taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
49A29F6899385331BB36E82080C2252B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247547	https://doi.org/10.3897/zookeys.1227.138100.figure133	Figure 133. Papuanatula (Papuafiliola) stenophylla sp. nov., larval exuviae (holotype; b – f in one magnification): a enlarged dorsal eye of male b parts of head c abdominal terga and sterna d caudalii e half of pronotum and mesonotum f leg g abdominal segment X (ventral view with paraprocts).	Figure 133. Papuanatula (Papuafiliola) stenophylla sp. nov., larval exuviae (holotype; b – f in one magnification): a enlarged dorsal eye of male b parts of head c abdominal terga and sterna d caudalii e half of pronotum and mesonotum f leg g abdominal segment X (ventral view with paraprocts).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
49A29F6899385331BB36E82080C2252B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247548	https://doi.org/10.3897/zookeys.1227.138100.figure134	Figure 134. Papuanatula (Papuafiliola) stenophylla sp. nov., larva (a – c holotype): a labrum b, c left and right mandibles d, e left and right mandibles with developing mandibles of next instar f maxilla g hypopharynx (lateral view) h, i labium j labial palp (dorsal view).	Figure 134. Papuanatula (Papuafiliola) stenophylla sp. nov., larva (a – c holotype): a labrum b, c left and right mandibles d, e left and right mandibles with developing mandibles of next instar f maxilla g hypopharynx (lateral view) h, i labium j labial palp (dorsal view).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
49A29F6899385331BB36E82080C2252B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247549	https://doi.org/10.3897/zookeys.1227.138100.figure135	Figure 135. Papuanatula (Papuafiliola) stenophylla sp. nov., larval exuviae (holotype): a leg b apex of tibia and base of tarsus (dorsal view) c claw d abdominal tergum VII (dry).	Figure 135. Papuanatula (Papuafiliola) stenophylla sp. nov., larval exuviae (holotype): a leg b apex of tibia and base of tarsus (dorsal view) c claw d abdominal tergum VII (dry).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
49A29F6899385331BB36E82080C2252B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247550	https://doi.org/10.3897/zookeys.1227.138100.figure136	Figure 136. Papuanatula (Papuafiliola) stenophylla sp. nov., larva: a – c fore, middle, and hind legs d tergalius VII in natural position e – j tergalii of II – VII k anal rib of tergalius VII.	Figure 136. Papuanatula (Papuafiliola) stenophylla sp. nov., larva: a – c fore, middle, and hind legs d tergalius VII in natural position e – j tergalii of II – VII k anal rib of tergalius VII.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
49A29F6899385331BB36E82080C2252B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247552	https://doi.org/10.3897/zookeys.1227.138100.figure138	Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva.	Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
49A29F6899385331BB36E82080C2252B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247561	https://doi.org/10.3897/zookeys.1227.138100.figure145	Figure 145. a – e Papuanatula (Papuafiliola) tuberculata sp. nov., eggs (extracted from imago).	Figure 145. a – e Papuanatula (Papuafiliola) tuberculata sp. nov., eggs (extracted from imago).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
2D0E5D311DEE532F87CE60ED436663FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247546	https://doi.org/10.3897/zookeys.1227.138100.figure132	Figure 132. Papuanatula (Papuafiliola) stenophylla sp. nov., larvae.	Figure 132. Papuanatula (Papuafiliola) stenophylla sp. nov., larvae.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
2D0E5D311DEE532F87CE60ED436663FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247547	https://doi.org/10.3897/zookeys.1227.138100.figure133	Figure 133. Papuanatula (Papuafiliola) stenophylla sp. nov., larval exuviae (holotype; b – f in one magnification): a enlarged dorsal eye of male b parts of head c abdominal terga and sterna d caudalii e half of pronotum and mesonotum f leg g abdominal segment X (ventral view with paraprocts).	Figure 133. Papuanatula (Papuafiliola) stenophylla sp. nov., larval exuviae (holotype; b – f in one magnification): a enlarged dorsal eye of male b parts of head c abdominal terga and sterna d caudalii e half of pronotum and mesonotum f leg g abdominal segment X (ventral view with paraprocts).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
2D0E5D311DEE532F87CE60ED436663FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247548	https://doi.org/10.3897/zookeys.1227.138100.figure134	Figure 134. Papuanatula (Papuafiliola) stenophylla sp. nov., larva (a – c holotype): a labrum b, c left and right mandibles d, e left and right mandibles with developing mandibles of next instar f maxilla g hypopharynx (lateral view) h, i labium j labial palp (dorsal view).	Figure 134. Papuanatula (Papuafiliola) stenophylla sp. nov., larva (a – c holotype): a labrum b, c left and right mandibles d, e left and right mandibles with developing mandibles of next instar f maxilla g hypopharynx (lateral view) h, i labium j labial palp (dorsal view).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
2D0E5D311DEE532F87CE60ED436663FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247549	https://doi.org/10.3897/zookeys.1227.138100.figure135	Figure 135. Papuanatula (Papuafiliola) stenophylla sp. nov., larval exuviae (holotype): a leg b apex of tibia and base of tarsus (dorsal view) c claw d abdominal tergum VII (dry).	Figure 135. Papuanatula (Papuafiliola) stenophylla sp. nov., larval exuviae (holotype): a leg b apex of tibia and base of tarsus (dorsal view) c claw d abdominal tergum VII (dry).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
2D0E5D311DEE532F87CE60ED436663FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247550	https://doi.org/10.3897/zookeys.1227.138100.figure136	Figure 136. Papuanatula (Papuafiliola) stenophylla sp. nov., larva: a – c fore, middle, and hind legs d tergalius VII in natural position e – j tergalii of II – VII k anal rib of tergalius VII.	Figure 136. Papuanatula (Papuafiliola) stenophylla sp. nov., larva: a – c fore, middle, and hind legs d tergalius VII in natural position e – j tergalii of II – VII k anal rib of tergalius VII.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
2D0E5D311DEE532F87CE60ED436663FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247551	https://doi.org/10.3897/zookeys.1227.138100.figure137	Figure 137. Papuanatula (Papuafiliola) stenophylla sp. nov., male imago (a – c, f holotype): a head and thorax b abdomen c hind leg d male subimago e subimaginal gonostyli developing under larval cuticle f styliger and gonovectes of male imago.	Figure 137. Papuanatula (Papuafiliola) stenophylla sp. nov., male imago (a – c, f holotype): a head and thorax b abdomen c hind leg d male subimago e subimaginal gonostyli developing under larval cuticle f styliger and gonovectes of male imago.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
2D0E5D311DEE532F87CE60ED436663FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247557	https://doi.org/10.3897/zookeys.1227.138100.figure143	Figure 143. Papuanatula (Papuafiliola) tuberculata sp. nov., subimaginal exuviae (holotype): a half of mesonotum b mesopleuron c hind tarsus d fore tarsus.	Figure 143. Papuanatula (Papuafiliola) tuberculata sp. nov., subimaginal exuviae (holotype): a half of mesonotum b mesopleuron c hind tarsus d fore tarsus.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
2D0E5D311DEE532F87CE60ED436663FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247552	https://doi.org/10.3897/zookeys.1227.138100.figure138	Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva.	Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
2D0E5D311DEE532F87CE60ED436663FC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247567	https://doi.org/10.3897/zookeys.1227.138100.figure148	Figure 148. Distribution of Papuanatula species (x: type locality).	Figure 148. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EA88C84262745D8682259B48D59E7A64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247552	https://doi.org/10.3897/zookeys.1227.138100.figure138	Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva.	Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EA88C84262745D8682259B48D59E7A64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247553	https://doi.org/10.3897/zookeys.1227.138100.figure139	Figure 139. Papuanatula (Papuafiliola) tuberculata sp. nov., larval exuviae (holotype, in one magnification): a head b fore, middle, and hind legs (densely covered with diatom algae) c half of pronotum and mesonotum d abdomen.	Figure 139. Papuanatula (Papuafiliola) tuberculata sp. nov., larval exuviae (holotype, in one magnification): a head b fore, middle, and hind legs (densely covered with diatom algae) c half of pronotum and mesonotum d abdomen.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EA88C84262745D8682259B48D59E7A64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247554	https://doi.org/10.3897/zookeys.1227.138100.figure140	Figure 140. Papuanatula (Papuafiliola) tuberculata sp. nov., larva (a – c, e, h, i holotype): a labrum b, c left and right mandibles d, e maxillae f, g hypopharynx h – j labium (i focus on dorsal side).	Figure 140. Papuanatula (Papuafiliola) tuberculata sp. nov., larva (a – c, e, h, i holotype): a labrum b, c left and right mandibles d, e maxillae f, g hypopharynx h – j labium (i focus on dorsal side).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EA88C84262745D8682259B48D59E7A64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247555	https://doi.org/10.3897/zookeys.1227.138100.figure141	Figure 141. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a leg b tarsus of fore leg (holotype).	Figure 141. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a leg b tarsus of fore leg (holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EA88C84262745D8682259B48D59E7A64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247556	https://doi.org/10.3897/zookeys.1227.138100.figure142	Figure 142. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a tergum III b paraprocts c antenna d tergalius IV e its enlarged anal rib (a – c holotype).	Figure 142. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a tergum III b paraprocts c antenna d tergalius IV e its enlarged anal rib (a – c holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EA88C84262745D8682259B48D59E7A64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247557	https://doi.org/10.3897/zookeys.1227.138100.figure143	Figure 143. Papuanatula (Papuafiliola) tuberculata sp. nov., subimaginal exuviae (holotype): a half of mesonotum b mesopleuron c hind tarsus d fore tarsus.	Figure 143. Papuanatula (Papuafiliola) tuberculata sp. nov., subimaginal exuviae (holotype): a half of mesonotum b mesopleuron c hind tarsus d fore tarsus.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EA88C84262745D8682259B48D59E7A64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247559	https://doi.org/10.3897/zookeys.1227.138100.figure144	Figure 144. Papuanatula (Papuafiliola) tuberculata sp. nov., female imago (holotype): a dorsal view b lateral view c – e fore, middle and hind legs f pterostigma.	Figure 144. Papuanatula (Papuafiliola) tuberculata sp. nov., female imago (holotype): a dorsal view b lateral view c – e fore, middle and hind legs f pterostigma.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EA88C84262745D8682259B48D59E7A64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247561	https://doi.org/10.3897/zookeys.1227.138100.figure145	Figure 145. a – e Papuanatula (Papuafiliola) tuberculata sp. nov., eggs (extracted from imago).	Figure 145. a – e Papuanatula (Papuafiliola) tuberculata sp. nov., eggs (extracted from imago).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EA88C84262745D8682259B48D59E7A64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247450	https://doi.org/10.3897/zookeys.1227.138100.figure36	Figure 36. Papuanatula (Papuanatula) cyclopomontana sp. nov. a head and thorax of male imago b, c fore and middle legs of male imago d subimaginal exuviae of fore leg e subimaginal exuviae of mesopleuron f subimaginal exuviae of mesonotum g abdomen of male imago h, i abdomen of female subimago, dorsally and ventrally (a – g holotype).	Figure 36. Papuanatula (Papuanatula) cyclopomontana sp. nov. a head and thorax of male imago b, c fore and middle legs of male imago d subimaginal exuviae of fore leg e subimaginal exuviae of mesopleuron f subimaginal exuviae of mesonotum g abdomen of male imago h, i abdomen of female subimago, dorsally and ventrally (a – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EA88C84262745D8682259B48D59E7A64.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247567	https://doi.org/10.3897/zookeys.1227.138100.figure148	Figure 148. Distribution of Papuanatula species (x: type locality).	Figure 148. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A8CB813B5D5B586EBAB13E874CF478B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247440	https://doi.org/10.3897/zookeys.1227.138100.figure26	Figure 26. Papuanatula (Papuanatula) balkei sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 26. Papuanatula (Papuanatula) balkei sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A8CB813B5D5B586EBAB13E874CF478B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247441	https://doi.org/10.3897/zookeys.1227.138100.figure27	Figure 27. Papuanatula (Papuanatula) balkei sp. nov., larva: a labrum b labrum: dorsal submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	Figure 27. Papuanatula (Papuanatula) balkei sp. nov., larva: a labrum b labrum: dorsal submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A8CB813B5D5B586EBAB13E874CF478B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247442	https://doi.org/10.3897/zookeys.1227.138100.figure28	Figure 28. Papuanatula (Papuanatula) balkei sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 50 µm (a); 10 µm (b, d); 20 µm (c).	Figure 28. Papuanatula (Papuanatula) balkei sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 50 µm (a); 10 µm (b, d); 20 µm (c).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A8CB813B5D5B586EBAB13E874CF478B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247443	https://doi.org/10.3897/zookeys.1227.138100.figure29	Figure 29. Papuanatula (Papuanatula) balkei sp. nov., larva: a middle leg b middle femur, ventral margin c middle tibia, ventral margin d middle tibia, posterior surface e middle tarsus, ventral margin f middle femur, posterior apex g fore claw (arrow: posterior seta) h male head, mature. Scale bars: 50 µm (a); 10 µm (b – g); 100 µm (h).	Figure 29. Papuanatula (Papuanatula) balkei sp. nov., larva: a middle leg b middle femur, ventral margin c middle tibia, ventral margin d middle tibia, posterior surface e middle tarsus, ventral margin f middle femur, posterior apex g fore claw (arrow: posterior seta) h male head, mature. Scale bars: 50 µm (a); 10 µm (b – g); 100 µm (h).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A8CB813B5D5B586EBAB13E874CF478B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247444	https://doi.org/10.3897/zookeys.1227.138100.figure30	Figure 30. Papuanatula (Papuanatula) balkei sp. nov., larva: a abdominal terga b paracercus c cercus d tergalius IV e tergalius IV, margin f paraproct g cercotractor. Scale bars: 10 µm (a, c, e – g); 20 µm (b, d).	Figure 30. Papuanatula (Papuanatula) balkei sp. nov., larva: a abdominal terga b paracercus c cercus d tergalius IV e tergalius IV, margin f paraproct g cercotractor. Scale bars: 10 µm (a, c, e – g); 20 µm (b, d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A8CB813B5D5B586EBAB13E874CF478B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247445	https://doi.org/10.3897/zookeys.1227.138100.figure31	Figure 31. Papuanatula (Papuanatula) balkei sp. nov., larva (SEM): a abdominal terga II – IV b abdominal terga V – VII c abdominal terga VII – X. Scale bars: 300 µm (a); 200 µm (b, c).	Figure 31. Papuanatula (Papuanatula) balkei sp. nov., larva (SEM): a abdominal terga II – IV b abdominal terga V – VII c abdominal terga VII – X. Scale bars: 300 µm (a); 200 µm (b, c).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A8CB813B5D5B586EBAB13E874CF478B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A8CB813B5D5B586EBAB13E874CF478B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247563	https://doi.org/10.3897/zookeys.1227.138100.figure146	Figure 146. Distribution of Papuanatula species (x: type locality).	Figure 146. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247415	https://doi.org/10.3897/zookeys.1227.138100.figure1	Figure 1. Papuanatula (Papuanatula) bessa, larva, habitus (type locality): a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 1. Papuanatula (Papuanatula) bessa, larva, habitus (type locality): a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247416	https://doi.org/10.3897/zookeys.1227.138100.figure2	Figure 2. Papuanatula (Papuanatula) bessa, larva (from Waena): a male larva b – g exuviae: b head c pronotum and mesonotum d – f fore, middle, and hind legs g abdomen.	Figure 2. Papuanatula (Papuanatula) bessa, larva (from Waena): a male larva b – g exuviae: b head c pronotum and mesonotum d – f fore, middle, and hind legs g abdomen.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247417	https://doi.org/10.3897/zookeys.1227.138100.figure3	Figure 3. Papuanatula (Papuanatula) bessa, larva (from Waena): a, b labrum c labium d hypopharynx e, f left and right mandibles g maxilla.	Figure 3. Papuanatula (Papuanatula) bessa, larva (from Waena): a, b labrum c labium d hypopharynx e, f left and right mandibles g maxilla.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247418	https://doi.org/10.3897/zookeys.1227.138100.figure4	Figure 4. Papuanatula (Papuanatula) bessa, larva (a – e from Wagau, Herzog Mts.): a foreleg b fore claw c hind tibia, inner margin d hind tibia, posterior surface e femur, posterior apex f, g tergalius IV (type locality) h paracercus (type locality) I cercus section (type locality). Scale bars: 50 µm (a); 10 µm (b – d, g, i); 20 µm (f, h).	Figure 4. Papuanatula (Papuanatula) bessa, larva (a – e from Wagau, Herzog Mts.): a foreleg b fore claw c hind tibia, inner margin d hind tibia, posterior surface e femur, posterior apex f, g tergalius IV (type locality) h paracercus (type locality) I cercus section (type locality). Scale bars: 50 µm (a); 10 µm (b – d, g, i); 20 µm (f, h).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247419	https://doi.org/10.3897/zookeys.1227.138100.figure5	Figure 5. Papuanatula (Papuanatula) bessa, larva, abdomen (from Waena): a tergum I b, c tergum II d apex of protuberance on tergum III e tergum IV f tergum V g tergum IX h paraprocts.	Figure 5. Papuanatula (Papuanatula) bessa, larva, abdomen (from Waena): a tergum I b, c tergum II d apex of protuberance on tergum III e tergum IV f tergum V g tergum IX h paraprocts.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247568	https://doi.org/10.3897/zookeys.1227.138100.figure6	Figure 6. Papuanatula (Papuanatula) bessa (from Waena): a, b male subimago with larval exuviae c abdomen of male subimago d subimaginal gonostyli developing under larval cuticle.	Figure 6. Papuanatula (Papuanatula) bessa (from Waena): a, b male subimago with larval exuviae c abdomen of male subimago d subimaginal gonostyli developing under larval cuticle.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247440	https://doi.org/10.3897/zookeys.1227.138100.figure26	Figure 26. Papuanatula (Papuanatula) balkei sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 26. Papuanatula (Papuanatula) balkei sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247479	https://doi.org/10.3897/zookeys.1227.138100.figure65	Figure 65. Papuanatula (Papuanatula) obscura sp. nov., larvae.	Figure 65. Papuanatula (Papuanatula) obscura sp. nov., larvae.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247480	https://doi.org/10.3897/zookeys.1227.138100.figure66	Figure 66. Papuanatula (Papuanatula) obscura sp. nov., larval exuviae: a, b abdomen with caudalii c head d half of pronotum and mesonotum e – g fore, middle, and hind legs h part of head with eye i tergalius III j flagellum of antenna k head with labrum, mandibles, and hypopharynx (a – g in one magnification, k holotype).	Figure 66. Papuanatula (Papuanatula) obscura sp. nov., larval exuviae: a, b abdomen with caudalii c head d half of pronotum and mesonotum e – g fore, middle, and hind legs h part of head with eye i tergalius III j flagellum of antenna k head with labrum, mandibles, and hypopharynx (a – g in one magnification, k holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247455	https://doi.org/10.3897/zookeys.1227.138100.figure41	Figure 41. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a hind leg b hind claw (arrow: posterior seta) c middle tibia, posterior surface d middle tarsus and claw e middle femur, ventral margin f hind tibia, ventral margin g hind femur: setae at dorsal margin h middle femur, posterior apex i male head, mature. Scale bars: 50 µm (a); 10 µm (b, c, e – h); 20 µm (d); 100 µm (i).	Figure 41. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a hind leg b hind claw (arrow: posterior seta) c middle tibia, posterior surface d middle tarsus and claw e middle femur, ventral margin f hind tibia, ventral margin g hind femur: setae at dorsal margin h middle femur, posterior apex i male head, mature. Scale bars: 50 µm (a); 10 µm (b, c, e – h); 20 µm (d); 100 µm (i).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247482	https://doi.org/10.3897/zookeys.1227.138100.figure68	Figure 68. Papuanatula (Papuanatula) obscura sp. nov., larva: a – d middle leg and its parts e tarsus and claw of last instar larva f tarsus and claw of young larva.	Figure 68. Papuanatula (Papuanatula) obscura sp. nov., larva: a – d middle leg and its parts e tarsus and claw of last instar larva f tarsus and claw of young larva.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247484	https://doi.org/10.3897/zookeys.1227.138100.figure70	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
3EF770306D6A5AF7B1A59EAC62EA7461.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247563	https://doi.org/10.3897/zookeys.1227.138100.figure146	Figure 146. Distribution of Papuanatula species (x: type locality).	Figure 146. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A7FE8EA98D5758D89C9B33EA0E6025CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247421	https://doi.org/10.3897/zookeys.1227.138100.figure7	Figure 7. Papuanatula (Papuanatula) copis, larva, habitus (type locality): a dorsal view b lateral view. Scale bars: 1 mm.	Figure 7. Papuanatula (Papuanatula) copis, larva, habitus (type locality): a dorsal view b lateral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A7FE8EA98D5758D89C9B33EA0E6025CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247422	https://doi.org/10.3897/zookeys.1227.138100.figure8	Figure 8. Papuanatula (Papuanatula) copis, larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 8. Papuanatula (Papuanatula) copis, larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A7FE8EA98D5758D89C9B33EA0E6025CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247423	https://doi.org/10.3897/zookeys.1227.138100.figure9	Figure 9. Papuanatula (Papuanatula) copis, larva (type locality): a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola (arrow: subtriangular process). Scale bars: 20 µm (a, c, d, f); 10 µm (b).	Figure 9. Papuanatula (Papuanatula) copis, larva (type locality): a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola (arrow: subtriangular process). Scale bars: 20 µm (a, c, d, f); 10 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A7FE8EA98D5758D89C9B33EA0E6025CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247424	https://doi.org/10.3897/zookeys.1227.138100.figure10	Figure 10. Papuanatula (Papuanatula) copis, larva: a labium (type locality) b labial palp (type locality) c maxilla d maxillary palp e male head, mature. Scale bars: 50 µm (a); 10 µm (b, d); 20 µm (c); 100 µm (e).	Figure 10. Papuanatula (Papuanatula) copis, larva: a labium (type locality) b labial palp (type locality) c maxilla d maxillary palp e male head, mature. Scale bars: 50 µm (a); 10 µm (b, d); 20 µm (c); 100 µm (e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A7FE8EA98D5758D89C9B33EA0E6025CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247425	https://doi.org/10.3897/zookeys.1227.138100.figure11	Figure 11. Papuanatula (Papuanatula) copis, larva: a foreleg (type locality) b middle femur, apex posterior (type locality) c middle femur, ventral margin (type locality) d middle tibia, ventral margin (type locality) e middle tibia, posterior apex (type locality) f middle tarsus, ventral margin g, h middle claw I, j tergalius IV, margin. Scale bars: 50 µm (a); 10 µm (b – e, g, h, j); 20 µm (i).	Figure 11. Papuanatula (Papuanatula) copis, larva: a foreleg (type locality) b middle femur, apex posterior (type locality) c middle femur, ventral margin (type locality) d middle tibia, ventral margin (type locality) e middle tibia, posterior apex (type locality) f middle tarsus, ventral margin g, h middle claw I, j tergalius IV, margin. Scale bars: 50 µm (a); 10 µm (b – e, g, h, j); 20 µm (i).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A7FE8EA98D5758D89C9B33EA0E6025CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247426	https://doi.org/10.3897/zookeys.1227.138100.figure12	Figure 12. Papuanatula (Papuanatula) copis, larva: a metanotum and abdominal terga b abdominal terga c paracercus (type locality) d cercus (type locality) e paraproct (type locality) f cercotractor (type locality) g paraproct, distal margin (type locality). Scale bars: 100 µm (a); 10 µm (b, d – g); 20 µm (c).	Figure 12. Papuanatula (Papuanatula) copis, larva: a metanotum and abdominal terga b abdominal terga c paracercus (type locality) d cercus (type locality) e paraproct (type locality) f cercotractor (type locality) g paraproct, distal margin (type locality). Scale bars: 100 µm (a); 10 µm (b, d – g); 20 µm (c).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A7FE8EA98D5758D89C9B33EA0E6025CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247427	https://doi.org/10.3897/zookeys.1227.138100.figure13	Figure 13. Papuanatula (Papuanatula) copis, larva, SEM: a dorsal view b pronotum. Scale bars: 0.5 mm (a); 300 µm (b).	Figure 13. Papuanatula (Papuanatula) copis, larva, SEM: a dorsal view b pronotum. Scale bars: 0.5 mm (a); 300 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A7FE8EA98D5758D89C9B33EA0E6025CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247563	https://doi.org/10.3897/zookeys.1227.138100.figure146	Figure 146. Distribution of Papuanatula species (x: type locality).	Figure 146. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247446	https://doi.org/10.3897/zookeys.1227.138100.figure32	Figure 32. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae (in one magnification): a abdominal segments I – IX b segment X and caudalii c pronotum and mesonotum d head e – g fore, middle, and hind legs h abdominal segments I – IX (dry) (c – g holotype).	Figure 32. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae (in one magnification): a abdominal segments I – IX b segment X and caudalii c pronotum and mesonotum d head e – g fore, middle, and hind legs h abdominal segments I – IX (dry) (c – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247447	https://doi.org/10.3897/zookeys.1227.138100.figure33	Figure 33. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae (holotype): a labrum b hypopharynx c maxilla d labium.	Figure 33. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae (holotype): a labrum b hypopharynx c maxilla d labium.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247448	https://doi.org/10.3897/zookeys.1227.138100.figure34	Figure 34. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae (holotype): a, b fragment of cercus c apex of femur and base of tibia (middle leg) d apex of tibia and base of tarsus (fore leg) e tarsus and claw (fore leg).	Figure 34. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae (holotype): a, b fragment of cercus c apex of femur and base of tibia (middle leg) d apex of tibia and base of tarsus (fore leg) e tarsus and claw (fore leg).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247449	https://doi.org/10.3897/zookeys.1227.138100.figure35	Figure 35. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae: a abdominal terga b abdominal tergum III c paraprocts (b, c holotype).	Figure 35. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae: a abdominal terga b abdominal tergum III c paraprocts (b, c holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247450	https://doi.org/10.3897/zookeys.1227.138100.figure36	Figure 36. Papuanatula (Papuanatula) cyclopomontana sp. nov. a head and thorax of male imago b, c fore and middle legs of male imago d subimaginal exuviae of fore leg e subimaginal exuviae of mesopleuron f subimaginal exuviae of mesonotum g abdomen of male imago h, i abdomen of female subimago, dorsally and ventrally (a – g holotype).	Figure 36. Papuanatula (Papuanatula) cyclopomontana sp. nov. a head and thorax of male imago b, c fore and middle legs of male imago d subimaginal exuviae of fore leg e subimaginal exuviae of mesopleuron f subimaginal exuviae of mesonotum g abdomen of male imago h, i abdomen of female subimago, dorsally and ventrally (a – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247451	https://doi.org/10.3897/zookeys.1227.138100.figure37	Figure 37. a Papuanatula (Papuanatula) cyclopomontana sp. nov., genitalia (holotype) b – e eggs of subgenus Papuanatula: b Papuanatula (Papuanatula) cyclopomontana sp. nov. (extracted from subimago) c Papuanatula (Papuanatula) obscurella sp. nov. (extracted from imago) d, e Papuanatula (Papuanatula) obscura sp. nov. (extracted from imago).	Figure 37. a Papuanatula (Papuanatula) cyclopomontana sp. nov., genitalia (holotype) b – e eggs of subgenus Papuanatula: b Papuanatula (Papuanatula) cyclopomontana sp. nov. (extracted from subimago) c Papuanatula (Papuanatula) obscurella sp. nov. (extracted from imago) d, e Papuanatula (Papuanatula) obscura sp. nov. (extracted from imago).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247479	https://doi.org/10.3897/zookeys.1227.138100.figure65	Figure 65. Papuanatula (Papuanatula) obscura sp. nov., larvae.	Figure 65. Papuanatula (Papuanatula) obscura sp. nov., larvae.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247455	https://doi.org/10.3897/zookeys.1227.138100.figure41	Figure 41. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a hind leg b hind claw (arrow: posterior seta) c middle tibia, posterior surface d middle tarsus and claw e middle femur, ventral margin f hind tibia, ventral margin g hind femur: setae at dorsal margin h middle femur, posterior apex i male head, mature. Scale bars: 50 µm (a); 10 µm (b, c, e – h); 20 µm (d); 100 µm (i).	Figure 41. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a hind leg b hind claw (arrow: posterior seta) c middle tibia, posterior surface d middle tarsus and claw e middle femur, ventral margin f hind tibia, ventral margin g hind femur: setae at dorsal margin h middle femur, posterior apex i male head, mature. Scale bars: 50 µm (a); 10 µm (b, c, e – h); 20 µm (d); 100 µm (i).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247482	https://doi.org/10.3897/zookeys.1227.138100.figure68	Figure 68. Papuanatula (Papuanatula) obscura sp. nov., larva: a – d middle leg and its parts e tarsus and claw of last instar larva f tarsus and claw of young larva.	Figure 68. Papuanatula (Papuanatula) obscura sp. nov., larva: a – d middle leg and its parts e tarsus and claw of last instar larva f tarsus and claw of young larva.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247474	https://doi.org/10.3897/zookeys.1227.138100.figure60	Figure 60. Papuanatula (Papuanatula) heterochaeta sp. nov. a, b male imagines c abdomen of male imago d female subimago e subimaginal exuviae of mesonotum f subimaginal exuviae of head, prothorax, sterna, and pleura of mesothorax and metathorax g middle leg of male imago (c, e – g holotype).	Figure 60. Papuanatula (Papuanatula) heterochaeta sp. nov. a, b male imagines c abdomen of male imago d female subimago e subimaginal exuviae of mesonotum f subimaginal exuviae of head, prothorax, sterna, and pleura of mesothorax and metathorax g middle leg of male imago (c, e – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247484	https://doi.org/10.3897/zookeys.1227.138100.figure70	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
835CD3DA6248573393280208AFAD25D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247563	https://doi.org/10.3897/zookeys.1227.138100.figure146	Figure 146. Distribution of Papuanatula species (x: type locality).	Figure 146. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
77FD5C6C0E195936AA811122427BE6EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247452	https://doi.org/10.3897/zookeys.1227.138100.figure38	Figure 38. Papuanatula (Papuanatula) dumspinae sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 38. Papuanatula (Papuanatula) dumspinae sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
77FD5C6C0E195936AA811122427BE6EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247453	https://doi.org/10.3897/zookeys.1227.138100.figure39	Figure 39. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a labrum b labrum: dorsal submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola h left mandible: mediolateral setae. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	Figure 39. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a labrum b labrum: dorsal submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola h left mandible: mediolateral setae. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
77FD5C6C0E195936AA811122427BE6EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247454	https://doi.org/10.3897/zookeys.1227.138100.figure40	Figure 40. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 50 µm (a); 20 µm (b, c); 10 µm (d).	Figure 40. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 50 µm (a); 20 µm (b, c); 10 µm (d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
77FD5C6C0E195936AA811122427BE6EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247455	https://doi.org/10.3897/zookeys.1227.138100.figure41	Figure 41. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a hind leg b hind claw (arrow: posterior seta) c middle tibia, posterior surface d middle tarsus and claw e middle femur, ventral margin f hind tibia, ventral margin g hind femur: setae at dorsal margin h middle femur, posterior apex i male head, mature. Scale bars: 50 µm (a); 10 µm (b, c, e – h); 20 µm (d); 100 µm (i).	Figure 41. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a hind leg b hind claw (arrow: posterior seta) c middle tibia, posterior surface d middle tarsus and claw e middle femur, ventral margin f hind tibia, ventral margin g hind femur: setae at dorsal margin h middle femur, posterior apex i male head, mature. Scale bars: 50 µm (a); 10 µm (b, c, e – h); 20 µm (d); 100 µm (i).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
77FD5C6C0E195936AA811122427BE6EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247456	https://doi.org/10.3897/zookeys.1227.138100.figure42	Figure 42. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a abdominal terga b paracercus c, d cercus e, f tergalius IV, margin g paraproct h paraproct: distal margin (arrow: notched scales). Scale bars: 10 µm (a, c, d, f, h); 20 µm (b, g); 50 µm (e).	Figure 42. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a abdominal terga b paracercus c, d cercus e, f tergalius IV, margin g paraproct h paraproct: distal margin (arrow: notched scales). Scale bars: 10 µm (a, c, d, f, h); 20 µm (b, g); 50 µm (e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
77FD5C6C0E195936AA811122427BE6EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247457	https://doi.org/10.3897/zookeys.1227.138100.figure43	Figure 43. Papuanatula (Papuanatula) dumspinae sp. nov., larva (SEM): a abdominal terga III – V b abdominal terga VI – VIII c abdominal tergum III. Scale bars: 200 µm (a, b); 50 µm (c).	Figure 43. Papuanatula (Papuanatula) dumspinae sp. nov., larva (SEM): a abdominal terga III – V b abdominal terga VI – VIII c abdominal tergum III. Scale bars: 200 µm (a, b); 50 µm (c).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
77FD5C6C0E195936AA811122427BE6EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
77FD5C6C0E195936AA811122427BE6EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247563	https://doi.org/10.3897/zookeys.1227.138100.figure146	Figure 146. Distribution of Papuanatula species (x: type locality).	Figure 146. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
499A62E183BC5CA0A698FC1A4219FE9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247458	https://doi.org/10.3897/zookeys.1227.138100.figure44	Figure 44. Papuanatula (Papuanatula) duplex sp. nov., larva, habitus a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 44. Papuanatula (Papuanatula) duplex sp. nov., larva, habitus a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
499A62E183BC5CA0A698FC1A4219FE9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247459	https://doi.org/10.3897/zookeys.1227.138100.figure45	Figure 45. Papuanatula (Papuanatula) duplex sp. nov., larva a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola h labium i labial palp j maxilla k apex of maxillary palp. Scale bars: 20 µm (a, c, d, f, h, j); 10 µm (b, i, k).	Figure 45. Papuanatula (Papuanatula) duplex sp. nov., larva a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola h labium i labial palp j maxilla k apex of maxillary palp. Scale bars: 20 µm (a, c, d, f, h, j); 10 µm (b, i, k).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
499A62E183BC5CA0A698FC1A4219FE9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247460	https://doi.org/10.3897/zookeys.1227.138100.figure46	Figure 46. Papuanatula (Papuanatula) duplex sp. nov., larva: a foreleg b fore tibia, posterior surface c middle claw d abdominal terga VII – IX e abdominal tergum VI, posterior margin f paraproct g tergalius VII. Scale bars: 50 µm (a); 10 µm (b, c, e, f); 20 µm (d, g).	Figure 46. Papuanatula (Papuanatula) duplex sp. nov., larva: a foreleg b fore tibia, posterior surface c middle claw d abdominal terga VII – IX e abdominal tergum VI, posterior margin f paraproct g tergalius VII. Scale bars: 50 µm (a); 10 µm (b, c, e, f); 20 µm (d, g).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
499A62E183BC5CA0A698FC1A4219FE9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247461	https://doi.org/10.3897/zookeys.1227.138100.figure47	Figure 47. Papuanatula (Papuanatula) duplex sp. nov., larva (SEM): a pro-, meso-, metanotum, abdominal terga I – V b metanotum, abdominal terga I and II c pronotum d protuberance on metanotum. Scale bars: 500 µm (a); 200 µm (b); 100 µm (c); 40 µm (d).	Figure 47. Papuanatula (Papuanatula) duplex sp. nov., larva (SEM): a pro-, meso-, metanotum, abdominal terga I – V b metanotum, abdominal terga I and II c pronotum d protuberance on metanotum. Scale bars: 500 µm (a); 200 µm (b); 100 µm (c); 40 µm (d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
499A62E183BC5CA0A698FC1A4219FE9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
499A62E183BC5CA0A698FC1A4219FE9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247563	https://doi.org/10.3897/zookeys.1227.138100.figure146	Figure 146. Distribution of Papuanatula species (x: type locality).	Figure 146. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
106D5D67C34850D18EA366C9811F1CF0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247462	https://doi.org/10.3897/zookeys.1227.138100.figure48	Figure 48. Papuanatula (Papuanatula) epibessa sp. nov., larva, female, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 48. Papuanatula (Papuanatula) epibessa sp. nov., larva, female, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
106D5D67C34850D18EA366C9811F1CF0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247463	https://doi.org/10.3897/zookeys.1227.138100.figure49	Figure 49. Papuanatula (Papuanatula) epibessa sp. nov., larva, male, habitus: a dorsal view. Scale bar: 1 mm.	Figure 49. Papuanatula (Papuanatula) epibessa sp. nov., larva, male, habitus: a dorsal view. Scale bar: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
106D5D67C34850D18EA366C9811F1CF0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247464	https://doi.org/10.3897/zookeys.1227.138100.figure50	Figure 50. Papuanatula (Papuanatula) epibessa sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	Figure 50. Papuanatula (Papuanatula) epibessa sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
106D5D67C34850D18EA366C9811F1CF0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247465	https://doi.org/10.3897/zookeys.1227.138100.figure51	Figure 51. Papuanatula (Papuanatula) epibessa sp. nov., larva: a labium b labial palp c maxilla d maxillary palp e male head, mature. Scale bars: 50 µm (a, c); 20 µm (b); 10 µm (d); 100 µm (e).	Figure 51. Papuanatula (Papuanatula) epibessa sp. nov., larva: a labium b labial palp c maxilla d maxillary palp e male head, mature. Scale bars: 50 µm (a, c); 20 µm (b); 10 µm (d); 100 µm (e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
106D5D67C34850D18EA366C9811F1CF0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247466	https://doi.org/10.3897/zookeys.1227.138100.figure52	Figure 52. Papuanatula (Papuanatula) epibessa sp. nov., larva: a hind leg b hind femur: ventral margin c hind femur: posterior apex d hind tibia: ventral margin e hind tibia: posterior surface f hind tarsus: ventral surface g middle tarsus and claw h middle claw. Scale bars: a 50 µm (a); 10 µm (b – f, h); 20 µm (g).	Figure 52. Papuanatula (Papuanatula) epibessa sp. nov., larva: a hind leg b hind femur: ventral margin c hind femur: posterior apex d hind tibia: ventral margin e hind tibia: posterior surface f hind tarsus: ventral surface g middle tarsus and claw h middle claw. Scale bars: a 50 µm (a); 10 µm (b – f, h); 20 µm (g).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
106D5D67C34850D18EA366C9811F1CF0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247467	https://doi.org/10.3897/zookeys.1227.138100.figure53	Figure 53. Papuanatula (Papuanatula) epibessa sp. nov., larva: a metanotum and abdominal terga I – III b abdominal terga IV – IX c paracercus d cercus e tergalius III f tergalius III, margin g paraproct. Scale bars: 20 µm (a, c, g); 10 µm (b, d, f); 50 µm (e).	Figure 53. Papuanatula (Papuanatula) epibessa sp. nov., larva: a metanotum and abdominal terga I – III b abdominal terga IV – IX c paracercus d cercus e tergalius III f tergalius III, margin g paraproct. Scale bars: 20 µm (a, c, g); 10 µm (b, d, f); 50 µm (e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
106D5D67C34850D18EA366C9811F1CF0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247468	https://doi.org/10.3897/zookeys.1227.138100.figure54	Figure 54. Papuanatula (Papuanatula) epibessa sp. nov., larva (SEM): a abdominal tergum V b abdominal terga VIII – X. Scale bars: 30 µm (a); 100 µm (b).	Figure 54. Papuanatula (Papuanatula) epibessa sp. nov., larva (SEM): a abdominal tergum V b abdominal terga VIII – X. Scale bars: 30 µm (a); 100 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
106D5D67C34850D18EA366C9811F1CF0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
106D5D67C34850D18EA366C9811F1CF0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247563	https://doi.org/10.3897/zookeys.1227.138100.figure146	Figure 146. Distribution of Papuanatula species (x: type locality).	Figure 146. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0F1C9FB320815E0980C3B0BE679108BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247469	https://doi.org/10.3897/zookeys.1227.138100.figure55	Figure 55. Papuanatula (Papuanatula) heterochaeta sp. nov., larva: a dorsal view b, c posterior and anterior sides of fore leg d, e posterior and anterior sides of hind leg f, g hind leg in Canada balsam with focus on anterior and posterior sides.	Figure 55. Papuanatula (Papuanatula) heterochaeta sp. nov., larva: a dorsal view b, c posterior and anterior sides of fore leg d, e posterior and anterior sides of hind leg f, g hind leg in Canada balsam with focus on anterior and posterior sides.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0F1C9FB320815E0980C3B0BE679108BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247470	https://doi.org/10.3897/zookeys.1227.138100.figure56	Figure 56. Papuanatula (Papuanatula) heterochaeta sp. nov., larval cuticle: a head b half of pronotum and mesonotum c abdomen d, e fore, middle, and hind legs (enlarged) (b – f holotype).	Figure 56. Papuanatula (Papuanatula) heterochaeta sp. nov., larval cuticle: a head b half of pronotum and mesonotum c abdomen d, e fore, middle, and hind legs (enlarged) (b – f holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0F1C9FB320815E0980C3B0BE679108BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247471	https://doi.org/10.3897/zookeys.1227.138100.figure57	Figure 57. Papuanatula (Papuanatula) heterochaeta sp. nov., larva: a, b labrum c, d mandibles shortly after molt e, f mandibles before molt g maxilla before molt h, i labium j hypopharynx.	Figure 57. Papuanatula (Papuanatula) heterochaeta sp. nov., larva: a, b labrum c, d mandibles shortly after molt e, f mandibles before molt g maxilla before molt h, i labium j hypopharynx.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0F1C9FB320815E0980C3B0BE679108BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247472	https://doi.org/10.3897/zookeys.1227.138100.figure58	Figure 58. Papuanatula (Papuanatula) heterochaeta sp. nov., larva: a apex of femur b, c inner side of tarsus d outer side of tarsus and tibia e claw f claw and tarsus (b – f holotype).	Figure 58. Papuanatula (Papuanatula) heterochaeta sp. nov., larva: a apex of femur b, c inner side of tarsus d outer side of tarsus and tibia e claw f claw and tarsus (b – f holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0F1C9FB320815E0980C3B0BE679108BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247473	https://doi.org/10.3897/zookeys.1227.138100.figure59	Figure 59. Papuanatula (Papuanatula) heterochaeta sp. nov., larva: a abdominal tergum III (dry) b hind margin of abdominal tergum V c paraprocts and paracercus d – i tergalii II – VII (a – c holotype).	Figure 59. Papuanatula (Papuanatula) heterochaeta sp. nov., larva: a abdominal tergum III (dry) b hind margin of abdominal tergum V c paraprocts and paracercus d – i tergalii II – VII (a – c holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0F1C9FB320815E0980C3B0BE679108BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247474	https://doi.org/10.3897/zookeys.1227.138100.figure60	Figure 60. Papuanatula (Papuanatula) heterochaeta sp. nov. a, b male imagines c abdomen of male imago d female subimago e subimaginal exuviae of mesonotum f subimaginal exuviae of head, prothorax, sterna, and pleura of mesothorax and metathorax g middle leg of male imago (c, e – g holotype).	Figure 60. Papuanatula (Papuanatula) heterochaeta sp. nov. a, b male imagines c abdomen of male imago d female subimago e subimaginal exuviae of mesonotum f subimaginal exuviae of head, prothorax, sterna, and pleura of mesothorax and metathorax g middle leg of male imago (c, e – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0F1C9FB320815E0980C3B0BE679108BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247475	https://doi.org/10.3897/zookeys.1227.138100.figure61	Figure 61. Papuanatula (Papuanatula) heterochaeta sp. nov., male genitalia: a subimaginal genitalia developing under larval cuticle b subimaginal genitalia starting to spread c, d imaginal genitalia (d holotype).	Figure 61. Papuanatula (Papuanatula) heterochaeta sp. nov., male genitalia: a subimaginal genitalia developing under larval cuticle b subimaginal genitalia starting to spread c, d imaginal genitalia (d holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0F1C9FB320815E0980C3B0BE679108BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247446	https://doi.org/10.3897/zookeys.1227.138100.figure32	Figure 32. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae (in one magnification): a abdominal segments I – IX b segment X and caudalii c pronotum and mesonotum d head e – g fore, middle, and hind legs h abdominal segments I – IX (dry) (c – g holotype).	Figure 32. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae (in one magnification): a abdominal segments I – IX b segment X and caudalii c pronotum and mesonotum d head e – g fore, middle, and hind legs h abdominal segments I – IX (dry) (c – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0F1C9FB320815E0980C3B0BE679108BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247484	https://doi.org/10.3897/zookeys.1227.138100.figure70	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0F1C9FB320815E0980C3B0BE679108BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247565	https://doi.org/10.3897/zookeys.1227.138100.figure147	Figure 147. Distribution of Papuanatula species (x: type locality).	Figure 147. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F7F3649175985A59BCB1B831EB479A93.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247416	https://doi.org/10.3897/zookeys.1227.138100.figure2	Figure 2. Papuanatula (Papuanatula) bessa, larva (from Waena): a male larva b – g exuviae: b head c pronotum and mesonotum d – f fore, middle, and hind legs g abdomen.	Figure 2. Papuanatula (Papuanatula) bessa, larva (from Waena): a male larva b – g exuviae: b head c pronotum and mesonotum d – f fore, middle, and hind legs g abdomen.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F7F3649175985A59BCB1B831EB479A93.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247417	https://doi.org/10.3897/zookeys.1227.138100.figure3	Figure 3. Papuanatula (Papuanatula) bessa, larva (from Waena): a, b labrum c labium d hypopharynx e, f left and right mandibles g maxilla.	Figure 3. Papuanatula (Papuanatula) bessa, larva (from Waena): a, b labrum c labium d hypopharynx e, f left and right mandibles g maxilla.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F7F3649175985A59BCB1B831EB479A93.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247418	https://doi.org/10.3897/zookeys.1227.138100.figure4	Figure 4. Papuanatula (Papuanatula) bessa, larva (a – e from Wagau, Herzog Mts.): a foreleg b fore claw c hind tibia, inner margin d hind tibia, posterior surface e femur, posterior apex f, g tergalius IV (type locality) h paracercus (type locality) I cercus section (type locality). Scale bars: 50 µm (a); 10 µm (b – d, g, i); 20 µm (f, h).	Figure 4. Papuanatula (Papuanatula) bessa, larva (a – e from Wagau, Herzog Mts.): a foreleg b fore claw c hind tibia, inner margin d hind tibia, posterior surface e femur, posterior apex f, g tergalius IV (type locality) h paracercus (type locality) I cercus section (type locality). Scale bars: 50 µm (a); 10 µm (b – d, g, i); 20 µm (f, h).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F7F3649175985A59BCB1B831EB479A93.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F7F3649175985A59BCB1B831EB479A93.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247426	https://doi.org/10.3897/zookeys.1227.138100.figure12	Figure 12. Papuanatula (Papuanatula) copis, larva: a metanotum and abdominal terga b abdominal terga c paracercus (type locality) d cercus (type locality) e paraproct (type locality) f cercotractor (type locality) g paraproct, distal margin (type locality). Scale bars: 100 µm (a); 10 µm (b, d – g); 20 µm (c).	Figure 12. Papuanatula (Papuanatula) copis, larva: a metanotum and abdominal terga b abdominal terga c paracercus (type locality) d cercus (type locality) e paraproct (type locality) f cercotractor (type locality) g paraproct, distal margin (type locality). Scale bars: 100 µm (a); 10 µm (b, d – g); 20 µm (c).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F7F3649175985A59BCB1B831EB479A93.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247490	https://doi.org/10.3897/zookeys.1227.138100.figure76	Figure 76. Papuanatula (Papuanatula) obscurella sp. nov., male genitalia: a genitalia of imago b subimaginal genitalia developing under larval cuticle (a holotype).	Figure 76. Papuanatula (Papuanatula) obscurella sp. nov., male genitalia: a genitalia of imago b subimaginal genitalia developing under larval cuticle (a holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F7F3649175985A59BCB1B831EB479A93.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247451	https://doi.org/10.3897/zookeys.1227.138100.figure37	Figure 37. a Papuanatula (Papuanatula) cyclopomontana sp. nov., genitalia (holotype) b – e eggs of subgenus Papuanatula: b Papuanatula (Papuanatula) cyclopomontana sp. nov. (extracted from subimago) c Papuanatula (Papuanatula) obscurella sp. nov. (extracted from imago) d, e Papuanatula (Papuanatula) obscura sp. nov. (extracted from imago).	Figure 37. a Papuanatula (Papuanatula) cyclopomontana sp. nov., genitalia (holotype) b – e eggs of subgenus Papuanatula: b Papuanatula (Papuanatula) cyclopomontana sp. nov. (extracted from subimago) c Papuanatula (Papuanatula) obscurella sp. nov. (extracted from imago) d, e Papuanatula (Papuanatula) obscura sp. nov. (extracted from imago).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
1621D2C3529B5AFD9CA65A062B5BA7FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247428	https://doi.org/10.3897/zookeys.1227.138100.figure14	Figure 14. Papuanatula (Papuanatula) lenos, larva, habitus (type locality): a dorsal view. Scale bar: 1 mm.	Figure 14. Papuanatula (Papuanatula) lenos, larva, habitus (type locality): a dorsal view. Scale bar: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
1621D2C3529B5AFD9CA65A062B5BA7FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247429	https://doi.org/10.3897/zookeys.1227.138100.figure15	Figure 15. Papuanatula (Papuanatula) lenos, larva (type locality): a labrum b hypopharynx and superlinguae c right mandible d right mandible: margin between prostheca and mola e left mandible f left mandible: margin between prostheca and mola g labial palp h maxilla i maxillary palp. Scale bars: 20 µm (a – c, e, h); 10 µm (g, i).	Figure 15. Papuanatula (Papuanatula) lenos, larva (type locality): a labrum b hypopharynx and superlinguae c right mandible d right mandible: margin between prostheca and mola e left mandible f left mandible: margin between prostheca and mola g labial palp h maxilla i maxillary palp. Scale bars: 20 µm (a – c, e, h); 10 µm (g, i).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
1621D2C3529B5AFD9CA65A062B5BA7FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247430	https://doi.org/10.3897/zookeys.1227.138100.figure16	Figure 16. Papuanatula (Papuanatula) lenos, larva (type locality): a middle tibia, posterior surface b fore femur, ventral margin c abdominal terga d paracercus e tergalius II f tergalius IV, margin. Scale bars: 10 µm (a – c, f); 20 µm (d, e).	Figure 16. Papuanatula (Papuanatula) lenos, larva (type locality): a middle tibia, posterior surface b fore femur, ventral margin c abdominal terga d paracercus e tergalius II f tergalius IV, margin. Scale bars: 10 µm (a – c, f); 20 µm (d, e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
1621D2C3529B5AFD9CA65A062B5BA7FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247563	https://doi.org/10.3897/zookeys.1227.138100.figure146	Figure 146. Distribution of Papuanatula species (x: type locality).	Figure 146. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
44836D371795592F96323CCBE3EAB69A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247476	https://doi.org/10.3897/zookeys.1227.138100.figure62	Figure 62. Papuanatula (Papuanatula) normungulata sp. nov., larva (holotype): a head b maxilla c hypopharynx d labium.	Figure 62. Papuanatula (Papuanatula) normungulata sp. nov., larva (holotype): a head b maxilla c hypopharynx d labium.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
44836D371795592F96323CCBE3EAB69A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247477	https://doi.org/10.3897/zookeys.1227.138100.figure63	Figure 63. Papuanatula (Papuanatula) normungulata sp. nov., larva (holotype): larva a pronotum and mesonotum b abdomen c median spine on abdominal tergum VIII d metanotum and abdominal tergum I e abdominal tergum IX f tergalius II h abdominal tergum X and paracercus.	Figure 63. Papuanatula (Papuanatula) normungulata sp. nov., larva (holotype): larva a pronotum and mesonotum b abdomen c median spine on abdominal tergum VIII d metanotum and abdominal tergum I e abdominal tergum IX f tergalius II h abdominal tergum X and paracercus.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
44836D371795592F96323CCBE3EAB69A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247478	https://doi.org/10.3897/zookeys.1227.138100.figure64	Figure 64. Papuanatula (Papuanatula) normungulata sp. nov., larva (holotype): larva a, b fore and middle legs c apex of femur d outer side of tibia e tarsus and claw f outer side of femur.	Figure 64. Papuanatula (Papuanatula) normungulata sp. nov., larva (holotype): larva a, b fore and middle legs c apex of femur d outer side of tibia e tarsus and claw f outer side of femur.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
44836D371795592F96323CCBE3EAB69A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
44836D371795592F96323CCBE3EAB69A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247484	https://doi.org/10.3897/zookeys.1227.138100.figure70	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
44836D371795592F96323CCBE3EAB69A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247565	https://doi.org/10.3897/zookeys.1227.138100.figure147	Figure 147. Distribution of Papuanatula species (x: type locality).	Figure 147. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247479	https://doi.org/10.3897/zookeys.1227.138100.figure65	Figure 65. Papuanatula (Papuanatula) obscura sp. nov., larvae.	Figure 65. Papuanatula (Papuanatula) obscura sp. nov., larvae.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247480	https://doi.org/10.3897/zookeys.1227.138100.figure66	Figure 66. Papuanatula (Papuanatula) obscura sp. nov., larval exuviae: a, b abdomen with caudalii c head d half of pronotum and mesonotum e – g fore, middle, and hind legs h part of head with eye i tergalius III j flagellum of antenna k head with labrum, mandibles, and hypopharynx (a – g in one magnification, k holotype).	Figure 66. Papuanatula (Papuanatula) obscura sp. nov., larval exuviae: a, b abdomen with caudalii c head d half of pronotum and mesonotum e – g fore, middle, and hind legs h part of head with eye i tergalius III j flagellum of antenna k head with labrum, mandibles, and hypopharynx (a – g in one magnification, k holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247481	https://doi.org/10.3897/zookeys.1227.138100.figure67	Figure 67. Papuanatula (Papuanatula) obscura sp. nov., larva: a labrum b maxilla and hypopharynx c, d left and right mandibles e – g labium.	Figure 67. Papuanatula (Papuanatula) obscura sp. nov., larva: a labrum b maxilla and hypopharynx c, d left and right mandibles e – g labium.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247482	https://doi.org/10.3897/zookeys.1227.138100.figure68	Figure 68. Papuanatula (Papuanatula) obscura sp. nov., larva: a – d middle leg and its parts e tarsus and claw of last instar larva f tarsus and claw of young larva.	Figure 68. Papuanatula (Papuanatula) obscura sp. nov., larva: a – d middle leg and its parts e tarsus and claw of last instar larva f tarsus and claw of young larva.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247483	https://doi.org/10.3897/zookeys.1227.138100.figure69	Figure 69. Papuanatula (Papuanatula) obscura sp. nov., larval exuviae: a paraprocts b, c abdominal tergum IV d posterior margin of tergum VII.	Figure 69. Papuanatula (Papuanatula) obscura sp. nov., larval exuviae: a paraprocts b, c abdominal tergum IV d posterior margin of tergum VII.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247484	https://doi.org/10.3897/zookeys.1227.138100.figure70	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247485	https://doi.org/10.3897/zookeys.1227.138100.figure71	Figure 71. Papuanatula (Papuanatula) obscura sp. nov., male genitalia: a subimaginal genitalia developing under larval cuticle b, c imaginal genitalia (c holotype).	Figure 71. Papuanatula (Papuanatula) obscura sp. nov., male genitalia: a subimaginal genitalia developing under larval cuticle b, c imaginal genitalia (c holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247474	https://doi.org/10.3897/zookeys.1227.138100.figure60	Figure 60. Papuanatula (Papuanatula) heterochaeta sp. nov. a, b male imagines c abdomen of male imago d female subimago e subimaginal exuviae of mesonotum f subimaginal exuviae of head, prothorax, sterna, and pleura of mesothorax and metathorax g middle leg of male imago (c, e – g holotype).	Figure 60. Papuanatula (Papuanatula) heterochaeta sp. nov. a, b male imagines c abdomen of male imago d female subimago e subimaginal exuviae of mesonotum f subimaginal exuviae of head, prothorax, sterna, and pleura of mesothorax and metathorax g middle leg of male imago (c, e – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247450	https://doi.org/10.3897/zookeys.1227.138100.figure36	Figure 36. Papuanatula (Papuanatula) cyclopomontana sp. nov. a head and thorax of male imago b, c fore and middle legs of male imago d subimaginal exuviae of fore leg e subimaginal exuviae of mesopleuron f subimaginal exuviae of mesonotum g abdomen of male imago h, i abdomen of female subimago, dorsally and ventrally (a – g holotype).	Figure 36. Papuanatula (Papuanatula) cyclopomontana sp. nov. a head and thorax of male imago b, c fore and middle legs of male imago d subimaginal exuviae of fore leg e subimaginal exuviae of mesopleuron f subimaginal exuviae of mesonotum g abdomen of male imago h, i abdomen of female subimago, dorsally and ventrally (a – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247451	https://doi.org/10.3897/zookeys.1227.138100.figure37	Figure 37. a Papuanatula (Papuanatula) cyclopomontana sp. nov., genitalia (holotype) b – e eggs of subgenus Papuanatula: b Papuanatula (Papuanatula) cyclopomontana sp. nov. (extracted from subimago) c Papuanatula (Papuanatula) obscurella sp. nov. (extracted from imago) d, e Papuanatula (Papuanatula) obscura sp. nov. (extracted from imago).	Figure 37. a Papuanatula (Papuanatula) cyclopomontana sp. nov., genitalia (holotype) b – e eggs of subgenus Papuanatula: b Papuanatula (Papuanatula) cyclopomontana sp. nov. (extracted from subimago) c Papuanatula (Papuanatula) obscurella sp. nov. (extracted from imago) d, e Papuanatula (Papuanatula) obscura sp. nov. (extracted from imago).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4D22BC4BCBD5ACA8A6742D9317FD67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247565	https://doi.org/10.3897/zookeys.1227.138100.figure147	Figure 147. Distribution of Papuanatula species (x: type locality).	Figure 147. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247486	https://doi.org/10.3897/zookeys.1227.138100.figure72	Figure 72. Papuanatula (Papuanatula) obscurella sp. nov., larva: a – f larval exuviae (holotype) a abdomen b – d fore, middle, and hind legs e head f half of pronotum and mesonotum g male larva h, i female and male larvae with hypodermal subimaginal coloration of abdomen visible through larval cuticle.	Figure 72. Papuanatula (Papuanatula) obscurella sp. nov., larva: a – f larval exuviae (holotype) a abdomen b – d fore, middle, and hind legs e head f half of pronotum and mesonotum g male larva h, i female and male larvae with hypodermal subimaginal coloration of abdomen visible through larval cuticle.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247487	https://doi.org/10.3897/zookeys.1227.138100.figure73	Figure 73. Papuanatula (Papuanatula) obscurella sp. nov., larva: a, b left and right mandibles of penultimate instar with mandibles of last instar developing inside c, d left and right mandibles with complete incisors e labrum f hypopharynx with superlinguae g labium h maxilla (g holotype).	Figure 73. Papuanatula (Papuanatula) obscurella sp. nov., larva: a, b left and right mandibles of penultimate instar with mandibles of last instar developing inside c, d left and right mandibles with complete incisors e labrum f hypopharynx with superlinguae g labium h maxilla (g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247488	https://doi.org/10.3897/zookeys.1227.138100.figure74	Figure 74. Papuanatula (Papuanatula) obscurella sp. nov., larva: a fore tarsus and claw b hind leg c tergum IX d paraprocts (c holotype).	Figure 74. Papuanatula (Papuanatula) obscurella sp. nov., larva: a fore tarsus and claw b hind leg c tergum IX d paraprocts (c holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247489	https://doi.org/10.3897/zookeys.1227.138100.figure75	Figure 75. Papuanatula (Papuanatula) obscurella sp. nov.: a, b head and thorax of female imago dorsally and ventrally c abdomen of male imago d, e male imago f pterostigma g subimaginal exuviae of half of mesonotum h subimaginal exuviae of mesopleuron I, j fore and hind legs (c – f, I, j holotype).	Figure 75. Papuanatula (Papuanatula) obscurella sp. nov.: a, b head and thorax of female imago dorsally and ventrally c abdomen of male imago d, e male imago f pterostigma g subimaginal exuviae of half of mesonotum h subimaginal exuviae of mesopleuron I, j fore and hind legs (c – f, I, j holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247490	https://doi.org/10.3897/zookeys.1227.138100.figure76	Figure 76. Papuanatula (Papuanatula) obscurella sp. nov., male genitalia: a genitalia of imago b subimaginal genitalia developing under larval cuticle (a holotype).	Figure 76. Papuanatula (Papuanatula) obscurella sp. nov., male genitalia: a genitalia of imago b subimaginal genitalia developing under larval cuticle (a holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247455	https://doi.org/10.3897/zookeys.1227.138100.figure41	Figure 41. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a hind leg b hind claw (arrow: posterior seta) c middle tibia, posterior surface d middle tarsus and claw e middle femur, ventral margin f hind tibia, ventral margin g hind femur: setae at dorsal margin h middle femur, posterior apex i male head, mature. Scale bars: 50 µm (a); 10 µm (b, c, e – h); 20 µm (d); 100 µm (i).	Figure 41. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a hind leg b hind claw (arrow: posterior seta) c middle tibia, posterior surface d middle tarsus and claw e middle femur, ventral margin f hind tibia, ventral margin g hind femur: setae at dorsal margin h middle femur, posterior apex i male head, mature. Scale bars: 50 µm (a); 10 µm (b, c, e – h); 20 µm (d); 100 µm (i).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247482	https://doi.org/10.3897/zookeys.1227.138100.figure68	Figure 68. Papuanatula (Papuanatula) obscura sp. nov., larva: a – d middle leg and its parts e tarsus and claw of last instar larva f tarsus and claw of young larva.	Figure 68. Papuanatula (Papuanatula) obscura sp. nov., larva: a – d middle leg and its parts e tarsus and claw of last instar larva f tarsus and claw of young larva.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247480	https://doi.org/10.3897/zookeys.1227.138100.figure66	Figure 66. Papuanatula (Papuanatula) obscura sp. nov., larval exuviae: a, b abdomen with caudalii c head d half of pronotum and mesonotum e – g fore, middle, and hind legs h part of head with eye i tergalius III j flagellum of antenna k head with labrum, mandibles, and hypopharynx (a – g in one magnification, k holotype).	Figure 66. Papuanatula (Papuanatula) obscura sp. nov., larval exuviae: a, b abdomen with caudalii c head d half of pronotum and mesonotum e – g fore, middle, and hind legs h part of head with eye i tergalius III j flagellum of antenna k head with labrum, mandibles, and hypopharynx (a – g in one magnification, k holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247474	https://doi.org/10.3897/zookeys.1227.138100.figure60	Figure 60. Papuanatula (Papuanatula) heterochaeta sp. nov. a, b male imagines c abdomen of male imago d female subimago e subimaginal exuviae of mesonotum f subimaginal exuviae of head, prothorax, sterna, and pleura of mesothorax and metathorax g middle leg of male imago (c, e – g holotype).	Figure 60. Papuanatula (Papuanatula) heterochaeta sp. nov. a, b male imagines c abdomen of male imago d female subimago e subimaginal exuviae of mesonotum f subimaginal exuviae of head, prothorax, sterna, and pleura of mesothorax and metathorax g middle leg of male imago (c, e – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247450	https://doi.org/10.3897/zookeys.1227.138100.figure36	Figure 36. Papuanatula (Papuanatula) cyclopomontana sp. nov. a head and thorax of male imago b, c fore and middle legs of male imago d subimaginal exuviae of fore leg e subimaginal exuviae of mesopleuron f subimaginal exuviae of mesonotum g abdomen of male imago h, i abdomen of female subimago, dorsally and ventrally (a – g holotype).	Figure 36. Papuanatula (Papuanatula) cyclopomontana sp. nov. a head and thorax of male imago b, c fore and middle legs of male imago d subimaginal exuviae of fore leg e subimaginal exuviae of mesopleuron f subimaginal exuviae of mesonotum g abdomen of male imago h, i abdomen of female subimago, dorsally and ventrally (a – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247484	https://doi.org/10.3897/zookeys.1227.138100.figure70	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247451	https://doi.org/10.3897/zookeys.1227.138100.figure37	Figure 37. a Papuanatula (Papuanatula) cyclopomontana sp. nov., genitalia (holotype) b – e eggs of subgenus Papuanatula: b Papuanatula (Papuanatula) cyclopomontana sp. nov. (extracted from subimago) c Papuanatula (Papuanatula) obscurella sp. nov. (extracted from imago) d, e Papuanatula (Papuanatula) obscura sp. nov. (extracted from imago).	Figure 37. a Papuanatula (Papuanatula) cyclopomontana sp. nov., genitalia (holotype) b – e eggs of subgenus Papuanatula: b Papuanatula (Papuanatula) cyclopomontana sp. nov. (extracted from subimago) c Papuanatula (Papuanatula) obscurella sp. nov. (extracted from imago) d, e Papuanatula (Papuanatula) obscura sp. nov. (extracted from imago).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
A4617B4D05B95EF280888D8E2A29A34B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247565	https://doi.org/10.3897/zookeys.1227.138100.figure147	Figure 147. Distribution of Papuanatula species (x: type locality).	Figure 147. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
D638F3D2F20A5E3886580FBAF1717051.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247491	https://doi.org/10.3897/zookeys.1227.138100.figure77	Figure 77. Papuanatula (Papuanatula) parabessa sp. nov., larva, habitus: a dorsal view b lateral view c dorsolateral view (arrows: paired humps) d ventral view. Scale bars: 1 mm.	Figure 77. Papuanatula (Papuanatula) parabessa sp. nov., larva, habitus: a dorsal view b lateral view c dorsolateral view (arrows: paired humps) d ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
D638F3D2F20A5E3886580FBAF1717051.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247492	https://doi.org/10.3897/zookeys.1227.138100.figure78	Figure 78. Papuanatula (Papuanatula) parabessa sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	Figure 78. Papuanatula (Papuanatula) parabessa sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
D638F3D2F20A5E3886580FBAF1717051.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247493	https://doi.org/10.3897/zookeys.1227.138100.figure79	Figure 79. Papuanatula (Papuanatula) parabessa sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 50 µm (a, c); 10 µm (b, d).	Figure 79. Papuanatula (Papuanatula) parabessa sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 50 µm (a, c); 10 µm (b, d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
D638F3D2F20A5E3886580FBAF1717051.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247494	https://doi.org/10.3897/zookeys.1227.138100.figure80	Figure 80. Papuanatula (Papuanatula) parabessa sp. nov., larva: a hind leg b hind femur: ventral margin c hind femur: posterior apex d hind tibia, ventral margin e hind tarsus: ventral margin f hind tibia: posterior surface g fore claw h male head, mature i developing gonostyli. Scale bars: 50 µm (a); 10 µm (b – g, i); 100 µm (h).	Figure 80. Papuanatula (Papuanatula) parabessa sp. nov., larva: a hind leg b hind femur: ventral margin c hind femur: posterior apex d hind tibia, ventral margin e hind tarsus: ventral margin f hind tibia: posterior surface g fore claw h male head, mature i developing gonostyli. Scale bars: 50 µm (a); 10 µm (b – g, i); 100 µm (h).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
D638F3D2F20A5E3886580FBAF1717051.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247495	https://doi.org/10.3897/zookeys.1227.138100.figure81	Figure 81. Papuanatula (Papuanatula) parabessa sp. nov., larva: a metanotum and abdominal terga b abdominal terga c paracercus d cercus e tergalius IV f tergalius IV, margin. Scale bars: 20 µm (a, c); 10 µm (b, d, f); 50 µm (e).	Figure 81. Papuanatula (Papuanatula) parabessa sp. nov., larva: a metanotum and abdominal terga b abdominal terga c paracercus d cercus e tergalius IV f tergalius IV, margin. Scale bars: 20 µm (a, c); 10 µm (b, d, f); 50 µm (e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
D638F3D2F20A5E3886580FBAF1717051.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247496	https://doi.org/10.3897/zookeys.1227.138100.figure82	Figure 82. Papuanatula (Papuanatula) parabessa sp. nov., larva: a paraproct b cercotractor c paraproct, distal margin d abdominal terga IV – VI (SEM; arrows: paired humps) e abdominal terga (SEM). Scale bars: 20 µm (a); 10 µm (b, c); 200 µm (d); 50 µm (e).	Figure 82. Papuanatula (Papuanatula) parabessa sp. nov., larva: a paraproct b cercotractor c paraproct, distal margin d abdominal terga IV – VI (SEM; arrows: paired humps) e abdominal terga (SEM). Scale bars: 20 µm (a); 10 µm (b, c); 200 µm (d); 50 µm (e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
D638F3D2F20A5E3886580FBAF1717051.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
D638F3D2F20A5E3886580FBAF1717051.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247502	https://doi.org/10.3897/zookeys.1227.138100.figure88	Figure 88. Papuanatula (Papuanatula) paracopis sp. nov., larva: a metanotum and abdominal terga I – V b abdominal terga VI – IX c abdominal terga d paracercus e, f cercus g paraproct. Scale bars: 100 µm (a, b); 10 µm (c, e, f); 20 µm (d, g).	Figure 88. Papuanatula (Papuanatula) paracopis sp. nov., larva: a metanotum and abdominal terga I – V b abdominal terga VI – IX c abdominal terga d paracercus e, f cercus g paraproct. Scale bars: 100 µm (a, b); 10 µm (c, e, f); 20 µm (d, g).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
D638F3D2F20A5E3886580FBAF1717051.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247565	https://doi.org/10.3897/zookeys.1227.138100.figure147	Figure 147. Distribution of Papuanatula species (x: type locality).	Figure 147. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
5D94675EDBEE563F91DF3A37F5829D8D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247497	https://doi.org/10.3897/zookeys.1227.138100.figure83	Figure 83. Papuanatula (Papuanatula) paracopis sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 83. Papuanatula (Papuanatula) paracopis sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
5D94675EDBEE563F91DF3A37F5829D8D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247498	https://doi.org/10.3897/zookeys.1227.138100.figure84	Figure 84. Papuanatula (Papuanatula) paracopis sp. nov., immature larva, habitus: a dorsal view b lateral view. Scale bars: 1 mm.	Figure 84. Papuanatula (Papuanatula) paracopis sp. nov., immature larva, habitus: a dorsal view b lateral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
5D94675EDBEE563F91DF3A37F5829D8D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247499	https://doi.org/10.3897/zookeys.1227.138100.figure85	Figure 85. Papuanatula (Papuanatula) paracopis sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola h left mandible: mediolateral setae. Scale bars: 20 µm (a, c); 10 µm (b); 50 µm (d, f).	Figure 85. Papuanatula (Papuanatula) paracopis sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola h left mandible: mediolateral setae. Scale bars: 20 µm (a, c); 10 µm (b); 50 µm (d, f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
5D94675EDBEE563F91DF3A37F5829D8D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247500	https://doi.org/10.3897/zookeys.1227.138100.figure86	Figure 86. Papuanatula (Papuanatula) paracopis sp. nov., larva: a labium b labial palp c maxilla d male head, mature. Scale bars: 50 µm (a, c); 20 µm (b); 100 µm (d).	Figure 86. Papuanatula (Papuanatula) paracopis sp. nov., larva: a labium b labial palp c maxilla d male head, mature. Scale bars: 50 µm (a, c); 20 µm (b); 100 µm (d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
5D94675EDBEE563F91DF3A37F5829D8D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247501	https://doi.org/10.3897/zookeys.1227.138100.figure87	Figure 87. Papuanatula (Papuanatula) paracopis sp. nov., larva: a middle leg b middle tarsus and claw c middle tibia, posterior surface d middle femur, ventral margin e middle tibia, ventral margin f tergalius II g tergalius II, margin. Scale bars: 50 µm (a); 10 µm (b – f, h); 20 µm (g).	Figure 87. Papuanatula (Papuanatula) paracopis sp. nov., larva: a middle leg b middle tarsus and claw c middle tibia, posterior surface d middle femur, ventral margin e middle tibia, ventral margin f tergalius II g tergalius II, margin. Scale bars: 50 µm (a); 10 µm (b – f, h); 20 µm (g).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
5D94675EDBEE563F91DF3A37F5829D8D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247502	https://doi.org/10.3897/zookeys.1227.138100.figure88	Figure 88. Papuanatula (Papuanatula) paracopis sp. nov., larva: a metanotum and abdominal terga I – V b abdominal terga VI – IX c abdominal terga d paracercus e, f cercus g paraproct. Scale bars: 100 µm (a, b); 10 µm (c, e, f); 20 µm (d, g).	Figure 88. Papuanatula (Papuanatula) paracopis sp. nov., larva: a metanotum and abdominal terga I – V b abdominal terga VI – IX c abdominal terga d paracercus e, f cercus g paraproct. Scale bars: 100 µm (a, b); 10 µm (c, e, f); 20 µm (d, g).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
5D94675EDBEE563F91DF3A37F5829D8D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247503	https://doi.org/10.3897/zookeys.1227.138100.figure89	Figure 89. Papuanatula (Papuanatula) paracopis sp. nov., larva (SEM): a mesonotum, metanotum, abdominal terga I – IX b metanotum, abdominal terga I – II c abdominal terga III – V d abdominal terga VII – IX e abdominal tergum VI. Scale bars: 500 µm (a); 200 µm (b); 300 µm (c); 100 µm (d); 30 µm (e).	Figure 89. Papuanatula (Papuanatula) paracopis sp. nov., larva (SEM): a mesonotum, metanotum, abdominal terga I – IX b metanotum, abdominal terga I – II c abdominal terga III – V d abdominal terga VII – IX e abdominal tergum VI. Scale bars: 500 µm (a); 200 µm (b); 300 µm (c); 100 µm (d); 30 µm (e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
5D94675EDBEE563F91DF3A37F5829D8D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247565	https://doi.org/10.3897/zookeys.1227.138100.figure147	Figure 147. Distribution of Papuanatula species (x: type locality).	Figure 147. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
7506516145BE5F8DA017EF837EBC172B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247504	https://doi.org/10.3897/zookeys.1227.138100.figure90	Figure 90. Papuanatula (Papuanatula) paralenos sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 90. Papuanatula (Papuanatula) paralenos sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
7506516145BE5F8DA017EF837EBC172B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247505	https://doi.org/10.3897/zookeys.1227.138100.figure91	Figure 91. Papuanatula (Papuanatula) paralenos sp. nov., larva, habitus: a dorsal view. Scale bar: 1 mm.	Figure 91. Papuanatula (Papuanatula) paralenos sp. nov., larva, habitus: a dorsal view. Scale bar: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
7506516145BE5F8DA017EF837EBC172B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247506	https://doi.org/10.3897/zookeys.1227.138100.figure92	Figure 92. Papuanatula (Papuanatula) paralenos sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	Figure 92. Papuanatula (Papuanatula) paralenos sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
7506516145BE5F8DA017EF837EBC172B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247507	https://doi.org/10.3897/zookeys.1227.138100.figure93	Figure 93. Papuanatula (Papuanatula) paralenos sp. nov., larva: a labium b labial palp c maxilla d maxillary palp e male head, mature f tergalius IV g tergalius IV, margin. Scale bars: 50 µm (a, c); 10 µm (b, d, g); 100 µm (e); 20 µm (f).	Figure 93. Papuanatula (Papuanatula) paralenos sp. nov., larva: a labium b labial palp c maxilla d maxillary palp e male head, mature f tergalius IV g tergalius IV, margin. Scale bars: 50 µm (a, c); 10 µm (b, d, g); 100 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
7506516145BE5F8DA017EF837EBC172B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247508	https://doi.org/10.3897/zookeys.1227.138100.figure94	Figure 94. Papuanatula (Papuanatula) paralenos sp. nov., larva: a fore leg b fore femur: ventral margin c fore femur: posterior apex d fore tibia: ventral margin e hind tibia: posterior surface f fore tarsus: ventral margin g fore claw h fore claw (arrow: posterior seta) i developing gonostyli. Scale bars: 50 µm (a); 10 µm (b – h); 20 µm (i).	Figure 94. Papuanatula (Papuanatula) paralenos sp. nov., larva: a fore leg b fore femur: ventral margin c fore femur: posterior apex d fore tibia: ventral margin e hind tibia: posterior surface f fore tarsus: ventral margin g fore claw h fore claw (arrow: posterior seta) i developing gonostyli. Scale bars: 50 µm (a); 10 µm (b – h); 20 µm (i).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
7506516145BE5F8DA017EF837EBC172B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247509	https://doi.org/10.3897/zookeys.1227.138100.figure95	Figure 95. Papuanatula (Papuanatula) paralenos sp. nov., larva: a – c abdominal terga d paracercus e cercus f paraproct. Scale bars: 10 µm (a – c, e, f); 20 µm (d).	Figure 95. Papuanatula (Papuanatula) paralenos sp. nov., larva: a – c abdominal terga d paracercus e cercus f paraproct. Scale bars: 10 µm (a – c, e, f); 20 µm (d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
7506516145BE5F8DA017EF837EBC172B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247510	https://doi.org/10.3897/zookeys.1227.138100.figure96	Figure 96. Papuanatula (Papuanatula) paralenos sp. nov., larva (SEM): a abdominal terga I – IV b abdominal terga V – VII c abdominal terga VII – IX d abdominal tergum V e abdominal tergum VII f labrum: dorsal submarginal setae. Scale bars: 300 µm (a, b); 100 µm (c); 40 µm (d, e); 10 µm (f).	Figure 96. Papuanatula (Papuanatula) paralenos sp. nov., larva (SEM): a abdominal terga I – IV b abdominal terga V – VII c abdominal terga VII – IX d abdominal tergum V e abdominal tergum VII f labrum: dorsal submarginal setae. Scale bars: 300 µm (a, b); 100 µm (c); 40 µm (d, e); 10 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
7506516145BE5F8DA017EF837EBC172B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247565	https://doi.org/10.3897/zookeys.1227.138100.figure147	Figure 147. Distribution of Papuanatula species (x: type locality).	Figure 147. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
69934095D95F52A181A8F295840862F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247511	https://doi.org/10.3897/zookeys.1227.138100.figure97	Figure 97. Papuanatula (Papuanatula) paratuber sp. nov., larva, habitus: a dorsal view, female b dorsal view, male c lateral view, male d ventral view, male. Scale bars: 1 mm.	Figure 97. Papuanatula (Papuanatula) paratuber sp. nov., larva, habitus: a dorsal view, female b dorsal view, male c lateral view, male d ventral view, male. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
69934095D95F52A181A8F295840862F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247512	https://doi.org/10.3897/zookeys.1227.138100.figure98	Figure 98. Papuanatula (Papuanatula) paratuber sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	Figure 98. Papuanatula (Papuanatula) paratuber sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
69934095D95F52A181A8F295840862F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247513	https://doi.org/10.3897/zookeys.1227.138100.figure99	Figure 99. Papuanatula (Papuanatula) paratuber sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 20 µm (a, c); 10 µm (b, d).	Figure 99. Papuanatula (Papuanatula) paratuber sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 20 µm (a, c); 10 µm (b, d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
69934095D95F52A181A8F295840862F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247514	https://doi.org/10.3897/zookeys.1227.138100.figure100	Figure 100. Papuanatula (Papuanatula) paratuber sp. nov., larva: a hind leg b hind femur, ventral margin c hind tibia: ventral margin d hind tibia: posterior surface e middle claw f cercus g paracercus h male head, mature. Scale bars: 50 µm (a); 10 µm (b – f); 20 µm (g); 100 µm (h).	Figure 100. Papuanatula (Papuanatula) paratuber sp. nov., larva: a hind leg b hind femur, ventral margin c hind tibia: ventral margin d hind tibia: posterior surface e middle claw f cercus g paracercus h male head, mature. Scale bars: 50 µm (a); 10 µm (b – f); 20 µm (g); 100 µm (h).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
69934095D95F52A181A8F295840862F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247515	https://doi.org/10.3897/zookeys.1227.138100.figure101	Figure 101. Papuanatula (Papuanatula) paratuber sp. nov., larva: a metanotum, abdominal terga I – IX b abdominal terga c tergalius IV d tergalius IV, margin e paraproct. Scale bars: 100 µm (a); 10 µm (b, d, e); 20 µm (c).	Figure 101. Papuanatula (Papuanatula) paratuber sp. nov., larva: a metanotum, abdominal terga I – IX b abdominal terga c tergalius IV d tergalius IV, margin e paraproct. Scale bars: 100 µm (a); 10 µm (b, d, e); 20 µm (c).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
69934095D95F52A181A8F295840862F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247516	https://doi.org/10.3897/zookeys.1227.138100.figure102	Figure 102. Papuanatula (Papuanatula) paratuber sp. nov., larva (SEM): a metanotum, abdominal terga I – III b abdominal terga IV – VII c abdominal terga VII – X d pronotum e abdominal tergum VII. Scale bars: 400 µm (a); 300 µm (b – d); 50 µm (e).	Figure 102. Papuanatula (Papuanatula) paratuber sp. nov., larva (SEM): a metanotum, abdominal terga I – III b abdominal terga IV – VII c abdominal terga VII – X d pronotum e abdominal tergum VII. Scale bars: 400 µm (a); 300 µm (b – d); 50 µm (e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
69934095D95F52A181A8F295840862F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
69934095D95F52A181A8F295840862F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247565	https://doi.org/10.3897/zookeys.1227.138100.figure147	Figure 147. Distribution of Papuanatula species (x: type locality).	Figure 147. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
43ABA01CE8F55DD4A468D4334657B92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247517	https://doi.org/10.3897/zookeys.1227.138100.figure103	Figure 103. Papuanatula (Papuanatula) parvatubera sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 103. Papuanatula (Papuanatula) parvatubera sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
43ABA01CE8F55DD4A468D4334657B92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247518	https://doi.org/10.3897/zookeys.1227.138100.figure104	Figure 104. Papuanatula (Papuanatula) parvatubera sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 10 µm (a – c); 20 µm (d, f).	Figure 104. Papuanatula (Papuanatula) parvatubera sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 10 µm (a – c); 20 µm (d, f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
43ABA01CE8F55DD4A468D4334657B92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247519	https://doi.org/10.3897/zookeys.1227.138100.figure105	Figure 105. Papuanatula (Papuanatula) parvatubera sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 20 µm (a, c); 10 µm (b, d).	Figure 105. Papuanatula (Papuanatula) parvatubera sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 20 µm (a, c); 10 µm (b, d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
43ABA01CE8F55DD4A468D4334657B92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247520	https://doi.org/10.3897/zookeys.1227.138100.figure106	Figure 106. Papuanatula (Papuanatula) parvatubera sp. nov., larva: a hind leg b middle tarsus and claw (arrow: posterior seta) c male head, mature d prosternum e paracercus f cercus, dorsal view g cercus, lateral view. Scale bars: 50 µm (a); 10 µm (b, f, g); 100 µm (c); 20 µm (d, e).	Figure 106. Papuanatula (Papuanatula) parvatubera sp. nov., larva: a hind leg b middle tarsus and claw (arrow: posterior seta) c male head, mature d prosternum e paracercus f cercus, dorsal view g cercus, lateral view. Scale bars: 50 µm (a); 10 µm (b, f, g); 100 µm (c); 20 µm (d, e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
43ABA01CE8F55DD4A468D4334657B92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247521	https://doi.org/10.3897/zookeys.1227.138100.figure107	Figure 107. Papuanatula (Papuanatula) parvatubera sp. nov., larva: a abdominal terga III – V b abdominal tergum V (arrows: minute, acute spines) c paraproct d tergalius IV e developing gonostyli. Scale bars: 20 µm (a); 10 µm (b – e).	Figure 107. Papuanatula (Papuanatula) parvatubera sp. nov., larva: a abdominal terga III – V b abdominal tergum V (arrows: minute, acute spines) c paraproct d tergalius IV e developing gonostyli. Scale bars: 20 µm (a); 10 µm (b – e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
43ABA01CE8F55DD4A468D4334657B92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
43ABA01CE8F55DD4A468D4334657B92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247565	https://doi.org/10.3897/zookeys.1227.138100.figure147	Figure 147. Distribution of Papuanatula species (x: type locality).	Figure 147. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
BBCFDF5234BD518EA41075D89BEB1557.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247523	https://doi.org/10.3897/zookeys.1227.138100.figure109	Figure 109. Papuanatula (Papuanatula) pilosa sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 109. Papuanatula (Papuanatula) pilosa sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
BBCFDF5234BD518EA41075D89BEB1557.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247524	https://doi.org/10.3897/zookeys.1227.138100.figure110	Figure 110. Papuanatula (Papuanatula) pilosa sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	Figure 110. Papuanatula (Papuanatula) pilosa sp. nov., larva: a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
BBCFDF5234BD518EA41075D89BEB1557.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247525	https://doi.org/10.3897/zookeys.1227.138100.figure111	Figure 111. Papuanatula (Papuanatula) pilosa sp. nov., larva: a labium b labial palp c maxilla d maxillary palp segment III e paraproct f tergalius IV g tergalius IV: margin. Scale bars: 20 µm (a, c); 10 µm (b, d, e, g); 50 µm (f).	Figure 111. Papuanatula (Papuanatula) pilosa sp. nov., larva: a labium b labial palp c maxilla d maxillary palp segment III e paraproct f tergalius IV g tergalius IV: margin. Scale bars: 20 µm (a, c); 10 µm (b, d, e, g); 50 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
BBCFDF5234BD518EA41075D89BEB1557.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247526	https://doi.org/10.3897/zookeys.1227.138100.figure112	Figure 112. Papuanatula (Papuanatula) pilosa sp. nov., larva: a middle leg b middle claw c middle tibia: outer margin d middle tibia: inner margin e middle femur: inner margin f paracercus g, h cercus. Scale bars: 50 µm (a); 10 µm (b – e, g, h); 20 µm (f).	Figure 112. Papuanatula (Papuanatula) pilosa sp. nov., larva: a middle leg b middle claw c middle tibia: outer margin d middle tibia: inner margin e middle femur: inner margin f paracercus g, h cercus. Scale bars: 50 µm (a); 10 µm (b – e, g, h); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
BBCFDF5234BD518EA41075D89BEB1557.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247527	https://doi.org/10.3897/zookeys.1227.138100.figure113	Figure 113. Papuanatula (Papuanatula) pilosa sp. nov., larva (SEM): a abdominal terga III – IX b pronotum c abdominal terga III – VI d abdominal terga VII – IX e abdominal terga VIII – IX f abdominal tergum IX, posterior margin. Scale bars: 400 µm (a); 300 µm (b, c); 100 µm (d); 50 µm (e); 20 µm (f).	Figure 113. Papuanatula (Papuanatula) pilosa sp. nov., larva (SEM): a abdominal terga III – IX b pronotum c abdominal terga III – VI d abdominal terga VII – IX e abdominal terga VIII – IX f abdominal tergum IX, posterior margin. Scale bars: 400 µm (a); 300 µm (b, c); 100 µm (d); 50 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
BBCFDF5234BD518EA41075D89BEB1557.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
BBCFDF5234BD518EA41075D89BEB1557.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247565	https://doi.org/10.3897/zookeys.1227.138100.figure147	Figure 147. Distribution of Papuanatula species (x: type locality).	Figure 147. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EF608808121A551A880CA064B7D9A843.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247431	https://doi.org/10.3897/zookeys.1227.138100.figure17	Figure 17. Papuanatula (Papuanatula) plana, larva, habitus (type locality): a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 17. Papuanatula (Papuanatula) plana, larva, habitus (type locality): a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EF608808121A551A880CA064B7D9A843.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247432	https://doi.org/10.3897/zookeys.1227.138100.figure18	Figure 18. Papuanatula (Papuanatula) plana, larva (type locality): a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 10 µm (a, b); 20 µm (c, d, f).	Figure 18. Papuanatula (Papuanatula) plana, larva (type locality): a labrum b labrum: dorsal, submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 10 µm (a, b); 20 µm (c, d, f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EF608808121A551A880CA064B7D9A843.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247433	https://doi.org/10.3897/zookeys.1227.138100.figure19	Figure 19. Papuanatula (Papuanatula) plana, larva (type locality): a labium b labial palp c maxilla d maxillary palp. Scale bars: 20 µm (a, c); 10 µm (b, d).	Figure 19. Papuanatula (Papuanatula) plana, larva (type locality): a labium b labial palp c maxilla d maxillary palp. Scale bars: 20 µm (a, c); 10 µm (b, d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EF608808121A551A880CA064B7D9A843.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247434	https://doi.org/10.3897/zookeys.1227.138100.figure20	Figure 20. Papuanatula (Papuanatula) plana, larva (type locality): a foreleg b fore tibia, ventral margin c fore tarsus, ventral margin d fore claw (arrow: posterior seta) e male head, mature. Scale bars: 50 µm (a, e); 10 µm (b, c, d).	Figure 20. Papuanatula (Papuanatula) plana, larva (type locality): a foreleg b fore tibia, ventral margin c fore tarsus, ventral margin d fore claw (arrow: posterior seta) e male head, mature. Scale bars: 50 µm (a, e); 10 µm (b, c, d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EF608808121A551A880CA064B7D9A843.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247435	https://doi.org/10.3897/zookeys.1227.138100.figure21	Figure 21. Papuanatula (Papuanatula) plana, larva (type locality): a abdominal terga b paracercus c cercus d paraproct e tergalius V f developing gonostyli. Scale bars: 10 µm (a, c – e); 20 µm (b, f).	Figure 21. Papuanatula (Papuanatula) plana, larva (type locality): a abdominal terga b paracercus c cercus d paraproct e tergalius V f developing gonostyli. Scale bars: 10 µm (a, c – e); 20 µm (b, f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EF608808121A551A880CA064B7D9A843.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247436	https://doi.org/10.3897/zookeys.1227.138100.figure22	Figure 22. Papuanatula (Papuanatula) plana, larva, SEM: a, b abdominal terga c abdominal tergum VI d abdominal tergum VI. Scale bars: 100 µm (a, b); 40 µm (c); 10 µm (d).	Figure 22. Papuanatula (Papuanatula) plana, larva, SEM: a, b abdominal terga c abdominal tergum VI d abdominal tergum VI. Scale bars: 100 µm (a, b); 40 µm (c); 10 µm (d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EF608808121A551A880CA064B7D9A843.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247437	https://doi.org/10.3897/zookeys.1227.138100.figure23	Figure 23. Papuanatula (Papuanatula) plana, male imago, habitus (type locality): a dorsal view b lateral view. Scale bars: 1 mm.	Figure 23. Papuanatula (Papuanatula) plana, male imago, habitus (type locality): a dorsal view b lateral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EF608808121A551A880CA064B7D9A843.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247438	https://doi.org/10.3897/zookeys.1227.138100.figure24	Figure 24. Papuanatula (Papuanatula) plana, male imago (type locality): a fore wing b abdomen c genitalia. Scale bars: 100 µm (a, b); 20 µm (c).	Figure 24. Papuanatula (Papuanatula) plana, male imago (type locality): a fore wing b abdomen c genitalia. Scale bars: 100 µm (a, b); 20 µm (c).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EF608808121A551A880CA064B7D9A843.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EF608808121A551A880CA064B7D9A843.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247563	https://doi.org/10.3897/zookeys.1227.138100.figure146	Figure 146. Distribution of Papuanatula species (x: type locality).	Figure 146. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EDD2250FCAF45D02B9FDA7198F527BE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247528	https://doi.org/10.3897/zookeys.1227.138100.figure114	Figure 114. Papuanatula (Papuanatula) pluresetae sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 114. Papuanatula (Papuanatula) pluresetae sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EDD2250FCAF45D02B9FDA7198F527BE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247529	https://doi.org/10.3897/zookeys.1227.138100.figure115	Figure 115. Papuanatula (Papuanatula) pluresetae sp. nov., larva: a labrum b labrum: dorsal submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	Figure 115. Papuanatula (Papuanatula) pluresetae sp. nov., larva: a labrum b labrum: dorsal submarginal setae c hypopharynx and superlinguae d right mandible e right mandible: margin between prostheca and mola f left mandible g left mandible: margin between prostheca and mola. Scale bars: 20 µm (a, c, d, f); 10 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EDD2250FCAF45D02B9FDA7198F527BE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247530	https://doi.org/10.3897/zookeys.1227.138100.figure116	Figure 116. Papuanatula (Papuanatula) pluresetae sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 50 µm (a); 20 µm (b, c); 10 µm (d).	Figure 116. Papuanatula (Papuanatula) pluresetae sp. nov., larva: a labium b labial palp c maxilla d maxillary palp. Scale bars: 50 µm (a); 20 µm (b, c); 10 µm (d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EDD2250FCAF45D02B9FDA7198F527BE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247531	https://doi.org/10.3897/zookeys.1227.138100.figure117	Figure 117. Papuanatula (Papuanatula) pluresetae sp. nov., larva: a fore leg b fore claw (arrow: posterior setae) c fore tarsus and claw d middle femur, posterior apex e male head, mature f paracercus g cercus. Scale bars: 50 µm (a); 10 µm (b – g); 100 µm (h).	Figure 117. Papuanatula (Papuanatula) pluresetae sp. nov., larva: a fore leg b fore claw (arrow: posterior setae) c fore tarsus and claw d middle femur, posterior apex e male head, mature f paracercus g cercus. Scale bars: 50 µm (a); 10 µm (b – g); 100 µm (h).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EDD2250FCAF45D02B9FDA7198F527BE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247532	https://doi.org/10.3897/zookeys.1227.138100.figure118	Figure 118. Papuanatula (Papuanatula) pluresetae sp. nov., larva: a tergalius IV b tergalius IV, margin c paraproct d cercotractor e section of paraproct with notched scales f paraproct, distal margin. Scale bars: 50 µm (a); 10 µm (b, d); 20 µm (c).	Figure 118. Papuanatula (Papuanatula) pluresetae sp. nov., larva: a tergalius IV b tergalius IV, margin c paraproct d cercotractor e section of paraproct with notched scales f paraproct, distal margin. Scale bars: 50 µm (a); 10 µm (b, d); 20 µm (c).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EDD2250FCAF45D02B9FDA7198F527BE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247533	https://doi.org/10.3897/zookeys.1227.138100.figure119	Figure 119. Papuanatula (Papuanatula) pluresetae sp. nov., larva: a abdominal terga. Scale bar: 10 µm (a).	Figure 119. Papuanatula (Papuanatula) pluresetae sp. nov., larva: a abdominal terga. Scale bar: 10 µm (a).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EDD2250FCAF45D02B9FDA7198F527BE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247534	https://doi.org/10.3897/zookeys.1227.138100.figure120	Figure 120. Papuanatula (Papuanatula) pluresetae sp. nov., larva (SEM): a pro- and mesonotum b abdominal terga II – IV c abdominal terga IV – VI d abdominal terga VI – VIII. Scale bars: 200 µm.	Figure 120. Papuanatula (Papuanatula) pluresetae sp. nov., larva (SEM): a pro- and mesonotum b abdominal terga II – IV c abdominal terga IV – VI d abdominal terga VI – VIII. Scale bars: 200 µm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EDD2250FCAF45D02B9FDA7198F527BE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247440	https://doi.org/10.3897/zookeys.1227.138100.figure26	Figure 26. Papuanatula (Papuanatula) balkei sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 26. Papuanatula (Papuanatula) balkei sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EDD2250FCAF45D02B9FDA7198F527BE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
EDD2250FCAF45D02B9FDA7198F527BE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247567	https://doi.org/10.3897/zookeys.1227.138100.figure148	Figure 148. Distribution of Papuanatula species (x: type locality).	Figure 148. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4E3297889355B8F9D0BCA0E10E01329.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247439	https://doi.org/10.3897/zookeys.1227.138100.figure25	Figure 25. Papuanatula (Papuanatula) tuber, larva, habitus (holotype): a lateral view b male head, mature. Scale bar: 1 mm; (photographs C. C. Wirth, Purdue University).	Figure 25. Papuanatula (Papuanatula) tuber, larva, habitus (holotype): a lateral view b male head, mature. Scale bar: 1 mm; (photographs C. C. Wirth, Purdue University).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
F4E3297889355B8F9D0BCA0E10E01329.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247563	https://doi.org/10.3897/zookeys.1227.138100.figure146	Figure 146. Distribution of Papuanatula species (x: type locality).	Figure 146. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0C5C2BE9C0A756DF9D30E0C0C10D840B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247535	https://doi.org/10.3897/zookeys.1227.138100.figure121	Figure 121. Papuanatula (Papuanatula) webbi sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 121. Papuanatula (Papuanatula) webbi sp. nov., larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0C5C2BE9C0A756DF9D30E0C0C10D840B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247536	https://doi.org/10.3897/zookeys.1227.138100.figure122	Figure 122. Papuanatula (Papuanatula) webbi sp. nov., larva: a labrum b hypopharynx and superlinguae c right mandible d right mandible: margin between prostheca and mola e left mandible f left mandible: margin between prostheca and mola. Scale bars: 10 µm (a, b); 20 µm (c, e).	Figure 122. Papuanatula (Papuanatula) webbi sp. nov., larva: a labrum b hypopharynx and superlinguae c right mandible d right mandible: margin between prostheca and mola e left mandible f left mandible: margin between prostheca and mola. Scale bars: 10 µm (a, b); 20 µm (c, e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0C5C2BE9C0A756DF9D30E0C0C10D840B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247537	https://doi.org/10.3897/zookeys.1227.138100.figure123	Figure 123. Papuanatula (Papuanatula) webbi sp. nov., larva: a labium b labial palp c maxilla d maxillary palp e male head, mature. Scale bars: 20 µm (a, c); 10 µm (b, d); 100 µm (e).	Figure 123. Papuanatula (Papuanatula) webbi sp. nov., larva: a labium b labial palp c maxilla d maxillary palp e male head, mature. Scale bars: 20 µm (a, c); 10 µm (b, d); 100 µm (e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0C5C2BE9C0A756DF9D30E0C0C10D840B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247538	https://doi.org/10.3897/zookeys.1227.138100.figure124	Figure 124. Papuanatula (Papuanatula) webbi sp. nov., larva: a hind leg b hind tarsus and claw (arrow: posterior seta) c hind femur, ventral margin d hind tibia, ventral margin e hind tibia, posterior surface f pronotum g metanotum with rudimentary hind protopteron h fore protopteron i fore protopteron protuberance. Scale bars: 50 µm (a, h); 10 µm (b – g, i).	Figure 124. Papuanatula (Papuanatula) webbi sp. nov., larva: a hind leg b hind tarsus and claw (arrow: posterior seta) c hind femur, ventral margin d hind tibia, ventral margin e hind tibia, posterior surface f pronotum g metanotum with rudimentary hind protopteron h fore protopteron i fore protopteron protuberance. Scale bars: 50 µm (a, h); 10 µm (b – g, i).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0C5C2BE9C0A756DF9D30E0C0C10D840B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247539	https://doi.org/10.3897/zookeys.1227.138100.figure125	Figure 125. Papuanatula (Papuanatula) webbi sp. nov., larva: a abdominal terga I – IX b abdominal terga VI – VII (arrows: minute, acute spines) c paracercus d cercus e paraproct f tergalius III. Scale bars: 50 µm (a); 10 µm (b, d – f); 20 µm (c).	Figure 125. Papuanatula (Papuanatula) webbi sp. nov., larva: a abdominal terga I – IX b abdominal terga VI – VII (arrows: minute, acute spines) c paracercus d cercus e paraproct f tergalius III. Scale bars: 50 µm (a); 10 µm (b, d – f); 20 µm (c).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0C5C2BE9C0A756DF9D30E0C0C10D840B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247540	https://doi.org/10.3897/zookeys.1227.138100.figure126	Figure 126. Papuanatula (Papuanatula) webbi sp. nov., larva (SEM): a pronotum b abdominal terga VII – X c abdominal tergum VII, posterior margin d spine on posterior margin of abdominal tergum IX. Scale bars: 100 µm (a, b); 20 µm (c); 5 µm (d).	Figure 126. Papuanatula (Papuanatula) webbi sp. nov., larva (SEM): a pronotum b abdominal terga VII – X c abdominal tergum VII, posterior margin d spine on posterior margin of abdominal tergum IX. Scale bars: 100 µm (a, b); 20 µm (c); 5 µm (d).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0C5C2BE9C0A756DF9D30E0C0C10D840B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
0C5C2BE9C0A756DF9D30E0C0C10D840B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247567	https://doi.org/10.3897/zookeys.1227.138100.figure148	Figure 148. Distribution of Papuanatula species (x: type locality).	Figure 148. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247541	https://doi.org/10.3897/zookeys.1227.138100.figure127	Figure 127. Papuanatula (Papuanatula) zebrata sp. nov.: a male larva b cuticle of larval abdomen c abdomen of male subimago extracted from larva d larval abdomen with hypodermal coloration (b, c holotype).	Figure 127. Papuanatula (Papuanatula) zebrata sp. nov.: a male larva b cuticle of larval abdomen c abdomen of male subimago extracted from larva d larval abdomen with hypodermal coloration (b, c holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247542	https://doi.org/10.3897/zookeys.1227.138100.figure128	Figure 128. Papuanatula (Papuanatula) zebrata sp. nov., larva: a labrum b, c labium d – f left and right mandibles g maxilla h hypopharynx with superlinguae (a, b holotype).	Figure 128. Papuanatula (Papuanatula) zebrata sp. nov., larva: a labrum b, c labium d – f left and right mandibles g maxilla h hypopharynx with superlinguae (a, b holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247543	https://doi.org/10.3897/zookeys.1227.138100.figure129	Figure 129. Papuanatula (Papuanatula) zebrata sp. nov., larva: a setae on inner side of middle tarsus b tarsus and claw c – e fore, middle, and hind legs with developing subimaginal legs inside f fore leg (a, c – e holotype).	Figure 129. Papuanatula (Papuanatula) zebrata sp. nov., larva: a setae on inner side of middle tarsus b tarsus and claw c – e fore, middle, and hind legs with developing subimaginal legs inside f fore leg (a, c – e holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247544	https://doi.org/10.3897/zookeys.1227.138100.figure130	Figure 130. Papuanatula (Papuanatula) zebrata sp. nov., larva: a antenna b abdominal tergum V c abdominal terga IX – X (b, c holotype).	Figure 130. Papuanatula (Papuanatula) zebrata sp. nov., larva: a antenna b abdominal tergum V c abdominal terga IX – X (b, c holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247545	https://doi.org/10.3897/zookeys.1227.138100.figure131	Figure 131. Papuanatula (Papuanatula) zebrata sp. nov., subimagines: a cuticle of female subimaginal mesonotum b male subimaginal genitalia developing under larval exuviae (b holotype).	Figure 131. Papuanatula (Papuanatula) zebrata sp. nov., subimagines: a cuticle of female subimaginal mesonotum b male subimaginal genitalia developing under larval exuviae (b holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247446	https://doi.org/10.3897/zookeys.1227.138100.figure32	Figure 32. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae (in one magnification): a abdominal segments I – IX b segment X and caudalii c pronotum and mesonotum d head e – g fore, middle, and hind legs h abdominal segments I – IX (dry) (c – g holotype).	Figure 32. Papuanatula (Papuanatula) cyclopomontana sp. nov., larval exuviae (in one magnification): a abdominal segments I – IX b segment X and caudalii c pronotum and mesonotum d head e – g fore, middle, and hind legs h abdominal segments I – IX (dry) (c – g holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247455	https://doi.org/10.3897/zookeys.1227.138100.figure41	Figure 41. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a hind leg b hind claw (arrow: posterior seta) c middle tibia, posterior surface d middle tarsus and claw e middle femur, ventral margin f hind tibia, ventral margin g hind femur: setae at dorsal margin h middle femur, posterior apex i male head, mature. Scale bars: 50 µm (a); 10 µm (b, c, e – h); 20 µm (d); 100 µm (i).	Figure 41. Papuanatula (Papuanatula) dumspinae sp. nov., larva: a hind leg b hind claw (arrow: posterior seta) c middle tibia, posterior surface d middle tarsus and claw e middle femur, ventral margin f hind tibia, ventral margin g hind femur: setae at dorsal margin h middle femur, posterior apex i male head, mature. Scale bars: 50 µm (a); 10 µm (b, c, e – h); 20 µm (d); 100 µm (i).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247482	https://doi.org/10.3897/zookeys.1227.138100.figure68	Figure 68. Papuanatula (Papuanatula) obscura sp. nov., larva: a – d middle leg and its parts e tarsus and claw of last instar larva f tarsus and claw of young larva.	Figure 68. Papuanatula (Papuanatula) obscura sp. nov., larva: a – d middle leg and its parts e tarsus and claw of last instar larva f tarsus and claw of young larva.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247480	https://doi.org/10.3897/zookeys.1227.138100.figure66	Figure 66. Papuanatula (Papuanatula) obscura sp. nov., larval exuviae: a, b abdomen with caudalii c head d half of pronotum and mesonotum e – g fore, middle, and hind legs h part of head with eye i tergalius III j flagellum of antenna k head with labrum, mandibles, and hypopharynx (a – g in one magnification, k holotype).	Figure 66. Papuanatula (Papuanatula) obscura sp. nov., larval exuviae: a, b abdomen with caudalii c head d half of pronotum and mesonotum e – g fore, middle, and hind legs h part of head with eye i tergalius III j flagellum of antenna k head with labrum, mandibles, and hypopharynx (a – g in one magnification, k holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247484	https://doi.org/10.3897/zookeys.1227.138100.figure70	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	Figure 70. Papuanatula (Papuanatula) obscura sp. nov.: a, b male imago c abdomen of male imago d subimaginal exuviae of half of mesonotum e subimaginal exuviae of mesopleuron f middle leg of male imago g, h female imago i subimaginal exuviae of middle tarsus of female (c, f holotype).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
94C0A0452CA0519E979CBDDEE814C6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247567	https://doi.org/10.3897/zookeys.1227.138100.figure148	Figure 148. Distribution of Papuanatula species (x: type locality).	Figure 148. Distribution of Papuanatula species (x: type locality).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
67323A685CF85B53A343ED508B4AE6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247417	https://doi.org/10.3897/zookeys.1227.138100.figure3	Figure 3. Papuanatula (Papuanatula) bessa, larva (from Waena): a, b labrum c labium d hypopharynx e, f left and right mandibles g maxilla.	Figure 3. Papuanatula (Papuanatula) bessa, larva (from Waena): a, b labrum c labium d hypopharynx e, f left and right mandibles g maxilla.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
67323A685CF85B53A343ED508B4AE6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247520	https://doi.org/10.3897/zookeys.1227.138100.figure106	Figure 106. Papuanatula (Papuanatula) parvatubera sp. nov., larva: a hind leg b middle tarsus and claw (arrow: posterior seta) c male head, mature d prosternum e paracercus f cercus, dorsal view g cercus, lateral view. Scale bars: 50 µm (a); 10 µm (b, f, g); 100 µm (c); 20 µm (d, e).	Figure 106. Papuanatula (Papuanatula) parvatubera sp. nov., larva: a hind leg b middle tarsus and claw (arrow: posterior seta) c male head, mature d prosternum e paracercus f cercus, dorsal view g cercus, lateral view. Scale bars: 50 µm (a); 10 µm (b, f, g); 100 µm (c); 20 µm (d, e).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
67323A685CF85B53A343ED508B4AE6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247522	https://doi.org/10.3897/zookeys.1227.138100.figure108	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	Figure 108. Papuanatula (Papuanatula) parvatubera sp. nov., larva (SEM): a pro-, meso-, and metasternum (arrows: protuberances) b hind claw c abdominal terga III – V d abdominal terga VII – X e abdominal tergum IV f abdominal tergum VIII. Scale bars: 200 µm (a); 40 µm (b); 100 µm (c, d); 30 µm (e); 20 µm (f).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
67323A685CF85B53A343ED508B4AE6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247418	https://doi.org/10.3897/zookeys.1227.138100.figure4	Figure 4. Papuanatula (Papuanatula) bessa, larva (a – e from Wagau, Herzog Mts.): a foreleg b fore claw c hind tibia, inner margin d hind tibia, posterior surface e femur, posterior apex f, g tergalius IV (type locality) h paracercus (type locality) I cercus section (type locality). Scale bars: 50 µm (a); 10 µm (b – d, g, i); 20 µm (f, h).	Figure 4. Papuanatula (Papuanatula) bessa, larva (a – e from Wagau, Herzog Mts.): a foreleg b fore claw c hind tibia, inner margin d hind tibia, posterior surface e femur, posterior apex f, g tergalius IV (type locality) h paracercus (type locality) I cercus section (type locality). Scale bars: 50 µm (a); 10 µm (b – d, g, i); 20 µm (f, h).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
67323A685CF85B53A343ED508B4AE6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247422	https://doi.org/10.3897/zookeys.1227.138100.figure8	Figure 8. Papuanatula (Papuanatula) copis, larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	Figure 8. Papuanatula (Papuanatula) copis, larva, habitus: a dorsal view b lateral view c ventral view. Scale bars: 1 mm.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
67323A685CF85B53A343ED508B4AE6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247426	https://doi.org/10.3897/zookeys.1227.138100.figure12	Figure 12. Papuanatula (Papuanatula) copis, larva: a metanotum and abdominal terga b abdominal terga c paracercus (type locality) d cercus (type locality) e paraproct (type locality) f cercotractor (type locality) g paraproct, distal margin (type locality). Scale bars: 100 µm (a); 10 µm (b, d – g); 20 µm (c).	Figure 12. Papuanatula (Papuanatula) copis, larva: a metanotum and abdominal terga b abdominal terga c paracercus (type locality) d cercus (type locality) e paraproct (type locality) f cercotractor (type locality) g paraproct, distal margin (type locality). Scale bars: 100 µm (a); 10 µm (b, d – g); 20 µm (c).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
67323A685CF85B53A343ED508B4AE6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247427	https://doi.org/10.3897/zookeys.1227.138100.figure13	Figure 13. Papuanatula (Papuanatula) copis, larva, SEM: a dorsal view b pronotum. Scale bars: 0.5 mm (a); 300 µm (b).	Figure 13. Papuanatula (Papuanatula) copis, larva, SEM: a dorsal view b pronotum. Scale bars: 0.5 mm (a); 300 µm (b).	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
67323A685CF85B53A343ED508B4AE6DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1247419	https://doi.org/10.3897/zookeys.1227.138100.figure5	Figure 5. Papuanatula (Papuanatula) bessa, larva, abdomen (from Waena): a tergum I b, c tergum II d apex of protuberance on tergum III e tergum IV f tergum V g tergum IX h paraprocts.	Figure 5. Papuanatula (Papuanatula) bessa, larva, abdomen (from Waena): a tergum I b, c tergum II d apex of protuberance on tergum III e tergum IV f tergum V g tergum IX h paraprocts.	2025-02-11	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kluge, Nikita J.;Gattolliat, Jean-Luc			
