identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9F96B36388CA50319B0A1BB68BCAB9BD.text	9F96B36388CA50319B0A1BB68BCAB9BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria anilii Sindhu Arya	<div><p>1. Drymaria anilii Sindhu Arya, Harsh Sing &amp; Iamonico, Plants 13: 3378. 2024.</p><p>Type.</p><p>India. Nagaland: Kohima District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.01358&amp;materialsCitation.latitude=25.626389" title="Search Plazi for locations around (long 95.01358/lat 25.626389)">Kisama Village</a>, 25°37'35"N, 94°60'48.9"E, alt. 1800 m, 20 October 2021, Arya 730 (holotype DMP n. v.; isotypes DMP n. v., RO n. v.) .</p><p>Distribution.</p><p>India.</p><p>Notes.</p><p>We suspect that these plants are hybrids between Drymaria cordata subsp. diandra and D. villosa; they require more study. This species is assessed as data deficient (DD) (Arya et al. 2024).</p></div>	https://treatment.plazi.org/id/9F96B36388CA50319B0A1BB68BCAB9BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
4303B7C6DE19595899A22BA38EDAFA06.text	4303B7C6DE19595899A22BA38EDAFA06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria anomala S. Watson	<div><p>2. Drymaria anomala S. Watson, Proc. Amer. Acad. Arts 25: 143. 1890.</p><p>Type.</p><p>Mexico. Coahuila: Carneros Pass, 9 September 1889, C. G. Pringle 2847 (holotype GH [00037715 image!]) .</p><p>Distribution.</p><p>Mexico.</p><p>Notes.</p><p>Duke (1961) mentioned that the holotype of Drymaria anomala is deposited in the US. However, this statement is an error, and the holotype is in fact the sheet at GH, where Sereno Watson worked. According to Eric Schuettpelz (pers. comm. 2024), the curator of the US, there are no specimens of C. G. Pringle 2847 in their herbarium.</p></div>	https://treatment.plazi.org/id/4303B7C6DE19595899A22BA38EDAFA06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
8C103D7E4A20598AA3D9B2EE6662D336.text	8C103D7E4A20598AA3D9B2EE6662D336.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria apetala Bartl.	<div><p>3. Drymaria apetala Bartl. in Presl, Reliq. Haenk. 2 (1): 7. 1831.</p><p>= Stellaria virgata Ser. ex DC., Prodr. 1: 396. 1824. Type: Without data (holotype G [G 00212357!]) .</p><p>= Drymaria macrantha A. Gray, U. S. Expl. Exped. 126. 1854. Type: Peru. [Lima]: Cobradillo [illegible, possibly misspelled as Obrajillo, town located in Canta, Lima], 1838, Wilkes Exploring Expedition s. n. (lectotype, designated by Duke 1961, pg. 234: US [10417/00103348!]; isolectotype: GH [00037755 image!]) .</p><p>= Drymaria virgata Briq., Annuaire Conserv. Jard. Bot. Genève 13: 370. 1911. Type: Peru (erroneously reported as Mexico in the protologue). Without data, H. Ruiz &amp; J. Pavon s. n. (holotype: G [G 00226765 image!]) .</p><p>Type.</p><p>Chile. Habitat in Chile, 1822, T. Haenke s. n. (lectotype, designated here: GOET [GOET 000576 image!]; isolectotypes: HAL [HAL 0117876 image!], PR [24309 image!]) .</p><p>Distribution.</p><p>Peru, Chile.</p><p>Notes.</p><p>In most cases, for new species described in Reliquiae Haenkeanae, the specimens in PR are considered holotypes. However, in the case of the treatment of Caryophyllaceae, which was prepared by Friedrich Gottlieb Bartling, there are also duplicates housed in GOET, of the Georg August Universität Göttingen. This is the institution where Bartling worked, and these specimens were likely available for study by him at the time of the species’ description. What we are uncertain of is whether he also examined the material at PR. Because of this doubt, we designated the specimen at GOET as the lectotype.</p><p>For Drymaria macrantha, Duke stated that the specimen in the US was the “ HOLOTYPE. ” However, both the specimen at US and that at GH were available to Asa Gray at the time of description and labeled by him as a new species. Thus, we treat Duke’s use of the term holotype as an error to be corrected into a lectotype (see Turland et al. (2018) art. 9.10).</p><p>The type of Drymaria virgata was reportedly collected by Pavon in Mexico, but these are probably duplicates from a collection in Peru, since there are no known specimens collected outside of Peru (Duke 1961), and Ruiz and Pavon did not collect in Mexico.</p></div>	https://treatment.plazi.org/id/8C103D7E4A20598AA3D9B2EE6662D336	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
229BA077FD0A505A870578727FF7D422.text	229BA077FD0A505A870578727FF7D422.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria arenarioides subsp. peninsularis (S. F. Blake) J. A. Duke	<div><p>4 b. Drymaria arenarioides subsp. peninsularis (S. F. Blake) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 198. 1961.</p><p>≡ Drymaria peninsularis S. F. Blake, J. Wash. Acad. Sci. 14 (13): 285. 1924. Type: Mexico. Baja California Sur: Cape Region, Jan. – Mar. 1901, C. A. Purpus 423 (holotype: US [470422/00103356 image!]; isotypes: UC [135816 / UC 135816 image!], MO [3343597 / 144084 image!]) .</p><p>= Drymaria johnstonii Wiggins, Proc. Calif. Acad. Sci. ser. 4, 25: 203. 1944. Type: Mexico. Baja California Sur: in crevices of rock on mesa near crest of island, the isthmus, Espiritu Santo Island, 31 May 1921, I. M. Johnston 3972 (holotype: CAS [82814/0002334 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/229BA077FD0A505A870578727FF7D422	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
F0089F01CE8A586DA33AA7EF9FAECC86.text	F0089F01CE8A586DA33AA7EF9FAECC86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria arenarioides Willd. Later, Britton and Millspaugh 1920	<div><p>4 a. Drymaria arenarioides Willd. in Roem. &amp; Schult., Syst. Veg. 5: 406. 1819. subsp. arenarioides</p><p>≡ Drymaria frankenioides Kunth, Nov. Gen. Sp. 6 [folio]: 18; [quarto]: 21; tab. 515. 1823. nom. illeg. superfl.</p><p>Type.</p><p>Mexico. [Hidalgo]: Pachuca, [May 1803], A. von Humboldt &amp; A. J. A. Bonpland 4070 (lectotype, designated here: P [P 00679593!]; isolectotype: HAL [HAL 0117875 image!]) .</p><p>Distribution.</p><p>Mexico.</p><p>Notes.</p><p>Although authorship of Drymaria arenarioides is often given as “ Willd. ex Schult., ” the name and description occur in a single paragraph followed by “ Reliq. Willd. MS. ” This represents direct association of Willdenow with both the name and the description, and Willdenow should be cited as author (see Turland et al. (2018), art. 46.3 example 15). Because the holotype in B was destroyed (Duke 1961; Hiepko 1987), we here designate the P specimen as the lectotype.</p></div>	https://treatment.plazi.org/id/F0089F01CE8A586DA33AA7EF9FAECC86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
D645789CAD045FBE8A52617290C4C4AC.text	D645789CAD045FBE8A52617290C4C4AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria auriculipetala Mattf.	<div><p>5. Drymaria auriculipetala Mattf., Notizbl. Bot. Gart. Berlin-Dahlem 13 (118): 438. 1936.</p><p>Type.</p><p>Peru. Huánuco: Llata, alt. 2500 m, 21 August 1922, J. F. Macbride &amp; W. Featherstone 2264 (holotype: F [518748 / F 0042709 F image!]; isotypes: GH [00037748 image!], MO [1186085 / 492968 image!], NY [00342514 image!], US [00103329 image!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>The sheet at F was labeled by Duke as an isotype, but it is in fact the holotype. It was correctly cited as the holotype in Duke (1961). Currently, this species is listed as endangered (EN) (Cano and Sánchez 2006) with occurrences in C Peru.</p></div>	https://treatment.plazi.org/id/D645789CAD045FBE8A52617290C4C4AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
BEE4A094030956F988968A80448FC36F.text	BEE4A094030956F988968A80448FC36F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria axillaris Brandegee	<div><p>6. Drymaria axillaris Brandegee, Univ. Calif. Publ. Bot. 4 (11): 178. 1911.</p><p>Type.</p><p>Mexico. Coahuila: Sierra del Rey, June 1910, C. A. Purpus 4526 (holotype: UC [144769 / UC 144769 image!]; isotypes: E [E 00326061 image!], F [344117 / F 0053286 F image!], GH [00037716 image!], MO [3343598 / 144085 image!], US 841897/00103330 image!]) .</p><p>Distribution.</p><p>United States, Mexico.</p></div>	https://treatment.plazi.org/id/BEE4A094030956F988968A80448FC36F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
6E32E8B08AB55F5F8D3EE39C7085C8B8.text	6E32E8B08AB55F5F8D3EE39C7085C8B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria barkleyi J. A. Duke & Steyerm.	<div><p>7. Drymaria barkleyi J. A. Duke &amp; Steyerm., Ann. Missouri Bot. Gard. 48 (3): 199. 1961.</p><p>Type.</p><p>Mexico. Coahuila: 25 miles southwest of Monterrey, 1 December 1945, B. H. Warnock &amp; F. A. Barkley 14826 (holotype: F [1176679 / F 0053280 F image!]; isotypes: GH [00037717 image!], TEX [00370752 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/6E32E8B08AB55F5F8D3EE39C7085C8B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
1115BB0BD44B56C8A66361E6B7801724.text	1115BB0BD44B56C8A66361E6B7801724.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria coahuilana (I. M. Johnst.) B. L. Turner	<div><p>8. Drymaria coahuilana (I. M. Johnst.) B. L. Turner, Phytologia 78 (3): 200. 1995.</p><p>≡ Drymaria lyropetala var. coahuilana I. M. Johnst., J. Arnold Arbor. 31: 189. 1950. Type: Mexico. Coahuila: 2 km NW of St. Elena. Vicinity of Santa Elena Mines, eastern foothills of the Sierra de las Cruces, 21 June 1941, R. M. Stewart 567 (holotype: GH [00037732 image!]; isotypes: ANSM [ANSM 007430 image!], BM [BM 000522057 image!], DUKE [310217/10000517 image!], F [1757720 / F 0053298 F image!], LL [299864/00370754 image!], MEXU [204377 / MEXU 00204377!], UC [1416791 / UC 1416791 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/1115BB0BD44B56C8A66361E6B7801724	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
45AFA79A6FEF51D18907085734CDCC0D.text	45AFA79A6FEF51D18907085734CDCC0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria conzattii J. A. Duke	<div><p>9. Drymaria conzattii J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 224. 1961.</p><p>Type.</p><p>Mexico. Oaxaca: from Almoloya to Sta. Catarina, alt. 1000 m, 26 December 1906, C. Conzattii 1688 (holotype: MO [839425/144086 image!]; isotypes: US [572151/00103335 image!], MEXU [MEXU 00528401!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/45AFA79A6FEF51D18907085734CDCC0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
C5A6EF59AA0F5654B705E2EE2C4DA6D6.text	C5A6EF59AA0F5654B705E2EE2C4DA6D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria cordata subsp. diandra (Blume) J. A. Duke	<div><p>10 b. Drymaria cordata subsp. diandra (Blume) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 253. 1961.</p><p>≡ Drymaria diandra Blume, Bijdr. Fl. Ned. Ind. (2): 62. 1825. Type: Indonesia. Java: according to the protologue, “ in paludosus montanis Javae insulae, ” s. d., C. L. Blume 1549 (lectotype, designated by Mizushima 1957, pg. 81: L [L 0038673 image!]; isolectotypes: L [L 0038674 image!], MO [1765827 / 216778 image!]) .</p><p>= Drymaria retusa Wight &amp; Arn., Prodr. Fl. Ind. Orient. 359. 1834. nom. invalid. Type: India. Peninsula Ind. Orientalis, s. d., R. Wight 152 (holotype: G [G 00226767 image!]) .</p><p>= Drymaria extensa Wall., Fl. Brit. India 1 (2): 244. 1874. nom. invalid. Type: India. Ind. Orient ., 1829, N. Wallich 647 (holotype: G [G 00226769 image!]) .</p><p>= Drymaria gerontogea F. Muell., Descr. Notes Papuan Pl. 1 (5): 87. 1877. nom. invalid.</p><p>Distribution.</p><p>Subspecies of wide distribution throughout Africa, Asia, and Oceania.</p><p>Notes.</p><p>Drymaria retusa and Drymaria extensa are invalid names because they were merely cited as provisional synonyms of D. cordata and not accepted. Drymaria gerontogea has been considered validly published by various authors (e. g., Duke, 1961). However, it was not definitively accepted at the time of publication. Mueller merely states, “ Should further research prove Blume’s and Macfadyen’s plants distinct, then I would propose the species name D. gerontogea for the former. ”</p></div>	https://treatment.plazi.org/id/C5A6EF59AA0F5654B705E2EE2C4DA6D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
D8F73D761C08555CABA55761511AF3CF.text	D8F73D761C08555CABA55761511AF3CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria cordata (L.) Willd.	<div><p>10 a. Drymaria cordata (L.) Willd. in Roem. &amp; Schult., Syst. Veg. 5: 406. 1819. subsp. cordata</p><p>≡ Holosteum cordatum L., Sp. Pl. 1: 88. 1753. Type: Tab. 11 “ Alsine americana nummulariae folio ” in Hermann, Paradisus Batavus, 1698 (lectotype, designated by Burger in Cafferty and Jarvis 2004, pg. 1052).</p><p>= Holosteum diandrum Sw., Prod. 27. 1788. Type: Jamaica. s. d., O. Swartz s. n. (holotype: S [S 10-22103 image!]; isotypes: G [G 00215840 image!], S [S-R-2936 image!]) .</p><p>≡ Drymaria diandra (Sw.) Macfad., Fl. Jamaica 1: 52. 1837. nom. illeg. [non D. diandra Blume (1825)].</p><p>≡ Drymaria cordata var. diandra (Sw.) Griseb., Fl. Brit. W. I. 56. 1859.</p><p>= Drymaria cordata var. puberula Triana &amp; Planch., Ann. Sci. Nat., Bot., sér. 4, 17: 148. 1862. Type: Colombia. [Cundinamarca]: Bogota, alt. 2600 m, s. d., J. J. Triana s. n. (lectotype, designated by Duke 1961, pg. 251: P n. v.; isolectotype NY [3578908 image!]) .</p><p>= Drymaria procumbens Rose, Contr. U. S. Natl. Herb. 1 (9): 304. 1895. Type: Mexico. Colima: near water ditches about Colima, 9 Jan. – 6 Feb. 1891, E. Palmer 1165 (holotype: US [15185/00103359 image!]; isotypes: K [K 000471743 image!], NY [00342506 image!], US [1391060 / 00955563 image!]) .</p><p>= Drymaria adenophora Urb., Repert. Spec. Nov. Regni Veg. 21 (8–20): 213. 1925. Type: Cuba. Pinar del Río: Cortez, 27 March 1921, L. E. Ekman 12798 (lectotype, designated here: S [S-R-1684 image!]; isolectotypes: G [G 00226766 image!], NY [00073724 image!]) .</p><p>≡ Stellaria adenophora (Urb.) Leon, Contr. Ocas. Mus. Hist. Nat. Colegio “ De La Salle ” 9: 4. 1950.</p><p>= Drymaria sessilifolia Fiori, Anal Bot. Fieni Imp.: 20. 1939. Type: Ethiopia. Adis Abeba, Olettà, alt. 1500 m, 19 October 1937, E. Bartolozzi 13 (holotype: FT [FT 001060 image!]) .</p><p>= Drymaria cordata var. pacifica M. Mizush., J. Jap. Bot. 32 (3): 78. 1957. Type: Ecuador. Galapagos Islands: Academy Bay, Indefatigable Islands, 100 ft [alt. 30.48 m], 1 April 1930, H. K. Svenson 65 (holotype: GH [00037749 image!]) .</p><p>Distribution.</p><p>Subspecies of wide distribution, from the United States to South America and the Caribbean.</p><p>Notes.</p><p>Mizushima (1957) chose the specimen at Herb. Linn. no. 109-1 as the lectotype of Drymaria cordata . However, this sheet, as mentioned by Burger in Cafferty and Jarvis (2004), is a post- 1753 addition to the Linnaean herbarium and consequently cannot be considered original material and is ineligible for typification of the name.</p><p>Although the illustration designated as type is not of high quality in showing some details of the flowers, e. g., the petals appear undivided, there is no doubt that it corresponds to the concept of Drymaria cordata subsp. cordata as currently recognized.</p><p>There are two sheets of Drymaria diandra at S. The specimen considered here as an isotype is apparently a fragment of the holotype that was later removed from the specimen.</p><p>Regarding the type of Drymaria cordata var. puberula, Duke (1961) used the term holotype for the Triana specimen at P cited above; however, various syntypes were mentioned in the protologue without a formal designation of type. Thus, Duke’s statement should be treated as an error to be corrected to a lectotype.</p><p>Most of the type specimen deposited at B were destroyed in World War II (Hiepko 1987). Because the holotype in B of Drymaria adenophora was destroyed, we here designated the specimen at S as the lectotype.</p><p>For the same reason as with Drymaria arenarioides, authorship of the Drymaria cordata combination should be attributed to Willdenow (see Turland et al. (2018), art. 46.3 example 15).</p></div>	https://treatment.plazi.org/id/D8F73D761C08555CABA55761511AF3CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
CA68CE82A3395452919F86463E72B4B9.text	CA68CE82A3395452919F86463E72B4B9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria cubana Alain	<div><p>11. Drymaria cubana Alain, Phytologia 8: 369. 1962.</p><p>≡ Pinosia glandulosa Alain, Candollea 17: 107. 1960. Type: Cuba. Oriente: Farallón de Macambo, Vía Azul, km. 70, 28 December 1954, Alain &amp; López Figueiras 4202 (holotype: LS n. v.) .</p><p>Distribution.</p><p>Cuba.</p><p>Notes.</p><p>When Liogier (1962) transferred Pinosia glandulosa, the epithet “ glandulosa ” could not be used for this species due to the existence of Drymaria glandulosa C. Presl. He proposed the new name Drymaria cubana .</p></div>	https://treatment.plazi.org/id/CA68CE82A3395452919F86463E72B4B9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
6A3B6F9FB83053CCBE548D4B1C74131E.text	6A3B6F9FB83053CCBE548D4B1C74131E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria debilis Brandegee	<div><p>12. Drymaria debilis Brandegee, Proc. Calif. Acad. Sci., ser. 2, 2: 131. 1889.</p><p>= Drymaria polystachya Brandegee, Zoë 2 (1): 70. 1891. Type: Mexico. Baja California Sur: San José del Cabo, 24 September 1890, T. S. Brandegee 35 (holotype: UC [120336 / UC 120336 image!]; isotypes: NY [00342505 image!], POM [151790/0002392 image!], US [10418/00103357 image!, 735811/00931426 image!]) .</p><p>= Drymaria diffusa Rose, Contr. U. S. Natl. Herb. 1 (5): 130, pl. 12. 1892. Type: Mexico. Baja California Sur: Carmen Island, 1 November 1890, E. Palmer 819 (holotype: US [10399/00952505 image!]; isotypes: F [166343 / F 0053289 F image!], GH [00311114 image!], LE [LE 00001741 image!], UC [120326 / UC 120326 image!], US [1391051 / 00952506 image!]) .</p><p>≡ Drymaria polystachya var. diffusa (Rose) Wiggins, Proc. Calif. Acad. Sci., ser. 4, 25: 198. 1944.</p><p>Type.</p><p>Mexico. Baja California Sur: Purisima, 13 February 1889, T. S. Brandegee s. n. (holotype: UC [120315 / UC 120315 image!]; isotypes: CAS [0002340 image!], PH [01031254 / 00009747 image!], US [10402/00103336 image!]) .</p><p>Distribution.</p><p>Mexico.</p><p>Notes.</p><p>The isotype at CAS of Drymaria debilis represents fragments removed from the holotype.</p></div>	https://treatment.plazi.org/id/6A3B6F9FB83053CCBE548D4B1C74131E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
B030AA1CFCD453C0AD37B8C046CFD259.text	B030AA1CFCD453C0AD37B8C046CFD259.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria divaricata Kunth 2025	<div><p>13 a. Drymaria divaricata Kunth, Nov. Gen. Sp. 6 [folio]: 20; [quarto]: 24. 1823. var. divaricata</p><p>= Drymaria peruviana Muschl., Bot. Jahrb. Syst. 45: 451. 1911. Type: Peru. “ la Oroya, in departimento Junin, in formatione plantis pulvinaribus et caespitosis composita, 4300 m s. m., ” February 1903, Weberbauer 2601 (syntype B, destroyed) ; Departimento Junin, in montibus prope Tarma in formatione aperta plantis herbaceis (praesertim graminibus) et fruticibus composita, 3600–3700 m s. m., ” 10 February 1903, Weberbauer 2411 (syntype B, destroyed) ; “ Departimento Junin, La Oroya, in sabulosis, 3900 m s. m., ” February 1903, Weberbauer 2589 (syntype B, destroyed) ; Pocarà, statio viae ferreae, in saxosis, 3700 m s. m., ” 24–25 February 1902, Weberbauer 400 (syntype B, destroyed) .</p><p>Type.</p><p>Peru. Lima: crescit ad litora Oceani Pacifici, [1802], A. von Humboldt &amp; A. J. A. Bonpland s. n. (holotype: P [P 00679596!]) .</p><p>Distribution.</p><p>Peru, Bolivia.</p><p>Notes.</p><p>Most of the type specimen deposited at B were destroyed in World War II (Hiepko 1987). The four collections cited in the protologue of Drymaria peruviana were apparently destroyed, because they are no longer at B. We have not been able to locate any duplicates.</p></div>	https://treatment.plazi.org/id/B030AA1CFCD453C0AD37B8C046CFD259	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
F94681350442519EA0F6E4983E2F331D.text	F94681350442519EA0F6E4983E2F331D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria divaricata var. divergens J. A. Duke	<div><p>13 d. Drymaria divaricata var. divergens J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 243. 1961.</p><p>Type.</p><p>Peru. Lima: Pativilca, lomas de Pacar, alt. 150–700 m, 1 October 1949, O. V. Nuñez 2320 (holotype: US [2248528 / 00603655 image!]; isotype: MO [2174615 / 491441 image!]) .</p><p>Distribution.</p><p>Peru.</p></div>	https://treatment.plazi.org/id/F94681350442519EA0F6E4983E2F331D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
82F4E138D15D5441B458895880C6A223.text	82F4E138D15D5441B458895880C6A223.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria divaricata var. reflexiflora J. A. Duke	<div><p>13 e. Drymaria divaricata var. reflexiflora J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 244. 1961.</p><p>Type.</p><p>Peru. Lima: Chancay, lomas de Lachay, alt. 500–600 m, 5 November 1955, R. Ferreyra 11522 (holotype: MO [2174617 / 492967 image!]; isotype: USM [18367 / USM 000298 image!]) .</p><p>Distribution.</p><p>Peru.</p></div>	https://treatment.plazi.org/id/82F4E138D15D5441B458895880C6A223	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
5322387E228B50338FB1ECAE0D3CD66E.text	5322387E228B50338FB1ECAE0D3CD66E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria divaricata var. stricta (Rusby) J. A. Duke	<div><p>13 b. Drymaria divaricata var. stricta (Rusby) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 242. 1961.</p><p>≡ Drymaria stricta Rusby, Phytologia 1: 55. 1934. Type: Bolivia. [La Paz]: Pongo, 12000 ft [alt. 3657.6 m], 17 Feb. – 1 Mar. 1926, G. H. H. Tate 160 (holotype: NY [00342520 image!]) .</p><p>= Drymaria agapatensis Baehni &amp; J. F. Macbr., Publ. Field Mus. Nat. Hist., Bot. Ser. 13 (2 / 2): 619. 1937. Type: Peru. [Ayacucho]: Agapata, January 1854, W. Lechler 1947 (holotype: G [G 00226770 image!]; isotypes: L [L 00012078 image!], S [S-R-1685 image!]) .</p><p>Distribution.</p><p>Peru, Bolivia.</p></div>	https://treatment.plazi.org/id/5322387E228B50338FB1ECAE0D3CD66E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
A1740BA5F306579A8C4C5860891752A6.text	A1740BA5F306579A8C4C5860891752A6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria divaricata var. viscidula (A. Gray) J. A. Duke	<div><p>13 c. Drymaria divaricata var. viscidula (A. Gray) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 243. 1961.</p><p>≡ Drymaria viscidula A. Gray, U. S. Expl. Exped. 124. 1854. Type: Peru. [Lima]: Obrajillo, 1838, Wilkes Exploring Expedition s. n. (holotype: US [10427/00103369 image!]) .</p><p>Distribution.</p><p>Peru.</p></div>	https://treatment.plazi.org/id/A1740BA5F306579A8C4C5860891752A6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
37E8F6F4E589546E8FE4EEC8AE1DA9E2.text	37E8F6F4E589546E8FE4EEC8AE1DA9E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria effusa A. Gray	<div><p>14 a. Drymaria effusa A. Gray, Smithsonian Contr. Knowl. 5 (6): 19. 1853. var. effusa</p><p>Type.</p><p>Mexico. Sonora: mountains east of Santa Cruz, 28 September 1851, C. Wright 869 (holotype: GH [00037671 image!]; isotypes: G [G 00226771 image!], GH [00254586 image!], MO [3578826 / 216548 image!], PH [01031255 / 00009749 image!]) .</p><p>Distribution.</p><p>United States, Mexico.</p></div>	https://treatment.plazi.org/id/37E8F6F4E589546E8FE4EEC8AE1DA9E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
4EFCBEDF14D35533B070BFF6313068B2.text	4EFCBEDF14D35533B070BFF6313068B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria effusa var. confusa (Rose) J. A. Duke	<div><p>14 b. Drymaria effusa var. confusa (Rose) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 205–206. 1961.</p><p>≡ Drymaria confusa Rose, Contr. U. S. Natl. Herb. 5 (8): 133. 1897. Type: Mexico. Chihuahua: San José, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.066666&amp;materialsCitation.latitude=28.616667" title="Search Plazi for locations around (long -106.066666/lat 28.616667)">Southwestern</a> Chihuahua, [28°37'N, 106°04'W], 1885, E. Palmer 59 (holotype: US [129525/00103333 image!]; isotypes: GH [00037720 image!], K [K 000471741 image!], LE [LE 00001742 image!], MO [1807078 / 144088 image!], VT [UVMVT 0224423 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/4EFCBEDF14D35533B070BFF6313068B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
1720F9E34B545440B070344229A325F7.text	1720F9E34B545440B070344229A325F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria effusa var. depressa (Greene) J. A. Duke	<div><p>14 c. Drymaria effusa var. depressa (Greene) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 206–208. 1961.</p><p>≡ Drymaria depressa Greene, Leafl. Bot. Observ. Crit. 1 (11): 153. 1905. Type: United States. New Mexico: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-108.38333&amp;materialsCitation.latitude=32.783333" title="Search Plazi for locations around (long -108.38333/lat 32.783333)">around the south end of the Black Range. Sawyer’s Peak</a>. Grant County, [32°47'N, 108°23'W], 9500 ft [alt. 2895.6 m], 30 September 1904, O. B. Metcalfe 1430 (lectotype, designated here: MO [3578825 / 216549 image!]; isolectotypes: AC [10967/00319675 image!], CAS [82826/0033361 image!], COLO [0363028 image!], E [E 00318138 image!], F [187730 / F 0053281 F image!], GH [00037670 image!], NY [00342487 image!], MSC [193377 / MSC 0091647 image!], RM [90033 / RM 0002158 image!], POM [151505/0002397 image!], UNM [295 / UNM 00039 image!]) .</p><p>= Drymaria minuscula Standl. &amp; Steyerm., Publ. Field Mus. Nat. Hist., Bot. Ser. 23 (2): 52. 1944. Type: Guatemala. Huehuetenango: Vicinity of Chemal, summit of Sierra de Los Cuchumatanes, 8 August 1942, J. A. Steyemark 50243 (holotype: F [1129120 / F 0053300 F image!]; isotype: US [1920024 / 00103349 image!]) .</p><p>Distribution.</p><p>United States, Mexico, Guatemala.</p><p>Notes.</p><p>No herbaria were cited by Greene in the protologue for Drymaria depressa . Duke (1961) reported that the holotype was at ND, presumably because that is where Greene worked at the time he described D. depressa . According to Barbara Hellenthal (pers. comm. 2024), the curator of ND, there are no specimens of Metcalfe 1430 in their herbarium, and it is likely that the material used by Greene was subsequently returned to Metcalfe. In the absence of the holotype, we here designate the specimen at MO as the lectotype because it is the most complete.</p></div>	https://treatment.plazi.org/id/1720F9E34B545440B070344229A325F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
76A2B6C53B75506E9D3D2F159D750DEC.text	76A2B6C53B75506E9D3D2F159D750DEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria elata I. M. Johnst.	<div><p>15. Drymaria elata I. M. Johnst., J. Arnold Arbor. 21: 68. 1940.</p><p>Type.</p><p>Mexico. Coahuila: road from Mohovano northeast to Estancia Station: 6 miles south of Laguna del Rey, 21 September 1938, I. M. Johnston 7823 (holotype: GH [00037722 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/76A2B6C53B75506E9D3D2F159D750DEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
66F522FC262B5770B0CCC8A34E2D47D0.text	66F522FC262B5770B0CCC8A34E2D47D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria engleriana (Muschl.) Baehni & J. F. Macbr.	<div><p>16 a. Drymaria engleriana (Muschl.) Baehni &amp; J. F. Macbr., Publ. Field Mus. Nat. Hist., Bot. Ser. 13 (2 / 2): 621. 1937. var. engleriana</p><p>≡ Polycarpon englerianum Muschl., Bot. Jahrb. Syst. 45 (4): 452. 1911. Type: Peru. Ancash: Caraz, alt. 4300 m, 1906, A. Weberbauer 3101 (lectotype, designated here: MOL [00219/00000486 image!]; isolectotype: G [G 27707/00226772 image!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>Most of the type specimen deposited at B were destroyed in World War II (Hiepko 1987). Because the holotype in B was probably destroyed, we here designate the MOL specimen as the lectotype. Currently, this variety is listed as Endangered (EN) (Cano and Sánchez 2006).</p></div>	https://treatment.plazi.org/id/66F522FC262B5770B0CCC8A34E2D47D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
DF4A4FF6BC70561B96F8C418407EE32C.text	DF4A4FF6BC70561B96F8C418407EE32C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria engleriana var. devia (Baehni & J. F. Macbr.) J. A. Duke	<div><p>16 b. Drymaria engleriana var. devia (Baehni &amp; J. F. Macbr.) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 219. 1961.</p><p>≡ Drymaria devia Baehni &amp; J. F. Macbr., Publ. Field Mus. Nat. Hist., Bot. Ser. 13 (2 / 2): 620. 1937. Type: Peru. Lima: near Antaicocha, Cerro Colorado, east of Canta, alt. 3800–4000 m, 20 June 1925, F. W. Pennell 14655 (holotype: F [558629 / F 0042710 F!]; isotypes: GH [00037750 image!], PH [774610/00009748 image!], S [S-R-1686 image!], US [1340894 / 00103339 image!]) .</p><p>Distribution.</p><p>Peru, Chile.</p><p>Notes.</p><p>The sheet at F was labeled by Duke as an isotype, but it is in fact the holotype as cited in Duke (1961).</p></div>	https://treatment.plazi.org/id/DF4A4FF6BC70561B96F8C418407EE32C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
4C7BCC8FE307551C8A3252A4368A8139.text	4C7BCC8FE307551C8A3252A4368A8139.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria excisa Standl.	<div><p>17. Drymaria excisa Standl., Publ. Field Mus. Nat. Hist., Bot. Ser. 8 (1): 11. 1930.</p><p>= Drymaria grandis Bullock, Bull. Misc. Inform. Kew 1936 (6): 389. 1936. Type: Mexico. México: Temascaltepec, alt. 2440 m, 21 January 1934, G. B. Hinton 5427 (holotype: K [K 000471758 image!]; isotypes: BM [BM 000939014 image!], G [G 00226773 image!], GH [00037728 image!], NY [00342498 image!], NY [00342499 image!], RSA [350580 / RSA 0002395 image!], S [S-G-7134 / 7134 image!]) .</p><p>Type.</p><p>Mexico. Jalisco: Sierra Madre Occidental, Real Alto, trail to El Tajo de Santiago, alt. 2500 m, 23 February 1927, Y. E. J. Mexia 1748 (holotype: F [579891 / F 0053290 F image!]; isotypes: CAS [157122/0002333 image!], CAS [195467/0027349 image!], G [G 00226774 image!], GH [0003724 image!], MICH [1115007 image!], MO [954774/144089 image!], NY [00342496 image!], UC [350610 / UC 350610 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/4C7BCC8FE307551C8A3252A4368A8139	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
6BF2E039C57056A0BFAA3856E8A30131.text	6BF2E039C57056A0BFAA3856E8A30131.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria fasciculata A. Gray	<div><p>18. Drymaria fasciculata A. Gray, U. S. Expl. Exped., Phan.: 125. 1854.</p><p>Type.</p><p>Peru. [Lima]: Andes near Obrajillo, 1838, Wilkes Exploring Expedition s. n. (lectotype, designated by Duke 1961, pg. 217: US [10405/00103342 image!]; isolectotype GH [00037751 image!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>Duke stated that the specimen in the US was the “ HOLOTYPE. ” However, both the specimen at US and that at GH were available to Asa Gray at the time of description and labeled by him as a new species. Thus, we treat Duke’s use of the term holotype as an error to be corrected to lectotype.</p></div>	https://treatment.plazi.org/id/6BF2E039C57056A0BFAA3856E8A30131	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
9877251A44C85A4BB27B99847DF7565A.text	9877251A44C85A4BB27B99847DF7565A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria firmula Steyerm.	<div><p>19. Drymaria firmula Steyerm., Publ. Field Mus. Nat. Hist., Bot. Ser. 28 (1): 227. 1951.</p><p>Type.</p><p>Venezuela. Merida: Páramo de Los Colorados, between El Molino and San Isidro Alto, alt. 2745–2955 m, 14 May 1944, J. A. Steyermark 56536 (holotype F: [1340308 / 0053307 F image!]; isotypes: MO [2174628 / 216513 image!], NY [00342515 image!]) .</p><p>Distribution.</p><p>Venezuela.</p></div>	https://treatment.plazi.org/id/9877251A44C85A4BB27B99847DF7565A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
ACCE5D9D434E557780E5180E2E0772E4.text	ACCE5D9D434E557780E5180E2E0772E4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria frutescens Mattf.	<div><p>20. Drymaria frutescens Mattf., Notizbl. Bot. Gart. Berlin-Dahlem 13 (118): 439. 1936.</p><p>Type.</p><p>Peru. La Libertad: Santiago de Chuco, above Hacienda Angasmarca, alt. 3650 m, May 1916, A. Weberbauer 7203 (lectotype, designated here: F [628040 / F 0042712 F image!]; isolectotypes: F [549079 / F 0042713 F image!], GH [00037752 image!], MO [1768271 / 216512 image!], US [1495416 / 00103344 image!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>Duke mentions that the holotype was in B. However, the protologue (Mattfeld, 1936) clearly states that the type is in F. Due to the existence of two sheets at F, here we designated the specimen [628040/0042712 F] as the lectotype. This species is listed as Critically Endangered (CR) (Cano and Sánchez 2006).</p></div>	https://treatment.plazi.org/id/ACCE5D9D434E557780E5180E2E0772E4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
0C9AE0ED2FA250A1B5B58ADA7C774828.text	0C9AE0ED2FA250A1B5B58ADA7C774828.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria glaberrima Bartl.	<div><p>21. Drymaria glaberrima Bartl. in Presl, Reliq. Haenk. 2 (1): 7. 1831.</p><p>Type.</p><p>Peru. [Lima]: Habitat in Peruviae montanis huanoccensibus, 1822, T. Haenke s. n. (lectotype, designated here: GOET [GOET 000577!]; isolectotype: PR [24305 image!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>For the same reason as mentioned for Drymaria apetala, we designate the specimen in GOET as the lectotype. This species is listed as Endangered (EN) (Cano and Sánchez 2006).</p></div>	https://treatment.plazi.org/id/0C9AE0ED2FA250A1B5B58ADA7C774828	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
D1AD164213EB541F82BE2B6CA7B8F5EF.text	D1AD164213EB541F82BE2B6CA7B8F5EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria glandulosa Phil.	<div><p>22 a. Drymaria glandulosa Phil., Fl. Atacam. 10. 1860. var. glandulosa</p><p>= Drymaria ramosissima Schltdl., Linnaea 12 (2): 206. 1838. Type: Mexico. Prope urben Mexico, s. d., Hegewisch s. n. (holotype: HAL [HAL 0098412 image!]; isotype: CAS [82822/0002331 image!]) .</p><p>= Drymaria leptoclados Hemsl., Diagn. Pl. Nov. Mexic. 1: 2. 1878. Type: Guatemala. [Guatemala]: camino del Sapote, December 1865, K. G. Bernoulli 240 (holotype: K [3862 / K 000471752 image!]; isotypes: G [G 00226778 image!], NY [00342500 image!]) .</p><p>= Drymaria fendleri S. Watson, Proc. Amer. Acad. Arts 17: 328. 1882. Type: United States. New Mexico, 1 January 1847, A. Fendler 60 (lectotype, designated by Duke 1961, pg. 247: GH [00037672 image!]; isolectotypes: GH [00037673 image!, 00037674 image!], K [K 000723291 image!]) .</p><p>≡ Drymaria glandulosa var. fendleri (S. Watson) Fosberg, Contr. U. S. Natl. Herb. 29 (2): 96. 1957.</p><p>= Drymaria leptoclados var. peruviana Ball, J. Proc. Linn. Soc., Bot. 22: 32. 1885. Type: Peru. [Lima]: Chicla, 21 April 1882, J. Ball s. n. (holotype: K [K 000471733 image!]) .</p><p>= Drymaria blasiana M. E. Jones, Contr. W. Bot. 15: 125. 1929. Type: Mexico. Sinaloa: San Blas, 30 January 1927, M. E. Jones 22845 (holotype: POM [162740/0002399 image!]) .</p><p>= Drymaria fendleri var. perennis M. E. Jones, Contr. W. Bot. 18: 65. 1933. Type: Mexico: Baja California Sur: Laguna Mountains, La Laguna, 22 September 1930, M. E. Jones 27050 (holotype: POM [192443/0002396 image!]; isotypes: F [721383 / F 0053291 F image!], MIN [626682/1000875 image!], NY [00342497 image!], UC [522237 / UC 522237 image!]) .</p><p>≡ Drymaria glandulosa var. perennis (M. E. Jones) Fosberg, Contr. U. S. Natl. Herb. 29: 96. 1945.</p><p>Type.</p><p>Mexico: habitat in Mexico, s. d., T. Haenke s. n. (holotype: PR [24310 image!]).</p><p>Distribution.</p><p>United States, Mexico, Guatemala, Honduras, Peru, Bolivia, Argentina.</p><p>Notes.</p><p>For Drymaria fendleri, Duke stated that the specimen in GH was the “ HOLOTYPE. ” Thus, we treat Duke’s use of the term holotype as an error to be corrected to lectotype.</p></div>	https://treatment.plazi.org/id/D1AD164213EB541F82BE2B6CA7B8F5EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
2568A2D6BD815529B726C9D36253D64B.text	2568A2D6BD815529B726C9D36253D64B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria glandulosa var. galeottiana (Briq.) J. A. Duke	<div><p>22 b. Drymaria glandulosa var. galeottiana (Briq.) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 249. 1961.</p><p>≡ Drymaria galeottiana Briq., Annuaire Conserv. Jard. Bot. Genève 13: 373. 1911. Type: Mexico. Oaxaca: near Oaxaca, 1840, H. G. Galeotti 4408 (holotype: G [G 00226777 image!]; isotypes: K [K 3862/000471749 image!], LE [LE 00001740 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/2568A2D6BD815529B726C9D36253D64B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
31D5F152961259AD943FFE115BA82A0D.text	31D5F152961259AD943FFE115BA82A0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria gracilis Phil.	<div><p>23 a. Drymaria gracilis Phil., Fl. Atacam. 10. 1860. subsp. gracilis</p><p>≡ Drymaria cordata var. gracilis (Schltdl. &amp; Cham.) Rohrb., Fl. Bras. 14 (2): 260. (1872).</p><p>Type.</p><p>Mexico. Veracruz: Jalapam [ Xalapa], August 1827, C. J. W. Schiede &amp; F. Deppe 503 (holotype: HAL [HAL 098420/0098420 image!]; isotypes: GOET [GOET 000578!], LE [LE 00001739 image!]) .</p><p>Distribution.</p><p>Mexico, Guatemala, Honduras, El Salvador.</p><p>Notes.</p><p>Duke mentioned that the holotype of Drymaria gracilis was at B, but it is in fact the specimen at HAL, where Schlechtendal’s herbarium is now housed.</p></div>	https://treatment.plazi.org/id/31D5F152961259AD943FFE115BA82A0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
BCB0CF49B45751CCA45862304E78C31E.text	BCB0CF49B45751CCA45862304E78C31E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria gracilis subsp. carinata (Brandegee) J. A. Duke	<div><p>23 b. Drymaria gracilis subsp. carinata (Brandegee) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 247. 1961.</p><p>≡ Drymaria carinata Brandegee, Zoë 2 (1): 70. 1891. Type: Mexico. Baja California Sur: Sierra de la Laguna, 21 January 1890, T. S. Brandegee 34 (holotype: UC [120331 / UC 120331 image!]; isotypes: NY [00342491 image!], POM [101189/0002398 image!]) .</p><p>= Drymaria carinata var. perennis Wiggins, Proc. Calif. Acad. Sci., ser. 4, 25: 197. (1944). Type: Mexico. Baja California Sur: Sierra Laguna, La Laguna, 6000 ft [alt. 1828.8 m], 25 March 1939, H. S. Gentry 4415 (holotype: DS [263289/0002338 image!]; isotypes: ARIZ [268235 / ARIZ-BOT-0005308 image!, 319000 / ARIZ-BOT-0005625 image!], MO [1159044 / 144090 image!], UC [UC 708997 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/BCB0CF49B45751CCA45862304E78C31E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
576653AA661F5F6392903B979126F520.text	576653AA661F5F6392903B979126F520.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria grandiflora Bartl. From 1849	<div><p>24. Drymaria grandiflora Bartl. in Presl, Reliq. Haenk. 2 (1): 7. 1831.</p><p>Type.</p><p>Peru. [Huánuco]: Habitat in Chile and in Peruviae montanis huanoccensibus, 1822, T. Haenke s. n. (lectotype, designated here: GOET [GOET 000579 image!]; isolectotypes: GOET [GOET 000580 image!], HAL [HAL 0117872 image!], PR [24308 image!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>For the same reason as mentioned for Drymaria apetala, we designate the specimen at GOET as the lectotype. The protologue mentions that this species grows in Chile. In fact, this information is incorrect (Duke 1961). This species is listed as Near Threatened (NT) (Cano and Sánchez 2006).</p></div>	https://treatment.plazi.org/id/576653AA661F5F6392903B979126F520	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
D7877E93FE7550518CF28754ED28F9CF.text	D7877E93FE7550518CF28754ED28F9CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria holosteoides Benth.	<div><p>25 a. Drymaria holosteoides Benth., Bot. Voy. Sulphur: 16. 1844. var. holosteoides</p><p>≡ Mollugophytum holosteoides (Benth.) M. E. Jones, Contr. W. Bot. 18: 35. 1933.</p><p>= Drymaria veatchii Curran, Proc. Calif. Acad. Sci. 2 (1): 227. 1888. Type: Mexico. Baja California Sur: Magdalena Island, 1888, W. E. Bryant s. n. (holotype: UC [124950 / UC 124950 image!]) .</p><p>Type.</p><p>Mexico. Baja California Sur: Bay of Magdalena, 1841, R. B. Hinds s. n. (holotype K [K 000471757 image!]; isotype LE [LE 00001737 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/D7877E93FE7550518CF28754ED28F9CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
DC903FA959C15B33B0468ADB00B89129.text	DC903FA959C15B33B0468ADB00B89129.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria holosteoides var. crassifolia (Benth.) J. A. Duke	<div><p>25 b. Drymaria holosteoides var. crassifolia (Benth.) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 187. 1961.</p><p>≡ Drymaria crassifolia Benth., Bot. Voy. Sulphur: 16. 1844. Type: Mexico. Baja California Sur: Cape San Lucas, 1841, R. B. Hinds s. n. (holotype: K [3862 / K 000471763 image!]; isotype: LE [LE 00012079 image!]) .</p><p>≡ Mollugophytum crassifolium (Benth.) M. E. Jones, Contr. W. Bot. 18: 35. 1933.</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/DC903FA959C15B33B0468ADB00B89129	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
1962C97E2EC75117BD5643E71F41AD17.text	1962C97E2EC75117BD5643E71F41AD17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria hypericifolia Briq.	<div><p>26. Drymaria hypericifolia Briq., Annuaire Conserv. Jard. Bot. Genève 13–14: 369. 1911.</p><p>Type.</p><p>Mexico. Oaxaca: from Lecambre de Yolotepec to Juquila, southwest of Oaxaca, March 1845, C. Jürgensen 38 (lectotype, designated by Duke 1961, pg. 201: G [G 00226746 image!]; isolectotypes: F [869035 / F 0053294 F image!], K [K 000471756 image!]) .</p><p>Distribution.</p><p>Mexico, Guatemala, Honduras.</p><p>Notes.</p><p>Two collections were cited in the protologue, Jurgensen no. 38 and Galeotti no. 4425, and neither was designated as the type. Duke (1961) stated that the holotype was Jungersen´s collection.</p></div>	https://treatment.plazi.org/id/1962C97E2EC75117BD5643E71F41AD17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
63EE74160D945D6B8A3EA8A0DDAF0042.text	63EE74160D945D6B8A3EA8A0DDAF0042.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria jenniferae Villarreal & A. E. Estrada	<div><p>27. Drymaria jenniferae Villarreal &amp; A. E. Estrada, Brittonia 60: 329. 2008.</p><p>Type.</p><p>Mexico. Coahuila: mpio. Viesca, E of 5 de Mayo, alt. 1606 m, 30 September 2006, G. B. Hinton 28498 (holotype: MEXU [1393821 / MEXU 01393821!]; isotypes: ANSM [ANSM 087856 /087856 image!], CIIDIR [CIIDIR 022559 image!], GBH [GBH 028498 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/63EE74160D945D6B8A3EA8A0DDAF0042	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
06F531449A245DC9A9A2E241CDC45ACB.text	06F531449A245DC9A9A2E241CDC45ACB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria ladewii Rusby	<div><p>28. Drymaria ladewii Rusby, Phytologia 1 (2): 54. 1934.</p><p>Type.</p><p>Bolivia. Nequejahuira: Cordillera Real, 8000 ft [alt. 2438.4 m], 15–24 May 1926, G. H. H. Tate 652 (lectotype, designated here: NY [00342516 image!]; isolectotype: NY [00342517 image!]) .</p><p>Distribution.</p><p>Mexico, Guatemala, El Salvador, Costa Rica, Panama, Peru, Bolivia.</p><p>Notes.</p><p>There are two separate sheets of Tate 652 at NY, and neither was indicated as the holotype by Rusby. We select the better of the two as the lectotype.</p></div>	https://treatment.plazi.org/id/06F531449A245DC9A9A2E241CDC45ACB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
DA0328239CD65AC1922C055FECAD2A06.text	DA0328239CD65AC1922C055FECAD2A06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria laxiflora Benth.	<div><p>29. Drymaria laxiflora Benth., Pl. Hartw.: 73. 1841.</p><p>= Drymaria chihuahuensis Briq., Annuaire Conserv. Jard. Bot. Genève 13–14: 370. 1911. Type: Mexico. Chihuahua: near Chihuahua, 1 September 1885, C. G. Pringle 331 (holotype: G [G 00226751 image!]; isotypes: ARIZ [319001 / ARIZ-BOT-0005626 image!], CAS [94965/0002339 image!], E [E 00326059 image!, 00394858 image!], F [104212 / F 0053287 F image!, 870682/0053288 F image!], G [G 00226752 image!, 00226762 image!], GH [00037719 image!], JE [JE 00014348 image!], KFTA [KFTA 0000328 image!], LE [LE 00001743 image!, LE 00001744 image!], LECB [LECB 0000546 image!], LL [00370753 image!], MEL [MEL 2499824 image!, MEL 2499825 image!], MEXU [MEXU 00148765!], MICH [1115008 image!], NY [00342492 image!, 00342493 image!, 00342494 image!], UC [UC 41993 image!], US [10411/00931458 image!], VT [UVMVT 024422 image!]) .</p><p>Type.</p><p>Guatemala. Quetzaltenango: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.48333&amp;materialsCitation.latitude=14.783334" title="Search Plazi for locations around (long -91.48333/lat 14.783334)">In rupibus Sunil</a>, [14°47'N, 91°29'W], 1839, K. T. Hartweg 523 (holotype: K [K 000471754 image!]; isotypes: E [E 00326060 image!], F [871784 / F 0053295 F image!, 869007/0053296 F image!], FI [021732/005970 image!], G [G 00226750 image!, 00226763 image!], K [K 000471753 image!], LD [1216996 image!], LE [LE 00012081 image!], MO [6033460 / 2090484 image!]) .</p><p>Distribution.</p><p>United States, Mexico, Guatemala, Bolivia.</p><p>Notes.</p><p>Duke reported that the holotype of Drymaria laxiflora is at BM. However, we interpret the holotype to be the specimen at K ex Herbarium Benthamianum.</p><p>For Drymaria chihuahuensis, there is a sheet at VT marked as holotype. However, as mentioned by Duke (1961), the holotype is at G.</p></div>	https://treatment.plazi.org/id/DA0328239CD65AC1922C055FECAD2A06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
57CD08582A1D54169FDBBA781D16712E.text	57CD08582A1D54169FDBBA781D16712E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria leptophylla (Cham. & Schltdl.) Rohrb.	<div><p>30 a. Drymaria leptophylla (Cham. &amp; Schltdl.) Fenzl ex Rohrb., Linnaea 37 (2): 195. 1871 –1873 [1872]. var. leptophylla</p><p>≡ Arenaria leptophylla Cham. &amp; Schltdl., Linnaea 5 (2): 233. 1830. Type: Mexico. Hidalgo: al E de Pachuca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.6975&amp;materialsCitation.latitude=20.128056" title="Search Plazi for locations around (long -98.6975/lat 20.128056)">al lado de la Carretera Federal 105 hacia Mineral del Monte, Mpio. de Mineral de la Reforma</a>, 20°07'41"N, 98°41'51"W, alt. 2689 m, 28 September 2024, D. Valentín-Martínez &amp; V. W. Steinmann 1204 (neotype, designated here: IEB [274928!]) .</p><p>≡ Spergularia leptophylla (Schltdl. &amp; Cham.) G. Don, Gen. Hist. 1: 425. 1831.</p><p>≡ Spergula leptophylla (Schltdl. &amp; Cham.) D. Dietr., Syn. Plant. 2: 1598. 1840.</p><p>= Drymaria tenella A. Gray, Mem. Amer. Acad. Arts, n. s. 4 (1): 12. 1849. Type: United States. New Mexico: 8 miles W of Las Vegas, New Mexico, shady places in woodland in the mountain region, 1847, A. Fendler 56 (holotype: GH [00037680 image!]; isotype: MO [3578827 / 2196012 image!]) .</p><p>= Drymaria nodosa var. gracillima Hemsl., Diagn. Pl. Nov. Mexic. 2: 22. 1879. Type: Mexico. San Luis Potosi: Chiefly in the region of San Luis Potosi, 6000–8000 ft [alt. 1828.8–2438.4 m], 1878, C. C. Parry &amp; E. Palmer 60 (holotype: K [K 000471750!]; isotypes: GH [00037735 image!], MO [3343588 / 144092 image!], NY [00342502 image!], WIS [v 0254955 WIS image!]) .</p><p>≡ Drymaria gracillima (Hemsl.) Rose, Contr. U. S. Natl. Herb. 5: 132. 1897.</p><p>Distribution.</p><p>United States, Mexico.</p><p>Notes.</p><p>Most of the type specimen deposited at B were destroyed in World War II (Hiepko 1987). Because the holotype of Arenaria leptophylla deposited in B was destroyed and no duplicates of the original material were found in other herbaria such as HAL, where Schlechtendal’s herbarium is housed, we designated the specimen at IEB as a neotype. The species was described based on C. J. W. Schiede 511 from “ Montis Orizabae, ” Veracruz. In the absence of a good quality specimen from near the type locality, we have chosen the above specimen because it corresponds precisely to the application of the name Drymaria leptophylla and is from the neighboring state of Veracruz.</p></div>	https://treatment.plazi.org/id/57CD08582A1D54169FDBBA781D16712E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
DFDB7C7530165BCFA656845D2160DA0E.text	DFDB7C7530165BCFA656845D2160DA0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria leptophylla var. cognata (S. F. Blake) J. A. Duke	<div><p>30 b. Drymaria leptophylla var. cognata (S. F. Blake) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 208. 1961.</p><p>≡ Drymaria cognata S. F. Blake, J. Washington Acad. Sci. 14: 285. 1924. Type: Mexico. Durango: vicinity of Durango, April 1896, E. Palmer 912 (holotype: US [304403/00103332 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/DFDB7C7530165BCFA656845D2160DA0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
C9C83578E8415F8398A1A9F653C1E79A.text	C9C83578E8415F8398A1A9F653C1E79A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria leptophylla var. nodosa (Engelm.) J. A. Duke	<div><p>30 c. Drymaria leptophylla var. nodosa (Engelm.) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 209. 1961.</p><p>≡ Drymaria nodosa Engelm., Mem. Amer. Acad. Arts, ser. 2, 4 (1): 12. 1849. Type: Mexico. Chihuahua: Cosiquiriadi, s. d., F. A. Wislizenus s. n. (holotype: GH [00037734 image!]) .</p><p>≡ Drymaria tenella var. nodosa (Engelm.) Wiggins, Proc. Calif. Acad. Sci., ser. 4, 25: 205. 1944.</p><p>= Drymaria gentryi Fosberg, Proc. Biol. Soc. Washington 62: 147. 1949. Type: Mexico. Chihuahua: Rio Mayo, Los Cascarones, 11 September 1936, H. S. Gentry 2669 (holotype: US [1686921 / 00103345 image!]; isotypes: ARIZ [319002 / ARIZ-BOT-0005627 image!, 319003/0005628 image!], F [862589 / F 0053293 F image!], GH [00037726 image!], MO [1162012 / 144091 image!], S [S-G-7136 image!], UC [UC 582148 image!]) .</p><p>Distribution.</p><p>United States, Mexico.</p></div>	https://treatment.plazi.org/id/C9C83578E8415F8398A1A9F653C1E79A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
81C41D63382751DE8A41D4EFBAAD6932.text	81C41D63382751DE8A41D4EFBAAD6932.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria longepedunculata S. Watson	<div><p>31. Drymaria longepedunculata S. Watson, Proc. Amer. Acad. Arts 25: 142. 1890.</p><p>Type.</p><p>Mexico. Jalisco: cliffs of the barranca, near Guadalajara, 28 November 1888, C. G. Pringle 2121 (holotype: GH [00037730 image!]; isotypes: UC [UC 120340 image!], VT [UVMVT 024424 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/81C41D63382751DE8A41D4EFBAAD6932	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
22C2C12F24E7515EB12F796816C09346.text	22C2C12F24E7515EB12F796816C09346.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria lyropetala I. M. Johnst.	<div><p>32. Drymaria lyropetala I. M. Johnst., J. Arnold Arbor. 21: 68. 1940.</p><p>Type.</p><p>Mexico. San Luis Potosi: 2 miles south of Cedral, on the road from Matehuala north to San Miguel on the state-boundary, 11 September 1938, I. M. Johnston 7594 (holotype: GH [00037731 image!]; isotypes: F [1016593 / F 0053297 F image!], S [S-G-7133 image!], UC [UC 741256 image!], US [1823067 / 00103347 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/22C2C12F24E7515EB12F796816C09346	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
A8D9E7272B9F5130B6AF70522DD2AFD7.text	A8D9E7272B9F5130B6AF70522DD2AFD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria malachioides Briq.	<div><p>33. Drymaria malachioides Briq., Annuaire Conserv. Jard. Bot. Genève 13: 372. 1911.</p><p>Type.</p><p>Mexico. [Michoacán]: Cordillere d’ Ario, Jun. – Oct. 1840, H. G. Galeotti 4419 (holotype: G [G 00226779 image!]; isotypes: MO [1669694 / 2090290 image!], P [P 04936753 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/A8D9E7272B9F5130B6AF70522DD2AFD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
883F4FB36614527E8356BD57462C285B.text	883F4FB36614527E8356BD57462C285B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria molluginea (Schrad.) Didr.	<div><p>34. Drymaria molluginea (Schrad.) Didr., Linnaea 29: 738. 1858.</p><p>≡ Alsine molluginea Lag., Gen. Sp. Pl. 13. 1816. nom. illeg. [non A. molluginea Hornem., (1813)]. Type: Without locality information, s. d., M. Lagasca “ 1806 ” (lectotype, designated by Duke 1961, pg. 194: G-DC [00212545 image!]) .</p><p>≡ Arenaria molluginea Schrad., Index Seminum [Gottingen] 1819: 2. 1819.</p><p>≡ Arenaria molluginea Ser., Prodr. 1: 400. 1824. nom. illeg. superfl. [non A. molluginea Schrad., 1819].</p><p>≡ Stellaria molluginea (Schrad.) Sweet, Hort. Brit. ed. 2: 56. 1830.</p><p>≡ Spergularia molluginea (Schrad.) G. Don., Gen. Hist. 1: 425. 1831.</p><p>≡ Lepigonum mollugineum (Schrad.) Bartl., Index Seminum [Gottingen] 1838: 4. 1838.</p><p>= Alsine molluginea Hornem., Hort. Bot. Hafn. 1: 295. 1815 [1813]. Type: Cultivated at the Copenhagen Botanical Garden, s. d., J. W. Horneman s. n. (holotype: [C image!]) .</p><p>= Alsine molluginea Mart., Plant. Hort. Erlang. 82. 1814. nom. illeg. [non A. molluginea Hornem., (1813)]. Type: Cultivated at the Erlangen Botanical Garden, s. d., C. F. P. Martius s. n. (lectotype, designated here: BM [BM 001024715 image!] [ex. Herb. Roemer]) .</p><p>= Drymaria sperguloides A. Gray, Mem. Amer. Acad. Arts 4 (1): 10. 1849. Type: United Sates. New Mexico, 1847, A. Fendler 55 (holotype: GH [00037679 image!]; isotypes: MO [3578836 / 216516 image!]) .</p><p>≡ Mollugophytum sperguloides (A. Gray) M. E. Jones, Contr. W. Bot. 18: 35. 1933.</p><p>Distribution.</p><p>United States, Mexico.</p><p>Notes.</p><p>As mentioned above, the name Alsine molluginea was proposed independently three times for the same taxon but with different types. Seringe is often credited as first proposing a name in Arenaria in 1824. However, five years earlier, Schrader provided a legitimate name in Arenaria . Although he did not include a description or diagnosis, his name should be accepted because his citation of “ Alsine Lag. ” constitutes an indirect reference to Lagasca’s description published in 1816. (see Turland et al. (2018), art. 41.3).</p><p>Although the specimen of Alsine molluginea at G-DC does not provide information, the protologue states that it is based on plants grown in the “ H. R. M. ” [Real Jardín Botánico de Madrid] from seed gathered in Nueva Hispania by D. D. Sessé [sic] &amp; J. M. Mociño. Drymaria molluginea was first described as Alsine molluginea by the Danish botanist Jens Wilken Hornemann in an account of the plants found in the Copenhagen Botanical Garden. His morphological description was brief but stated that the species was an annual with divided petals and thick, linear, secund leaves. He also indicated that the habitat of the species is unknown, but that it was introduced in the garden in 1812 from material provided by the Vienna Botanical Garden. The specimen cited above was labeled by Horneman himself as Alsine molluginea (handwriting confirmed by Jens Soelberg, C), and is the only original material known. Martius independently described Alsine molluginea in his thesis on the plants from the Erlangen Botanical Garden. He stated that they came from the Paris Botanical Garden “ h. Par. ” The specimen at BM was labeled by Martius (handwriting confirmed by Han Joaquin Esser, M) as “ Alsine molluginea mihi enum. horti. Erlangensis. ” There are no specimens at M, where most of Martius’ collections can be found, and for that reason, we designate the collection at BM as the lectotype. The reason that we are attributing authorship of the basionym of Drymaria molluginea to Schrader and not to Hornemann is that Lagasca’s name was clearly mentioned when the combination to Drymaria was made by Didrichsen (1858), and no indication of Hornemann’s earlier proposal of Alsine molluginea was provided.</p></div>	https://treatment.plazi.org/id/883F4FB36614527E8356BD57462C285B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
81A127C9A91853D9A2DD6712A3F4C69F.text	81A127C9A91853D9A2DD6712A3F4C69F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria monticola Howell	<div><p>35. Drymaria monticola Howell, Proc. Calif. Acad. Sci., Ser. 4, 21: 329. 1935.</p><p>Type.</p><p>Ecuador. Galapagos Islands: Indefatigable Island, 9 May 1932, J. T. Howell 9243 (holotype: CAS [0002336 image!]; isotypes: B [B 10 0365633!], S [S-R-1687 image!]) .</p><p>Distribution.</p><p>Ecuador.</p><p>Notes.</p><p>This species is listed as Critically Endangered (CR) (León-Yáñez et al. 2011).</p></div>	https://treatment.plazi.org/id/81A127C9A91853D9A2DD6712A3F4C69F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
8B059505C5EE5E64845B4D843F8BA9F4.text	8B059505C5EE5E64845B4D843F8BA9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria multiflora Brandegee	<div><p>36. Drymaria multiflora Brandegee, Zoë 5 (11): 232. 1906.</p><p>Type.</p><p>Mexico. [México]: Salto de Agua, October 1905, C. A. Purpus 1653 (lectotype, designated by Duke 1961, pg. 224: UC [UC 125379 image!]; isolectotypes: F [193175 / F 0053301 F image!], GH [00037733 image!], MO [3578885 / 144093 image!], US [570477/00103350 image!]) .</p><p>Distribution.</p><p>Costa Rica, Guatemala, Mexico, Panama.</p></div>	https://treatment.plazi.org/id/8B059505C5EE5E64845B4D843F8BA9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
D827170E29ED5C3894E1A679BB7864B0.text	D827170E29ED5C3894E1A679BB7864B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria ortegioides Griseb.	<div><p>37. Drymaria ortegioides Griseb., Cat. Pl. Cub.: 21. 1866.</p><p>≡ Pinosia ortegioides (Griseb.) Urb., Ark. Bot. 23 A (5): 71. 1930.</p><p>Type.</p><p>Cuba. Cuba occ., 1863, C. Wright 2019 (holotype: GOET [GOET 000581!]; isotypes: BM [BM 000939016 image!], G [G 00226747 image!], GH [00037747 image!], K [K 000471735 image!], LE [LE 00001735 image!], NY [00073725 image!, 00073726 image!], S [S-R-1688 image!, S-R-1689 image!], US [10421/00103352 image!], YU [YU.001065 image!]) .</p><p>Distribution.</p><p>Cuba.</p></div>	https://treatment.plazi.org/id/D827170E29ED5C3894E1A679BB7864B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
A6FDC44C469A5C8D92A5B31143B310E1.text	A6FDC44C469A5C8D92A5B31143B310E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria ovata Willd.	<div><p>38. Drymaria ovata Willd. in Roem. &amp; Schult., Syst. Veg. 5: 406. 1819.</p><p>Type.</p><p>Ecuador. Quito, [1802], A. von Humboldt &amp; A. J. A. Bonpland s. n. (holotype: P [P 00274219!]; isotype: P [P 00335772!]) .</p><p>Distribution.</p><p>Ecuador, Venezuela, Peru, Bolivia, Argentina.</p><p>Notes.</p><p>The specimens deposited in P with the label “ Herbier Humboldt &amp; Bonpland ” come from Bonpland’s private collection. According to Stauffer et al. (2012), these are to be considered holotypes. In the case of Drymaria ovata, there are two specimens at P representing original material. The one with the barcode “ P 00274219 ” bears the label “ Herbier Humboldt &amp; Bonpland ” and represents the holotype.</p><p>For the same reason as with Drymaria arenarioides, authorship of Drymaria ovata should be attributed to Willdenow (see Turland et al. (2018), art. 46.3 example 15).</p></div>	https://treatment.plazi.org/id/A6FDC44C469A5C8D92A5B31143B310E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
A1CB1ADF704C5246955B921761F89F5A.text	A1CB1ADF704C5246955B921761F89F5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria pachyphylla Wooton & Standl.	<div><p>39. Drymaria pachyphylla Wooton &amp; Standl., Contr. U. S. Natl. Herb. 16 (3): 121. 1913.</p><p>Type.</p><p>United States. New Mexico: south of White Sands, 4100 ft [alt. 1249.6 m], 28 August 1897, E. O. Wooton 405 (holotype: US [330629/00103353 image!]; isotypes: CAS [93988/0033360 image!], E [E 00318137 image!], G [G 00226748 image!, 00226749 image!], LE [LE 00001734 image!], MO [1746734 / 216515 image!], NDG [49829 / 16745 image!], NY [00342490 image!], RM [RM 0002159 image!], US [735952/00931427 image!]) .</p><p>Distribution.</p><p>United States, Mexico.</p></div>	https://treatment.plazi.org/id/A1CB1ADF704C5246955B921761F89F5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
A029B3F8A2C75184BBAA5146617CFCD1.text	A029B3F8A2C75184BBAA5146617CFCD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria paposana Phil.	<div><p>40 a. Drymaria paposana Phil., Fl. Atacam. 10. 1860. var. paposana</p><p>Type.</p><p>Chile. [Antofagasta]: Prope Paposo in valle Tartal, s. d., R. A. Philippi s. n. (holotype: SGO [048885 / SGO 000001961 image!]) .</p><p>Distribution.</p><p>Bolivia, Chile.</p><p>Notes.</p><p>At MO, there is a photo of an isotype ([048885/166961!]) probably from the Austrian herbarium W.</p></div>	https://treatment.plazi.org/id/A029B3F8A2C75184BBAA5146617CFCD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
A84EC468A13D52BDBC02288FE8818253.text	A84EC468A13D52BDBC02288FE8818253.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria paposana var. serrulata J. A. Duke	<div><p>40 c. Drymaria paposana var. serrulata J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 238. 1961.</p><p>Type.</p><p>Peru. Arequipa: Caraveli, lomas de Atiquipa (Tamaira), alt. 500–550 m, 27 November 1958, R. Ferreyra 13490 (holotype: MO [2174593 / 216510 image!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>This variety is listed as vulnerable (VU) (Cano and Sánchez 2006).</p></div>	https://treatment.plazi.org/id/A84EC468A13D52BDBC02288FE8818253	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
4E69DC72519A5485ADC0939BD34A68DA.text	4E69DC72519A5485ADC0939BD34A68DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria paposana var. weberbaueri (Muschl.) J. A. Duke	<div><p>40 b. Drymaria paposana var. weberbaueri (Muschl.) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 238. 1961.</p><p>≡ Drymaria weberbaueri Muschl., Bot. Jahrb. Syst. 45 (4): 451. 1911. Type: Peru. [Lima]: Barranca prope Lima in formatione “ Loma ” dicta, alt. 50–200 m, s. d., A. Weberbaueri 1662 (lectotype, designated by Duke 1961, pg. 238: G [G 00226776 image!]) .</p><p>= Drymaria fenzliana Briq., Annuaire Conserv. Jard. Bot. Genève 13: 373. 1911. Type: Mexico [Peru]. Without data, J. A. Pavon s. n. (holotype: G [G 00226775 image!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>Most of the type specimen deposited at B were destroyed in World War II (Hiepko 1987). The holotype of Drymaria weberbaueri at B was probably destroyed; for that reason, Duke (1961) designated a lectotype. This variety is listed as vulnerable (VU) (Cano and Sánchez 2006).</p><p>The type of Drymaria fenzliana was reportedly collected by Pavon in Mexico, but these are probably duplicates from a collection in Peru; Ruiz and Pavon did not collect in Mexico (Duke 1961).</p></div>	https://treatment.plazi.org/id/4E69DC72519A5485ADC0939BD34A68DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
C824CA78D6C25D5487E1EF5EB18DB74E.text	C824CA78D6C25D5487E1EF5EB18DB74E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria pattersonii B. L. Turner	<div><p>41. Drymaria pattersonii B. L. Turner, Phytologia 78 (3): 201. 1995.</p><p>Type.</p><p>Mexico. Coahuila: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-100.78333&amp;materialsCitation.latitude=26.833334" title="Search Plazi for locations around (long -100.78333/lat 26.833334)">ca. 1 km E of the highway at Rancho Santa Lucia along highway 30 between Monclova and Candela, where the road turns SE, 2.3 mi NW of the turnoff to La Carrosa, 23 mi W of the intersection with highway</a>, 26°50'N, 100°47'W, alt. 800–900 m, 18 October 1993, A. Prather &amp; T. Patterson 1496 (holotype: TEX [00031240 image!] .</p><p>Distribution.</p><p>Mexico.</p><p>Notes.</p><p>In the protologue, an isotype was reported from MEXU. However, according to the person in charge of the type collection (Ma. del Rosario García Peña) (pers. comm., 2025), there is no specimen located there.</p></div>	https://treatment.plazi.org/id/C824CA78D6C25D5487E1EF5EB18DB74E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
D027F1DB072757E782A0892814C05B0F.text	D027F1DB072757E782A0892814C05B0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria perennis Gilli	<div><p>42. Drymaria perennis Gilli, Feddes Repert. 92: 677. 1981.</p><p>Type.</p><p>Ecuador. Cotopaxi: Zwischen El Corazon und Angamarca, alt. 2600 m, 2 June 1975, A. Gilli 306 (holotype: W [11292 image!]) .</p><p>Distribution.</p><p>Ecuador.</p></div>	https://treatment.plazi.org/id/D027F1DB072757E782A0892814C05B0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
01D9B472859F5A42929F7D030E09BF47.text	01D9B472859F5A42929F7D030E09BF47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria polycarpoides A. Gray	<div><p>43. Drymaria polycarpoides A. Gray, Mem. Amer. Acad. Arts 4: 12. 1849.</p><p>≡ Mollugophytum polycarpoides (A. Gray) M. E. Jones, Contr. W. Bot. 18: 35. 1933.</p><p>Type.</p><p>Mexico. Durango: Valley of Bolson of Mapimi, s. d., J. Gregg s. n. (holotype: GH [00037737 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/01D9B472859F5A42929F7D030E09BF47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
3E41CC54E20D58E1AFA8F9E91B944FC4.text	3E41CC54E20D58E1AFA8F9E91B944FC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria praecox Baehni & J. F. Macbr.	<div><p>44. Drymaria praecox Baehni &amp; J. F. Macbr., Publ. Field Mus. Nat. Hist., Bot. Ser. 13 (2 / 2): 625. 1937.</p><p>Type.</p><p>Peru. Paucartambo: Cuzco, between Pisac and Paucartambo, alt. 4100 m, 23 April 1914, A. Weberbauer 6916 (holotype: F [548919 / F 0042714 F image!]; isotypes: GH [00037754 image!], MO [2174610 / 216511 image!], MOL [00229/00000481!], US [1496204 / 00103358 image!], USM [18409 / USM 000299!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>The sheet at F was labeled by Duke as an isotype, but it is in fact the holotype, as noted in Duke (1961). This species is currently listed as Endangered (EN) (Cano and Sánchez 2006).</p></div>	https://treatment.plazi.org/id/3E41CC54E20D58E1AFA8F9E91B944FC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
917C802C32245845B12509C50ECA53FC.text	917C802C32245845B12509C50ECA53FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria pratheri B. L. Turner	<div><p>45. Drymaria pratheri B. L. Turner, Phytologia 78 (3): 200. 1995.</p><p>Type.</p><p>Mexico. Nuevo Leon: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-100.8&amp;materialsCitation.latitude=26.416666" title="Search Plazi for locations around (long -100.8/lat 26.416666)">seven miles NE of Las Estacas (about midway between Monclova and Minas on highway 53) on a private ranch road towards Lechuguillal</a>, 26°25'N, 100°48'W, alt. 650 m, 18 October 1993, A. Prather, T. Patterson, and A. Hempel 1483 (holotype: TEX [00031239 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/917C802C32245845B12509C50ECA53FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
3BCED369148A55ABBBE928FB405D0371.text	3BCED369148A55ABBBE928FB405D0371.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria rotundifolia A. Gray	<div><p>46 a. Drymaria rotundifolia A. Gray, U. S. Expl. Exped. 123. 1854. var. rotundifolia</p><p>Type.</p><p>Peru. [Lima]: Andes, Obrajillo, 1838, Wilkes Exploring Expedition s. n. (lectotype, designated by Duke 1961, pg. 239: US [10423/00103360 image!]; isolectotypes: GH [00037756 image!], P [P 04936735 image!]) .</p><p>Distribution.</p><p>Ecuador, Peru.</p><p>Notes.</p><p>Duke (1961) stated that the specimen in the US was the “ HOLOTYPE. ” However, the specimens at P, GH, and at US were available to Asa Gray at the time of description and labeled by him as a new species. Thus, we treat Duke’s use of the term holotype as an error to be corrected to lectotype.</p></div>	https://treatment.plazi.org/id/3BCED369148A55ABBBE928FB405D0371	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
1F3E4BFA2AAD5ABEB2DBF94804A511F5.text	1F3E4BFA2AAD5ABEB2DBF94804A511F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria rotundifolia var. mattfeldiana J. T. Howell	<div><p>46 c. Drymaria rotundifolia var. mattfeldiana J. T. Howell, Leaf. W. Bot. 10: 351. 1966.</p><p>Type.</p><p>Ecuador. Galapagos Islands: Isabela Island, 4000 ft [alt. 1219.2 m], 26 May 1932, J. T. Howell 9568 (holotype: CAS [213490/0002332 image!]; isotypes: B [B 10 1226983!], US [2814511 / 00103361 image!]) .</p><p>Distribution.</p><p>Ecuador.</p><p>Notes.</p><p>In the protologue, an isotype was reported from the PRC. However, according to the curator (Patrik Mráz) (pers. comm., 2024), there is no specimen located there.</p></div>	https://treatment.plazi.org/id/1F3E4BFA2AAD5ABEB2DBF94804A511F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
440629F26A905BDBAAA93445C73C9464.text	440629F26A905BDBAAA93445C73C9464.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria rotundifolia var. nitida (Ball) J. A. Duke	<div><p>46 b. Drymaria rotundifolia var. nitida (Ball) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 239. 1961.</p><p>≡ Drymaria nitida Ball, J. Proc. Linn. Soc., Bot. 22: 31. 1887. Type: Peru. [Lima]: Ex saxosis Andium Peruviae juxta pagum Chicla, 12000–13000 ft [alt. 3657.6–3962.4 m], 21–23 April 1882, J. Ball s. n (holotype: K [K 000471731!]; isotypes: GH [00037753 image!], LE [LE 00001736 image!], NY [00342518 image!]) .</p><p>Distribution.</p><p>Ecuador, Peru.</p></div>	https://treatment.plazi.org/id/440629F26A905BDBAAA93445C73C9464	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
4860E0C5ED2F57FF8C9639BDFC030F29.text	4860E0C5ED2F57FF8C9639BDFC030F29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria stellarioides Willd.	<div><p>47. Drymaria stellarioides Willd. in Roem. &amp; Schult., Syst. Veg. 5: 406. 1819.</p><p>Type.</p><p>Ecuador. [Tungurahua]: Crescit prope Hambato, alt. 1380 hex. [alt. 564 m] [ Regno Quitensi], [1802], A. von Humboldt &amp; A. J. A. Bonpland s. n. (lectotype, designated here: P [P 00335861!]; isolectotype P [P 00335860!]) .</p><p>Distribution.</p><p>Ecuador.</p><p>Notes.</p><p>The specimens deposited in P with the label “ Herbier Humboldt &amp; Bonpland ” come from Bonpland’s private collection. According to Stauffer et al. (2012), these are to be considered the holotypes. In the case of Drymaria stellarioides, there are two specimens at P representing original material. However, neither has the label “ Herbier Humboldt &amp; Bonpland. ” Therefore, we designate the one with the barcode “ P 00335861 ” as the lectotype because it is the better specimen and the only one with a flower.</p><p>Most of the type specimen deposited at B were destroyed in World War II (Hiepko 1987). Because the holotype at B was probably destroyed, here we designate one of the specimens of P as the lectotype.</p><p>For the same reason as with Drymaria arenarioides, authorship of Drymaria stellarioides should be attributed to Willdenow (see Turland et al. (2018), art. 46.3 example 15).</p><p>At F, there is a photo and a fragment probably from the holotype ([980359/0053308 F!]). This species is listed as Near Threatened (NT) (León-Yáñez et al. 2011).</p></div>	https://treatment.plazi.org/id/4860E0C5ED2F57FF8C9639BDFC030F29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
9C347F5F917F5242B95F266001C97400.text	9C347F5F917F5242B95F266001C97400.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria stereophylla Mattf. 1936	<div><p>48. Drymaria stereophylla Mattf., Notizbl. Bot. Gart. Berlin-Dahlem 13 (118): 436. 1936.</p><p>≡ Drymaria stereophylla Mattf. ex J. F. Macbride, Publ. Field Mus. Nat. Hist., Bot. Ser. 13 (2 / 2): 626. 1937. nom. illeg. [non D. stereophylla Mattf. (1936)].</p><p>= Drymaria stereophylla var. exstipulata Mattf., Notizbl. Bot. Gart. Berlin-Dahlem 13 (118): 437. 1936. Type: Peru. Junin: nordöstlich von Huancayo unterhalb der Hacienda Acopalca, alt. 3600–3700 m, 11 April 1913, A. Weberbauer 6600 (lectotype, designated here: US [1498310 / 00603668 image!]; isolectotypes: F [627961 / F 0042716 F!], MO [2174586 / 216509 image!], MOL [00000478 image!], US [1497167 / 00603667 image!]) .</p><p>Type.</p><p>Peru. Junin: La Oroya, 12000 ft [alt. 3657.6 m], 27 May – 7 Jun. 1922, J. F. Macbride &amp; W. Featherstone 962 (lectotype, designated here: F [517490 / F 0042715 F image!]; isolectotypes: GH [00037757 image!], NY [00342519 image!], US [1802047 / 00103363 image!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>Mattfeld described Drymaria stereophylla based on a series of specimens collected by A. Raimondi (2502, 2903) and J. F. Macbride and W. Featherstone (563, 962, 1761). However, the protologue does not mention which specimen is the holotype. Furthermore, it is likely that the specimens deposited at B were destroyed in World War II along with most of the type specimens of angiosperms as mentioned by Hiepko (1987) since there are no original specimens there. Thus, the Macbride &amp; Featherstone specimen deposited at F [517490 / F 0042715 F!] is designated as the lectotype.</p><p>Because the holotype of Drymaria stereophylla var. exstipulata at B was probably destroyed, here we designated one of the specimens at US as the lectotype.</p></div>	https://treatment.plazi.org/id/9C347F5F917F5242B95F266001C97400	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
05E49088152C5FD899A371BE8856EEAA.text	05E49088152C5FD899A371BE8856EEAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria stipitata Fosberg	<div><p>49. Drymaria stipitata Fosberg, Lloydia 4 (4): 281. 1941.</p><p>Type.</p><p>Mexico. Coahuila: mpio. of Sierra Mojada, Sierra Mojada, San Salvador Canyon, 14 September 1939, C. H. Muller 3301 (holotype: US [2133254 / 00103364 image!]; isotypes: GH [00037739 image!], LL [00370755 image!], MICH [1115006 image!], MO [3578889 / 144038 image!], UC [UC 719747 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/05E49088152C5FD899A371BE8856EEAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
3B821508C5DD508A90228C43104C263C.text	3B821508C5DD508A90228C43104C263C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria subumbellata I. M. Johnst.	<div><p>50. Drymaria subumbellata I. M. Johnst., J. Arnold Arbor. 31: 188. 1950.</p><p>Type.</p><p>Mexico. Coahuila: south end of Cañada Oscuro, near Tanque la Luz, 28 October 1941, I. M. Johnston 8489 (holotype: GH [00037740 image!]; isotypes: GH [00037741 image!], LL [00370756 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/3B821508C5DD508A90228C43104C263C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
6AD8DE70AA3A59D093B9629CC831B1C4.text	6AD8DE70AA3A59D093B9629CC831B1C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria suffruticosa A. Gray	<div><p>51. Drymaria suffruticosa A. Gray, Proc. Amer Acad. Arts. 17: 328. 1882.</p><p>Type.</p><p>Mexico. Coahuila: San Lorenzo de Laguna to Vicinity, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-102.18&amp;materialsCitation.latitude=25.439999" title="Search Plazi for locations around (long -102.18/lat 25.439999)">22 to 27 leagues SW from Parras</a>, (25°26'24"N, 102°10'48"W), 1–10 May 1880, E. Palmer 74 (holotype: GH [00037742 image!]; isotypes: F [302659 / F 0053304 F image!], K [K 000471742 image!], LE [LE 00001731 image!], MO [2174589 / 144042 image!], NY [00342507 image!, 00342508 image!], US [10422/00103366 image!], VT [UVMVT 024425 image!], YU [YU.068841 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/6AD8DE70AA3A59D093B9629CC831B1C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
35BC9EC8F0875499B5E5CD93871ED32F.text	35BC9EC8F0875499B5E5CD93871ED32F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria tenuis S. Watson	<div><p>52. Drymaria tenuis S. Watson, Proc. Amer. Acad. Arts 25: 142. 1890.</p><p>≡ Drymaria tenuis var. genuina Briq., Annuaire Conserv. Jard. Bot. Genève 13–14: 374. 1911. nom. invalid.</p><p>= Drymaria filiformis Seaton, Proc. Amer. Acad. Arts 28: 117. 1893. Type: Mexico. Veracruz: Orizaba, 9000 ft [alt. 2743.2 m], 8 August 1891, H. E. Seaton 267 (holotype: GH [00037725 image!]; isotypes: F [266658 / F 0053292 F image!], K [K 000471788!], MO [3578890 / 144039 image!], NY [00581302 image!], US [1391052 / 00931428 image!, 10397/00103343 image!]) .</p><p>= Drymaria tenuis var. jaliscana Briq., Annuaire Conserv. Jard. Bot. Genève 13–14: 374. 1911. Type: Mexico. Jalisco: near Guadalajara, September 1893, C. G. Pringle 4536 (holotype: G [G 00640059 image!]; isotypes: COLO [422760/00363044 image!], E [E 00394861 image!], F [264418 / F 0053305 F image!], GH [00037744 image!], KFTA [KFTA 0000330 image!], LECB [LECB 0000547 image!], MEL [MEL 2499838 image!], MICH [1115005 image!], MO [3578893 / 356353 image!], NY [00342509 image!], PUL [PUL 00000233 image!], UC [UC 120343 image!], US [75112/00103367 image!], VT [UVMVT 024427 image!]) .</p><p>Type.</p><p>Mexico. Jalisco: under ledges of the barranca, near Guadalajara, 28 November 1888, C. G. Pringle 2120 (holotype: GH [00037743 image!]; isotype: VT [UVMVT 024426 image!]).</p><p>Distribution.</p><p>Mexico.</p><p>Notes.</p><p>Duke (1961) mentioned that the holotype of Drymaria tenuis is deposited in the US. However, this statement is an error. We have been unable to find a specimen of Pringle 2120 in the US, and the holotype is in fact the sheet at GH, where Sereno Watson worked at the time the species was described.</p></div>	https://treatment.plazi.org/id/35BC9EC8F0875499B5E5CD93871ED32F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
2143F10BCEFC53CE8A4FBBF39D7BEF7F.text	2143F10BCEFC53CE8A4FBBF39D7BEF7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria veliziae Montesinos	<div><p>53. Drymaria veliziae Montesinos, PhytoKeys 140: 48. 2020.</p><p>Type.</p><p>Peru. Cajamarca: Encañada, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.505554&amp;materialsCitation.latitude=-6.832222" title="Search Plazi for locations around (long -78.505554/lat -6.832222)">Chanta baja</a>, 6°49'56"S, 78°30'20"W, alt. 3295 m, 6 June 2009, C. Tovar 1058 (holotype: CPUN [22705!]) .</p><p>Distribution.</p><p>Peru.</p><p>Notes.</p><p>Currently, this species is listed as endangered (EN) (Montesinos-Tubée et al. 2020).</p></div>	https://treatment.plazi.org/id/2143F10BCEFC53CE8A4FBBF39D7BEF7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
35969A6DE94B5038ACBE2BE167A97A33.text	35969A6DE94B5038ACBE2BE167A97A33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria villosa Schltdl. & Cham.	<div><p>54 a. Drymaria villosa Schltdl. &amp; Cham., Linnaea 5 (2): 232. 1830. subsp. villosa</p><p>≡ Drymaria cordata var. villosa (Schltdl. &amp; Cham.) Rohrb., Fl. Bras. 14 (2): 260. 1872.</p><p>= Drymaria hirsuta Bartl. in Presl, Reliq. Haenk. 2 (1): 8. 1831. Type: Peru: montanis huanoccensibus, 1822, T. Haenke s. n. (lectotype, designated here: GOET [GOET 000583 image!]; isolectotype: PR [24307 image!]) .</p><p>= Drymaria cubensis Regel, Allg. Gartenzeitung 8 (38): 298. 1840. Type: probably Cuba. [Probably cultivated in the Botanical Garden of Berlin from Cuban seed]. (holotype: K [K 000471737 image!]) .</p><p>= Drymaria ciliata C. A. Mey., Index Seminum (St. Petersburg) 9: 71. 1843. Type: This species was described on the basis of cultivated plants known in the horticultural trade.</p><p>= Drymaria cordata var. pilosa Schltdl., Linnaea 26: 374. 1853. Type: Mexico. This variety was described in a treatment of miscellaneous Mexican plants.</p><p>= Drymaria stylosa Backer, Bull. Jard. Bot. Buitenzorg, sér. 2 12: 15. 1913. Type: Indonesia: many localities in the mountains, s. d., C. A. Backer s. n. (without information about type specimens) .</p><p>= Drymaria tepicana M. E. Jones, Contr. W. Bot. 15: 124. 1929. Type: Mexico: Nayarit: Tepic, 16 February 1927, M. E. Jones 22847 (holotype: POM [162845/0002390 image!]) .</p><p>≡ Drymaria villosa f. tepicana (M. E. Jones) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 227. 1961.</p><p>= Drymaria barrancae M. E. Jones, Contr. W. Bot. 18: 65. 1933. Type: Mexico. Jalisco: La barranca, Guadalajara, 17 November 1930, M. E. Jones 27051 a (holotype POM [192857/0002400 image!]; isotype GH 00037718 image!]) .</p><p>Type.</p><p>Mexico. Veracruz: Jalapa, without date, C. J. W. Schiede s. n. (lectotype, designated by Athira and Maya 2025, pg. 197: LE [LE 00018305 image!]).</p><p>Distribution.</p><p>Subspecies of wide distribution throughout America and Asia.</p><p>Notes.</p><p>Drymaria villosa Cham. &amp; Schltdl., originally described in 1830, is equivalent to Drymaria cordata var. villosa (Cham. &amp; Schltdl.) Rohrb. from 1872. In 1961, Duke referred to a specimen at the Komarov Herbarium (LE) as an “ isotype, ” but according to Article 9.10 of the Shenzhen Code, this should be considered a lectotype. Arya et al. (2024) accepted this correction but incorrectly designated a specimen housed at LECB as the lectotype. However, Chamisso’s original material was transferred to LE (Komarov Institute) in 1840, not to LECB, making their lectotypification invalid (Berger 2018). Athira and Maya (2025) correctly designated specimen LE 00018305 from the Komarov Herbarium in St. Petersburg as the lectotype, as it matches Schlechtendal’s original description, ensuring nomenclatural stability. This approach follows Athira and Maya (2025) in recognizing LE 00018305 as the lectotype of D. villosa Schltdl. &amp; Cham. Finally, although recent plastome sequencing and phylogenetic studies place D. villosa within Caryophyllaceae (Chetri et al. 2024), these molecular data do not affect the nomenclatural typification.</p></div>	https://treatment.plazi.org/id/35969A6DE94B5038ACBE2BE167A97A33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
FB22C2C3170F5081ADDC6C0732389DD7.text	FB22C2C3170F5081ADDC6C0732389DD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria villosa subsp. palustris (Schltdl. & Cham.) J. A. Duke	<div><p>54 b. Drymaria villosa subsp. palustris (Schltdl. &amp; Cham.) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 227. 1961.</p><p>≡ Drymaria palustris Schltdl. &amp; Cham., Linnaea 5: 232. 1830. Type: Mexico. Veracruz: In paludosis prope Jalapam, s. d., C. J. W. Schiede &amp; F. Deppe 504 (lectotype, designated by Duke 1961, pg. 227: LE [LE 00001733 image!]; isolectotype LE [LE 00001732 image!]) .</p><p>≡ Drymaria cordata var. palustris (Schltdl. &amp; Cham.) Rohrb., Fl. Bras. 14 (2): 260. 1872.</p><p>= Drymaria pauciflora Bartl. in Presl, Reliq. Haenk. 2 (1): 8. 1831. Type: Peru: in Peruviae montanis huanoccensibus, 1822, T. Haenke s. n. (lectotype, designated here: GOET [GOET 000582 image!]; isolectotype: PR [24306 image!]) .</p><p>= Drymaria townsendii B. L. Rob., Bot. Gaz. 30 (1): 58. 1900. Type: Mexico. Chihuahua: Sierra Madre, 5 miles southeast of Colonia Garcia, 7400 ft [alt. 2255.5 m], 5 August 1899, C. H. T. Townsend &amp; C. M. Barber 231 (holotype: GH [00037745 image!]; isotypes: E [E 00326057 image!], F [102975 / F 0053306 F image!], K [K 000471738 image!], MO [3578839 / 144037 image!], NDG [16755 image!], NY [00342510 image!], US [735652/00103368 image!], US [00931459 image!]) .</p><p>≡ Drymaria villosa f. townsendii (B. L. Rob.) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 228. 1961.</p><p>= Drymaria nummularia Briq., Annuaire Conserv. Jard. Bot. Genève 13: 371. 1911. Type: Mexico. [Michoacán]: cordillere d’ Ario, 4000, Jun. – Oct. 1840, H. G. Galeotti 4416 (holotype: G [G 00226781 image!]; isotype: K [3862 / K 000471748 image!]) .</p><p>= Drymaria subsessilis M. E. Jones, Contr. W. Bot. 15: 125. 1929. Type: Mexico. Nayarit: in rivulets at Ixtlan, 19 February 1927, M. E. Jones 22848 (holotype: POM [162843/0002391 image!]; isotype: F [705822 / F 0053303 F image!]) .</p><p>= Drymaria sphagnophila Baehni &amp; J. F. Macbr., Publ. Field Mus. Nat. Hist., Bot. Ser. 13 (2 / 2): 625. 1937. Type: Peru. Huánuco: Mito, 9000 ft [alt. 2743.2 m], 8–22 July 1922, J. F. MacBride 1542 (holotype F n. v.; isotype: US [1779692 / 00103362 image!]) .</p><p>= Drymaria villosa subsp. palustris var. perennis J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 228. 1961. Type: México. San Luis Potosí: Álvarez, s. d., E. Palmer 187 (holotype: MO n. v.) .</p><p>Distribution.</p><p>Mexico, Guatemala, Honduras, El Salvador, Costa Rica, Panama, Venezuela, Colombia, Ecuador, Peru, Bolivia.</p><p>Notes.</p><p>Most of the type specimen deposited at B were destroyed in World War II (Hiepko 1987), and Duke (1961) mentioned that the holotype of Drymaria palustris at B was probably destroyed and designated as the lectotype the specimen cited above. This sheet bears an original label written by Chamisso stating “ Drymaria palustris n. sp. ” (handwriting confirmed by Denise Marx, HAL). However, the number “405” does not correspond to the number given in the protologue, which is “404.” Despite this discrepancy, apparently due to a labeling error, we believe that this specimen is original material and thus agree with Duke’s lectotypification. Although there are two specimens at LE, only one was labeled by Duke ([00001733]). We treat the other ([00001732]) as an isolectotype.</p><p>For the same reason as mentioned for Drymaria apetala, we designate the specimen in GOET as the lectotype of Drymaria pauciflora .</p></div>	https://treatment.plazi.org/id/FB22C2C3170F5081ADDC6C0732389DD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
D0C95C4C83885546AA740FCDC89F0EC2.text	D0C95C4C83885546AA740FCDC89F0EC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria villosa subsp. paramorum (S. F. Blake) J. A. Duke	<div><p>54 c. Drymaria villosa subsp. paramorum (S. F. Blake) J. A. Duke, Ann. Missouri Bot. Gard. 48 (3): 230. 1961.</p><p>≡ Drymaria paramorum S. F. Blake, Contr. U. S. Natl. Herb. 20: 521. 1924. Type: Venezuela. Trujillo: paramo de la cristalina, alt. 2900 m, 20 October 1910, A. Jahn 111 (holotype: US [602305/00103355 image!]) .</p><p>Distribution.</p><p>Colombia, Venezuela.</p></div>	https://treatment.plazi.org/id/D0C95C4C83885546AA740FCDC89F0EC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
5F4ACD1B4E8358A6802DAF0EEE472049.text	5F4ACD1B4E8358A6802DAF0EEE472049.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria viscosa S. Watson	<div><p>55. Drymaria viscosa S. Watson, Proc. Amer. Acad. Arts 22 (2): 469. 1887.</p><p>Type.</p><p>Mexico. Baja California: Socorro (“Socono”, “Socario”) Northern Lower California, 27 April 1886, C. R. Orcutt 1330 (holotype: GH [00037746 image!]; isotypes: F [208450 / F 0053282 F image!, 147780/0053283 F image!, 219882/0053284 F image!], ILL [ILL 00007352 image!], MICH [1115004 image!], MIN [410502/1002886 image!], MO [3578903 / 144036 image!], NY [00342511 image!, 00342512 image!], OS [173559/0000138 image!], UC [UC 8304 image!], US [1391067 / 00931429 image!, 10383/00103370 image!], VT [UVMVT 024429 image!, 024430 image!], YU [YU.001066 image!]) .</p><p>Distribution.</p><p>United States, Mexico.</p><p>Notes.</p><p>In Duke (1961), this species is cited as Drymaria viscosa S. Watson ex Orcutt. However, Orcutt did not describe the species nor attribute it to Watson, and, although his specimen represents the holotype, it is likely that Watson designated the epithet independently due to the presence of glandular trichomes that give it viscosity (Parfitt and Hodgson 1985).</p></div>	https://treatment.plazi.org/id/5F4ACD1B4E8358A6802DAF0EEE472049	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
644B4AF6149D534E845518DB5508836D.text	644B4AF6149D534E845518DB5508836D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria Willd.	<div><p>1. Drymaria Willd. in Roem. &amp; Schult., Syst. Veg. 5: 406. 1819. Type (designated by Fosberg 1945, pg. 96): Drymaria cordata (L.) Willd.</p><p>Pinosia Urb., Arkiv för Botanik 23 A (5): 70. 1930. Type: Pinosa ortegioides (Griseb.) Urb. (= Drymaria ortegioides Griseb.)</p><p>Mollugophytum M. E. Jones, Contr. W. Bot. 18: 35 (1933). Type, designated here: Mollugophytum holosteoides (Benth.) M. E. Jones (= Drymaria holosteoides Benth.)</p><p>Notes.</p><p>Although authorship of Drymaria is often given as “ Willd. ex Schult., ” the name and description occur in a single paragraph followed by “ Reliq. Willd. MS. ” This represents direct association of Willdenow with both the name and the description, and Willdenow should be cited as author (see Turland et al. (2018), art. 46.3 example 15). Drymaria arenarioides has been stated to be the type of Drymaria (e. g., Duke 1961, Hernández-Ledesma et al. 2015), with Britton and Millspaugh (1920) credited with having made the designation. However, according to Richard Rabeler (pers. comm. 2025), Britton and Millspaugh followed the largely mechanical method of choosing the first species as the type, and their choice is superseded by Fosberg’s designation of Drymaria cordata as the type (see Turland et al. (2018) art. 10.7), which was made in the context of discussing morphological characters to evaluate the status of Mollugophytum and did not follow the mechanical method.</p><p>Mollugophytum holosteoides is here designated the type because it is the most widespread species of those treated by Jones (1933) within Mollugophytum .</p></div>	https://treatment.plazi.org/id/644B4AF6149D534E845518DB5508836D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
A7D34945AC435BBE99E0BFBB0C662BA2.text	A7D34945AC435BBE99E0BFBB0C662BA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drymaria xerophylla A. Gray	<div><p>56. Drymaria xerophylla A. Gray, Smithsonian Contr. Knowl. 5 (6): 18. 1853.</p><p>Type.</p><p>Mexico. Without data, T. Coulter 722 (holotype: GH [00218128 image!]; isotype: NY [00342513 image!]) .</p><p>Distribution.</p><p>Mexico.</p></div>	https://treatment.plazi.org/id/A7D34945AC435BBE99E0BFBB0C662BA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Valentín-Martínez, Dagoberto;Steinmann, Victor W.;Flores-Olvera, Hilda;Borsch, Thomas;Montesinos-Tubée, Daniel B.	Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas, Montesinos-Tubée, Daniel B. (2025): A nomenclator of Drymaria (Caryophyllaceae). PhytoKeys 266: 139-175, DOI: 10.3897/phytokeys.266.162517
