identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9C99E1EE788C56A3AB128D65D377F4FF.text	9C99E1EE788C56A3AB128D65D377F4FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haplopteris yakushimensis C. W. Chen & Ebihara	<div><p>Haplopteris yakushimensis C. W. Chen &amp; Ebihara, Phytotaxa 156 (4): 232. 2014.</p><p>Fig. 1</p><p>Type.</p><p>Japan. • Kagoshima Pref., Yakushima Island,  Nakabase River, 20 Aug 1982, Nakaike s. n. (holotype TNS [VS-456666!])  .</p><p>Distribution and ecology.</p><p>This species was previously recorded only in Japan and Taiwan (Chen et al. 2014, TPG 2019). In Vietnam,  H. yakushimensis is found in damp evergreen broadleaf forests at elevations of 1200–1360 m, where it grows on rocks with thick compost near streams.</p><p>Specimens examined.</p><p>Vietnam. • Cao Bang Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.8804&amp;materialsCitation.latitude=22.589539" title="Search Plazi for locations around (long 105.8804/lat 22.589539)">Phia-Oac Phia-Den National Park</a>, 6 Dec 2013, Zhang et al. 6755 (MO, TAIF [499240!], VNMN)  .   Phia-Oac Phia-Den National Park, Nguyen Binh District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.8804&amp;materialsCitation.latitude=22.589539" title="Search Plazi for locations around (long 105.8804/lat 22.589539)">Thanh Cong Ward</a>, 22.589539°N, 105.880403°E, 1353 m, 8 Nov 2023, Chen Wade 6952 (TAIF!, VNMN!)  . •   Ha Giang Province: Vi Xuyen District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.880394&amp;materialsCitation.latitude=22.7675" title="Search Plazi for locations around (long 104.880394/lat 22.7675)">Cao Bo Ward</a>, 22.767500°N, 104.880394°E, 1200–1360 m, 12 Sep 2000, Harder et al. 5514 (UC [1763099!])  .</p><p>Note.</p><p>Haplopteris, a fern genus comprising approximately 40 species, is primarily found in tropical Africa, Asia, and the Pacific Islands (Schuettpelz et al. 2016). In Vietnam, eight  Haplopteris species were previously recorded:  H. angustifolia (Blume) E. H. Crane,  H. doniana (Mett. ex Hieron.) E. H. Crane,  H. elongata (Sw.) E. H. Crane,  H. ensata (Christ) C. W. Chen &amp; S. Linds.,  H. ensiformis (Sw.) E. H. Crane,  H. flexuosa (Fée) E. H. Crane,  H. hainanensis (C. Chr. ex Ching) E. H. Crane, and  H. sikkimensis (Kuhn) E. H. Crane (Chen et al. 2023) .  Haplopteris yakushimensis can be distinguished from these species by having fronds broader than 1 cm wide, costae that are grooved on the adaxial side and raised on the abaxial side, and the submarginal (ca. 1–2 mm away from the margins) sori lines.</p><p>Recent studies have identified two genetically distinct yet morphologically indistinguishable lineages in  H. yakushimensis (Kuo et al. 2017; TPG 2019). With the discovery of new populations in Vietnam, we are currently working to clarify these lineages using an integrated approach that includes reproductive biology, genome size estimation, and nuclear markers. The results of this research will be published in a separate paper.</p></div>	https://treatment.plazi.org/id/9C99E1EE788C56A3AB128D65D377F4FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chen, Cheng-Wei;Do, Van Truong;Truong, Quang Cuong;Dang, Viet Dai;Luu, Hong Truong;Chao, Yi-Shan;Huang, Yao-Moan;Chung, Kuo-Fang	Chen, Cheng-Wei, Do, Van Truong, Truong, Quang Cuong, Dang, Viet Dai, Luu, Hong Truong, Chao, Yi-Shan, Huang, Yao-Moan, Chung, Kuo-Fang (2025): Studies of Vietnamese Pteridophyte Flora 3. PhytoKeys 255: 143-169, DOI: 10.3897/phytokeys.255.141395
4A3F447C5DB15B358B5FCB3066F0443B.text	4A3F447C5DB15B358B5FCB3066F0443B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lindsaea kohkongensis I. C. Hwang, M. O. Moon & B. Y. Sun	<div><p>Lindsaea kohkongensis I. C. Hwang, M. O. Moon &amp; B. Y. Sun, Korean J. Pl. Taxon. 53 (4): 289. 2023.</p><p>Fig. 2</p><p>Type.</p><p>Cambodia • Koh Kong, Thma Bang District,  near Chamnar village, 22 Dec 2013, Sun et al. C 5520 (holotype HIBR; isotypes HIBR, KB)  .</p><p>Distribution and ecology.</p><p>This species was recently described from Cambodia and Malaysia. In Vietnam, it grows along a valley in damp tropical forests as lithophytes and is frequently submerged in water.</p><p>Specimens examined.</p><p>Vietnam. • Kien Giang Province: Phu Quoc district,  Phu Quoc National Park, 6 Mar 2018, Chen Wade 5034 (TAIF [514046!])  .</p><p>Note.</p><p>Morphologically, this species closely resembles  Lindsaea ensifolia . Although  L. kohkongensis is usually a much smaller species than  L. ensifolia, we did not identify any qualitative trait to distinguish them. Currently, subaquatic habitat preference appears to be the most reliable characteristic for differentiating this species from  L. ensifolia . Yun et al. (2023) described  L. kohkongensis as having free venation, in contrast to the anastomosing venation found in  L. ensifolia . However, this is not true. As demonstrated in Fig. 3 G of their paper and Fig. 2 F of the current study, the veins of  L. kohkongensis are connected near the lamina margin, forming areoles in fertile fronds.</p></div>	https://treatment.plazi.org/id/4A3F447C5DB15B358B5FCB3066F0443B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chen, Cheng-Wei;Do, Van Truong;Truong, Quang Cuong;Dang, Viet Dai;Luu, Hong Truong;Chao, Yi-Shan;Huang, Yao-Moan;Chung, Kuo-Fang	Chen, Cheng-Wei, Do, Van Truong, Truong, Quang Cuong, Dang, Viet Dai, Luu, Hong Truong, Chao, Yi-Shan, Huang, Yao-Moan, Chung, Kuo-Fang (2025): Studies of Vietnamese Pteridophyte Flora 3. PhytoKeys 255: 143-169, DOI: 10.3897/phytokeys.255.141395
FFBC7C92BCCB5197914306B00505EE9C.text	FFBC7C92BCCB5197914306B00505EE9C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pteris pseudowulaiensis Y. S. Chao	<div><p>Pteris pseudowulaiensis Y. S. Chao, Taiwania 66 (3): 314. 2021.</p><p>Fig. 3</p><p>Type.</p><p>Taiwan • New Taipei,  Mt. Pataoerh, 600–700 m, 29 Apr 2016, Hsu 8437 (holotype TAIF [497137!]; isotype TAIF [497138!])  .</p><p>Distribution and ecology.</p><p>This species, recently described from Taiwan, was initially reported from southern China and Taiwan (Chao et al. 2021). In Vietnam,  P. pseudowulaiensis was discovered in relatively dry evergreen broadleaf forests at lowland areas of Cuc Phuong National Park.</p><p>Specimens examined.</p><p>Vietnam • Ninh Binh Province: Cuc Phuong National Park,  Muong Khu trail, Y.-S. Chao 3509, 3515, 3517, 3531 (TAIF!, VNMN!)  .</p><p>Note.</p><p>This species belongs to the  Pteris fauriei Hieron. complex, a taxonomically challenging group due to its reticulate evolution involving hybridization, polyploidization, and apomixis (Chao et al. 2022). In Vietnam, five species from this complex have been recorded, namely  P. arisanensis Tagawa (Chao et al. 2022),  P. kawabatae Sa. Kurata (Chao et al. 2021),  P. latipinna Y. S. Chao &amp; W. L. Chiou (Chen et al. 2020 b),  P. oshimensis Hieron. (Liao et al. 2013), and  P. pseudowulaiensis . Morphologically,  P. pseudowulaiensis closely resembles  P. oshimensis but differs by having broader pinnae, measuring 2–3.5 cm compared to less than 2 cm in  P. oshimensis . According to Chao et al. (2022),  P. pseudowulaiensis is an apogamous diploid originating from hybridization of  P. wulaiensis C. M. Kuo (a species reported from Japan and Taiwan) and another unknown species.</p></div>	https://treatment.plazi.org/id/FFBC7C92BCCB5197914306B00505EE9C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chen, Cheng-Wei;Do, Van Truong;Truong, Quang Cuong;Dang, Viet Dai;Luu, Hong Truong;Chao, Yi-Shan;Huang, Yao-Moan;Chung, Kuo-Fang	Chen, Cheng-Wei, Do, Van Truong, Truong, Quang Cuong, Dang, Viet Dai, Luu, Hong Truong, Chao, Yi-Shan, Huang, Yao-Moan, Chung, Kuo-Fang (2025): Studies of Vietnamese Pteridophyte Flora 3. PhytoKeys 255: 143-169, DOI: 10.3897/phytokeys.255.141395
