of pectoral glands present on chest, elongated, oriented lateroposteriorly, covered with residuals of fine spines; single pair of axillary gland distinct, free from spines; remaining ventral body smooth. Coloration: In life, the dorsal background coloration is Warm Buff (Color 4) to Salmon Color (Color 58), and the remaining coloration patterns of the dorsal surfaces are all Brunt Umber (Color 48). A distinct eye stripe is present from the tip of snout to the corner of mouth along the inferior edge of supratympanic fold on each side. A single “ T ” - shaped pattern is situated on dorsal head, with the top of the T-figure located between eyes and tip of the T-figure extending till mid dorsum. Two parallel, somewhat irregular, dorsolateral stripes are present lateral of the T-figure from the temporal head to the groin. Dense circular spots and irregular patches of Brunt Umber (Color 48) are scattered on remaining areas of dorsum and dorsal limbs, with the lateral ones arranged in somewhat lines. The ventral background coloration is Rose Pink (Color 243). An intermixed Pale Pinksih Buff (Color 3) and Walnut Brown (Color 27) marble patterns are present on lower chin, ventrolateral body, and ventrolateral limbs. Nuptial spines on finger Jet Black (Color 300). Variation: Although only female is available, in addition to the lack of nuptial spines on fingers and the absence of pectoral glands, sexual dimorphism is evident, where the female is slightly larger than males (SVL 56.4 mm vs. 51.1 – 52.8 mm), has thinner arms (FAW 22 % FAL vs. 25.2 – 32.8 %), shorter arms and hands (LFHL 43 % SVL vs. 46.9 – 49.9 %), shorter hind limbs (HLL 123.3 % SVL vs. 138.9 – 145.5 %). Among male specimens, only the holotype has completely developed nuptial spines on fingers (i. e. nuptial spines completely covered with a continuous black keratin layer), where spines of all remaining males (KIZ 038628, 038629, 038631, 038640, 038644, and 039283) are under development. KIZ 038640 has only a dark vertebral stripe, where the dorsolateral stripe on each side of the vertebral stripe is replaced by dorsolateral series of spots. Webbing formula remains consistent among males and does not differ between the two sexes. Natural history and conservation: Currently S. pardalotus sp. nov. is known from the two localities only, which are about 40 km apart in linear distance along the Gaoligong Mountain Range (Fig. 1). It is likely to occur in additional sites along the mountain range in both China and Myanmar. The only known amphibian species sympatric with the new species is Amolops yangi. Most individuals of S. pardalotus sp. nov. were found along shoreline of lakes at night (Fig. 12, D and E), while a few were spotted in the forests at night close to the lake. Newly hatched tadpoles were found under rocks in the water in late October, and the female paratype (KIZ 038641) collected in September was gravid, so the breeding season might be between August and October. Individuals were found commonly infested with green aquatic leaches. With little known on the natural history and distribution range of the species, we recommend list it as Data Deficient (DD) for its conservation status, pending on future studies.	en	Wang, Kai, Yu, Zhongbin, Wu, Yunhe, Hou, Shao-Bing, Wang, De, Xiong, Yun, Ye, Xin-Long, Duan, Jian-Ping, Li, Hai-Jun, Li, Jie, Jin, Jie-Qiong, Yang, Shen- Pin, Che, Jing (2025): Three New Species of the Genus Scutiger (Amphibia: Anura: Megophryidae) from the Gaoligong Mountain Range in China, with An Updated Key to the Chinese Congeners. Zootaxa 5725 (2): 203-230, DOI: 10.11646/zootaxa.5725.2.2, URL: https://doi.org/10.11646/zootaxa.5725.2.2
3A4687F46A71FF84FF1CFE4DBBCBF575.taxon	materials_examined	Holotype: KIZ 51834, adult male, from the western slope of the Gaoligong Mountain, facing to the Dulong Valley, Gongshan County, Nujiang Prefecture, Yunnan Province, China (98.46359 ºE, 27.84004 ºN, elevation 2874 m, GCJ 02). Collected by Kai Wang, Zhuo-Yu Lu, Yi-Lin Zhang, and Xian-Kun Huang on 27 May, 2023. Paratype (one adult male, two adult females): KIZ 51836, adult female, share the same collection information as the holotype; KIZ 038604, adult male, KIZ 038603, adult female, collected by Zhong-Bin Yu, Dong An, and Tian-En Chen, from the Dulong Valley (98.50400 ºE, 27.79778 ºN, elevation 3006 m, GCJ 02), collected by Zhongbin Yu, Dong An, and Xian-Kun Huang on 27 July 2021.	en	Wang, Kai, Yu, Zhongbin, Wu, Yunhe, Hou, Shao-Bing, Wang, De, Xiong, Yun, Ye, Xin-Long, Duan, Jian-Ping, Li, Hai-Jun, Li, Jie, Jin, Jie-Qiong, Yang, Shen- Pin, Che, Jing (2025): Three New Species of the Genus Scutiger (Amphibia: Anura: Megophryidae) from the Gaoligong Mountain Range in China, with An Updated Key to the Chinese Congeners. Zootaxa 5725 (2): 203-230, DOI: 10.11646/zootaxa.5725.2.2, URL: https://doi.org/10.11646/zootaxa.5725.2.2
3A4687F46A71FF84FF1CFE4DBBCBF575.taxon	etymology	Etymology: The species name, ching, is derived from the language Trung, the language of the local ethic minority group (Dulong) at the type locality of the new species. It means small and slender, which describes the diagnostic body shape of the new species. We chose the native language of the Dulong People to honor the harmony between the local ethnic culture and nature ecosystem. We suggest Elf Lazy Toad as its English common name, and “ ƜÑÊ ẊḆ ” (Pinyin: Shan Jing Chi Tu Chan) as its Chinese common name.	en	Wang, Kai, Yu, Zhongbin, Wu, Yunhe, Hou, Shao-Bing, Wang, De, Xiong, Yun, Ye, Xin-Long, Duan, Jian-Ping, Li, Hai-Jun, Li, Jie, Jin, Jie-Qiong, Yang, Shen- Pin, Che, Jing (2025): Three New Species of the Genus Scutiger (Amphibia: Anura: Megophryidae) from the Gaoligong Mountain Range in China, with An Updated Key to the Chinese Congeners. Zootaxa 5725 (2): 203-230, DOI: 10.11646/zootaxa.5725.2.2, URL: https://doi.org/10.11646/zootaxa.5725.2.2
3A4687F46A71FF84FF1CFE4DBBCBF575.taxon	diagnosis	Diagnosis: Scutiger ching sp. nov. can be diagnosed from congeners by a combination of the following characteristics: (1) body size small, SVL 37.5 – 45.1 mm; (2) forearm and hand length long, LFHL 50.4 – 54.7 % SVL in males; (3) posterior head short, PHL 40.4 – 43.2 % HL; (4) inter-nare distance large, IND 26.3 – 30.6 % HW; (5) snout long, SEL 36.7 – 43.2 % HL; (6) eyes large, ED 28.8 – 33.1 % HL; (5) single pair of pectoral glands and axillary glands present in males; (6) fine black spines present on pectoral glands, axillary glands, base of ventral forelimbs, peripheral ventral chin, ventral cloaca, and all tubercles on dorsal head, body, and limbs in breeding males; (7) vomerine teeth absence; (8) maxillary teeth fine, presence; (9) tongue slightly notched; (9) nuptial pads present on first three fingers in males, covered with enlarged black spines; (10) distinct enlarged, raised tubercles scattered on dorsal and lateral forearms and anterior brachium; (11) smaller, raised tubercles present on dorsal eyelids and supralabial as well as posterior jaw, covered with dense fine spinules during mating season in males; (12) vocal sacs present in males; (13) supratympanic fold strongly developed, extend posterior of axillary; (14) toes feebly webbed with weak dermal fringes; (15) parietal head flat; and (16) distinct, raised tubercles densely, irregularly scattered on dorsum and dorsal limbs, some large some fine. Comparison: Morphologically, S. ching sp. nov. is most similar to S. tengchongensis, in which both species are closely distributed and have small body size, nuptial spines on the first three fingers, and spinules on dorsal surfaces of head, body, and limbs in males. However, S. ching sp. nov. can be diagnosed readily from S. tengchongensis by the presence of vocal sacs in males (vs. absence), irregularly scattered tubercles on dorsum (vs. regularly aligned as lateral rows), and by having longer hind limbs in males (HLL 148.3 – 156.5 % SVL in males vs. 131.9 – 141.8 %). Additionally, S. ching sp. nov. is similar to S. gongshanensis, in which both species are closely distributed and having similar body size and vocal sacs in males. However, S. ching sp. nov. differs from S. gongshanensis by having nuptial spines on first three fingers in males (vs. only first two fingers) and much longer hind limbs in males (HLL HLL 148.3 – 156.5 % SVL in males vs. 125.5 – 139.0 %). For remaining recognized species of the genus, S. ching sp. nov. differs from most recognized congeners (i. e. S. adungensis, S. biluoensis, S. glandulatus, S. jiulongensis, S. kanjiroba, S. maculatus, S. mammatus, S. meiliensis, S. muliensis, S. nyingchiensis, S. occitentalis, S. pingwuensis, S. spinosus, S. tuberculatus, S. wanglangensis) by having a smaller adult body size (SVL <40 mm in males, <46 mm in female vs.> 55 mm in both sexes). For the remaining congeners with similar body size, S. ching sp. nov. differs from the S. boulengeri complex (including S. boulengeri and S. bangdaensis, as well as the two synonyms of S. boulengeri, namely Cophophryne alticola and Aelurophryne tainingensis), as well as S. bhutanensis, S. chintingensis, S. feiliangi, S. liupanensis, S. luozhaensis, S. ningshanensis, S. sikkimensis, and S. wolong by the presence of vocal sacs in males (vs. absence). For the new species described above, S. ching sp. nov. differs from S. lisu sp. nov. by the presence of vocal sacs in males (vs. absence) and having a much smaller body size (SVL <46 mm vs.> 75 mm); and from S. pardalotus sp. nov. by having more slender forelimbs (thick and more robust), a distinct body coloration (nearly uniform Olive Horn Color [Color 16] with faint, irregular Clay Color [Color 18] patches vs. Pale Pinkish Buff (Color 3) to Light Orange Yellow (Color 7) with Brunt Umber (Color 48) spots and dorsolateral stripes), and much weaker webbing on toes (rudimentary or no webbing vs. half-webbed).	en	Wang, Kai, Yu, Zhongbin, Wu, Yunhe, Hou, Shao-Bing, Wang, De, Xiong, Yun, Ye, Xin-Long, Duan, Jian-Ping, Li, Hai-Jun, Li, Jie, Jin, Jie-Qiong, Yang, Shen- Pin, Che, Jing (2025): Three New Species of the Genus Scutiger (Amphibia: Anura: Megophryidae) from the Gaoligong Mountain Range in China, with An Updated Key to the Chinese Congeners. Zootaxa 5725 (2): 203-230, DOI: 10.11646/zootaxa.5725.2.2, URL: https://doi.org/10.11646/zootaxa.5725.2.2
3A4687F46A71FF84FF1CFE4DBBCBF575.taxon	description	Description of the holotype: Adult male, slim, small, SVL 37.5 mm; head moderate, width about equal to length, HW 98.5 % HL; parietal region flat; snout relatively long, projecting beyond jaw, SEL 40.4 % HL; lateral supralabial convex posterior to nares; nares about equal between eye and snout, facing dorsolaterally, further apart from each other, IN 30.6 % HW; loreal slightly concave; canthus rostralis distinct; eye raised, large, ED 33.1 % HL, DEW 25.0 % HL; iris ellipse shaped, vertically oriented; tympanum concealed; frontal and temporal flat; supratympanic folds well developed, raised, extending posteriorly to posterior jaw only. Limbs slender; forearm long, FAL 54.7 % SVL, slightly thicken toward elbows, FAW 24.4 % FAL; fingers slender, free of webbing, finger length I <II <IV <III; enlarged, raised nuptial spines present dorsally on first two fingers and anterolaterally on third finger; distinctively raised tubercles present on forearm, dorsal hand, and on third and fourth fingers dorsally; metacarpal tubercles distinct, inner larger than outer; subdigital tubercles indistinct. Hindlimb relatively long, heels not meeting anteriorly and reaching posterior eyes with tip of longest toe when adpressed, HLL 156.5 % SVL, FEM 45.6 % SVL, TFL 70.9 % SVL, TIB 40.0 % SVL; foot longer than tibia, FL 126.7 % TIL; toe tips rounded, toe length I <II <V <III <IV; dermal fringes feeble, webbing rudimentary, webbing formula I 2 - – 2 + II 2 – 3 - III 3 – 4 - IV 4 - – 3 - V; subdigital tubercles absent; subarticular tubercles absent, subdigital ridges distinct; inner metatarsal tubercle distinct, oval shaped, longer than first toe, outer metacarpal tubercle absent. Dorsal skins of head, body, and limbs rough, with fine to medium sized tubercles densely scattered, each with one or cluster of black spinules; tubercles densely gathered on dorsal body, but smaller, fewer, and looser toward head and flank. Tubercles on basal branchium dense, irregular, singular and regular on forearm. Spinules present on ventral lower lip; single pair of pectoral glands present on chest, ovel shape, elongated laterally, covered with fine spines; single pair of axillary gland present, distinct, smaller than pectoral, also covered with spines; few patches of spines also present on base of ventral brachium. Ventral cloaca also covered with scattered spinules. Remaining ventral surfaces smooth. Coloration: The background coloration is Pale Horn Color (Color 11) to Olive Horn Color (Color 16). The lateral snout, loreal, and supratympanic folds are Walnut Brown (Color 27). In addition to the Walnut Brown (Color 27) transverse streak between eyes dorsally, these patterns together form a Pale Horn Color (Color 11) triangular patch on dorsal snout. The densely tuberculated region of the dorsum is Antique Brown (Color 24). Indistinct Antique Brown (Color 24) patches or streaks are present on dorsal limb, particularly prominent on dorsal hind limbs. The background coloration of ventral surfaces of head, limbs, and body are Pale Pinkish Buff (Color 3), which are heavily marbled with Flesh Color (Color 249), irregular patterns. Variation: Sexual dimorphism is evident in the new species: in addition to the lack of vocal sacs, pectoral glands, and nuptial spines, the female paratype has more regularly aligned tubercles on dorsum (vs. irregularly scattered), thinner forearms (FAL 15.9 % FAW vs. 21.5 – 24.4 % in males), and shorter hind limbs (HLL 135.7 % SVL vs. 148.3 – 156.5 % in males). The paratype male (KIZ 038604) is not in full breeding mode, and its sexually dimorphic characters are different from the holotype: all dorsal tubercles of the paratype lack black spinules, and the spines of pectoral gland, axillary glands, and dorsal fingers all lack black keratin layer. Webbing formula is consistent among all individuals, without signs of sexual dimorphism. Natural history and conservation: Currently the new species is known from the type locality only, but it is likely to occur in similar alpine habitat further south along the Gaoligong Mountain. The holotype, the paratype female (KIZ 51836), and other uncollected individuals were found gathered in very shallow streams (less than 1 cm deep), and males were calling with a low voice from complex burrows at night. Because the paratype male (KIZ 038604) does not have nuptial spines on pectoral and axillary glands in late July, the breeding season of S. ching sp. nov. is likely between early May and early July. Individuals were extremely shy and stopped calling when disturbed slightly. Although S. lisu sp. nov., Zhangixalus cf. puerensis, and Rana suchinae were found sympatric with the new species, they were found in different microhabitats (larger streams and small rivers for S. lisu sp. nov., deep puddles or slow-flowing pools of streams for Zhangixalus cf. puerensis, and Rana suchinae; Fig. 12 C).	en	Wang, Kai, Yu, Zhongbin, Wu, Yunhe, Hou, Shao-Bing, Wang, De, Xiong, Yun, Ye, Xin-Long, Duan, Jian-Ping, Li, Hai-Jun, Li, Jie, Jin, Jie-Qiong, Yang, Shen- Pin, Che, Jing (2025): Three New Species of the Genus Scutiger (Amphibia: Anura: Megophryidae) from the Gaoligong Mountain Range in China, with An Updated Key to the Chinese Congeners. Zootaxa 5725 (2): 203-230, DOI: 10.11646/zootaxa.5725.2.2, URL: https://doi.org/10.11646/zootaxa.5725.2.2
