taxonID	type	description	language	source
04D0C3A351C65D268C90A544C0E81A76.taxon	description	Figs 2 A – L, 8 L	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
04D0C3A351C65D268C90A544C0E81A76.taxon	diagnosis	Diagnosis. The male tegmina maculae of this species most closely resemble those of P. wooi (Qiu, Che & Wang, 2018). However, in natural posture, the medial black maculae on both tegmina are fused in this species, and the abdomen (excluding the terminal segment) is light yellowish brown. In contrast, P. wooi exhibits non-fused medial black maculae on the tegmina in natural posture, with the abdomen (excluding the terminal segment) being dark yellowish brown. Furthermore, females of this species possess four symmetrical yellow maculae near the anterior margin on both the mesonotum and metanotum, whereas P. wooi females bear only two conspicuous symmetrical yellow maculae solely on the metanotum.	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
04D0C3A351C65D268C90A544C0E81A76.taxon	description	Description. Holotype. Measurements (mm). Overall length (including tegmen): 26.27; body length: 17.82; body width (tegmina not included): 8.46; tegmen length × width: 21.73 × 7.74; pronotum length × width: 7.42 × 4.85. Coloration. Pronotum black, anterior margin white. Tegmina and hind wings yellowish brown, maculae black (Fig. 2 A, E). Eyes, vertex, post-clypeus and spaces between ocelli black. Ocelli and antennal sockets white. Ante-clypeus yellowish white. Antennae and labrum yellowish brown. Labial palpi brown (Fig. 2 G). Legs black brown, spines yellowish brown to black. Pulvilli and arolia white. Sterna yellowish brown, margins and distal part black (Fig. 2 B). Body. Head: Sub-rounded, nearly completely hidden under pronotum. Eyes and ocelli well-developed. Interocular space narrower than the distance between ocelli, the latter narrower than the distance between antennal sockets (Fig. 2 G). Pronotum: Oval-shaped, widest near the middle. Surface densely covered with short setae and long pubescence, central portion bearing symmetrical black stripes. Anterior whitish margin narrow, clearly delineated from black areas (Fig. 2 E). Tegmina and hind wings: Densely covered with black maculae, markings on the basal lateral margins and mid-region more densely distributed than other areas (Fig. 2 A). Legs: Slender, front femur Type C 1. Pulvilli and arolia present (Fig. 2 B). Abdomen: Smooth. Supra-anal plate transverse, pubescent, posterior margin protruded medially. Paraprocts simple (Fig. 2 I). Subgenital plate with short setae, hind margin slightly asymmetric. Styli thin and long (Fig. 2 J). Genitalia: Well-sclerotized. L 1 apically bearing two short branches; the two basal branches distinct and elongated. L 2 arcuately curved. Genital hook (L 3) medially swollen, tapering toward hooked apex. L 4 M broadly lamellate. L 4 N well-developed; pda and paa strongly curved. L 8 irregular, narrow and plate-like. R 1 M stoutly expanded terminally. R 1 L elongate and banded. R 2 divided into two chunks. R 3 broadly concave (Fig. 2 K, L). Female paratype (mm). Body length: 20.54; body width: 12.02; pronotum length × width: 8.91 × 5.58. Coloration. Terga reddish brown to blackish brown, with four symmetrical yellow maculae at the anterior margin of the meso- and metanotum (Fig. 2 C). Pronotum dark reddish brown, pubescence light reddish brown (Fig. 2 F). Vertex, eyes, space between ocelli and post-clypeus black. Antennal sockets white. Antennae yellowish brown. Ocelli, ante-clypeus, and basal part of labrum pale yellow. Middle and distal part of labrum yellowish brown. Legs dark brown, spines reddish brown to black. Sterna brown to yellowish brown. Subgenital plate black (Fig. 2 D, H). Body. The widest point of pronotum near the hind margin. Anterior whitish margin absent, bearing symmetrical black stripe medially (Fig. 2 F). Ocelli degraded to two spots. Interocular space almost equal to the distance between antennal sockets, both wider than the distance between ocelli. Front femur Type C 1. Arolia and pulvilli absent (Fig. 2 D, H). Nymph. Pronotum also with four symmetrical yellow maculae, the rest similar to the female. Ootheca. Unknown.	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
04D0C3A351C65D268C90A544C0E81A76.taxon	etymology	Etymology. This species is named after the collector, Mr Quan-Fu Duan, in recognition of his contribution to its discovery.	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
F7002A181F5E5E719F739A102C1B1695.taxon	description	Figs 3 A – L, 8 K, 10 A, B, D – G	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
F7002A181F5E5E719F739A102C1B1695.taxon	diagnosis	Diagnosis. This species exhibits highly similar tegmina maculae in males and dorsal maculae in females to those of P. daweishana. The primary diagnostic differences are: (1) Males of this species possess a light yellowish-brown abdomen, whereas P. daweishana males have a nearly uniform black abdomen; 2) The anterior whitish margin on the male pronotum is extremely narrow in this species, contrasting with the significantly wider counterpart in P. daweishana males; and (3) The sclerite paa of this species is broad with a short finger-like projection at the right posterior margin, while in P. daweishana, the paa bears a long finger-like projection.	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
F7002A181F5E5E719F739A102C1B1695.taxon	description	Description. Holotype. Measurements (mm). Overall length (including tegmen): 25.63; body length: 17.26; body width (tegmina not included): 8.75; tegmen length × width: 23.02 × 8.62; pronotum length × width: 7.73 × 4.64. Coloration. Body dark brown (Fig. 3 A, B). Pronotum reddish brown, with yellowish-brown pubescence along the outer margin, anterior margin partially yellow (Fig. 3 E). Tegmina and hind wings yellowish brown, with dark brown maculae (Fig. 3 A, B). Vertex, eyes, and spaces between ocelli black. Ocelli and antennal sockets white. Antennae yellow, ante-clypeus pale yellow, post-clypeus black. Labrum yellowish brown, middle part dark brown (Fig. 3 G). Legs dark brown, spines yellowish brown to black. Pulvilli and arolia white. Sterna dark brown (Fig. 3 B). Body. Head: Sub-rounded, almost entirely hidden under pronotum. Eyes and ocelli well-developed. Ocelli ridge indistinct, bearing a row of slender setae along the upper edge. Interocular space narrower than the distance between ocelli, the latter narrower than the distance between antennal sockets (Fig. 4 G). Pronotum: Oval-shaped, widest near the middle. Surface densely covered with short setae and long pubescence, with symmetrical black stripes medially. Anterior whitish margin extremely narrow, medially almost interrupted (Fig. 3 E). Tegmina and hind wings: Tegmina densely covered with small and diffuse maculae; maculae at basal and medial regions more densely distributed than at apical region (Fig. 3 A, B). Legs: Slender, front femur Type C 1. Pulvilli and arolia present (Fig. 3 B). Abdomen: Smooth, light yellowish brown. Supra-anal plate narrow, distinctly pubescent. Cerci slender. Subgenital plate slightly asymmetrical. Styli long. Genitalia: Well-sclerotized. L 1 apically swollen and bearing one short branch that pointed to the top right corner; the two basal branches thin and elongated. L 2 arcuately curved. Genital hook (L 3) straight and short. L 4 M broadly lamellate. L 4 N well-developed; pda narrow, paa wide and with a finger-like protrusion at the right bottom. L 8 plate-like. R 1 M stoutly expanded terminally. R 1 L elongate and thin. R 2 divided into two chunks. R 3 broadly concave (Fig. 3 K, L). Female (pre-adult) paratype (mm). Body length: 17.72; body width: 10.56; pronotum length × width: 8.23 × 4.90. Coloration. Terga yellow to brownish yellow, densely covered with brownish-yellow maculae, median with a black-brown longitudinal line (Fig. 3 C). Pronotum brownish yellow, densely covered with short setae (Fig. 3 F). Vertex, eyes, and the space between ocelli and post-clypeus are black. Antennal sockets and ante-clypeus white. Ocelli and the areas surrounding the ocelli and antennal sockets yellowish white. Antennae yellowish brown. Labrum blackish brown. Legs blackish brown, with spines ranging from yellowish brown to black. Sterna yellow, densely covered with brownish-yellow maculae, median with a wide longitudinal yellow line. Middle part of subgenital plate black (Fig. 3 D, H). Body. The widest point of pronotum near the hind margin. Anterior whitish margin absent (Fig. 3 F). Ocelli degraded to two spots. Interocular space almost equal to the distance between antennal sockets, both wider than the distance between ocelli. Front femur Type C 1. Arolia and pulvilli absent (Fig. 3 D, H). Female. Unknown Nymph. Similar to the pre-adult female. Ootheca. Unknown.	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
F7002A181F5E5E719F739A102C1B1695.taxon	etymology	Etymology. The name of this species comes from the type locality, Menglian County, Pu’er City, Yunnan Province.	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
74819DD952CD5889986B6D27AEED1DBF.taxon	description	Figs 6 A – L, 8 C, F, 9 A – E	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
80CD9127DC8D521695E630C195288935.taxon	description	Figs 4 A – L, 8 B, E, G, H	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
80CD9127DC8D521695E630C195288935.taxon	diagnosis	Diagnosis. The males of this species resemble certain Pseudoeupolyphaga congeners with densely maculate tegmina, such as Pseudoeupolyphaga daweishana (Qiu, Che & Wang, 2018), Pseudoeupolyphaga hengduana (Woo & Feng, 1992), and Pseudoeupolyphaga fengi yongshengensis (Qiu, Che & Wang, 2018). However, it is readily distinguished from P. daweishana and P. hengduana by the conspicuously sparser maculae in the anal field of the tegmina compared to adjacent areas. Furthermore, it differs from P. fengi yongshengensis by the absence of a longitudinal yellow stripe on the mid-ventral sternum — a diagnostic trait presents in the latter. Additional key diagnostic characters include: denser maculae on the tegmina of P. spelunca males, concentrated primarily basally and medially, but sparse apically. In contrast, the three congeneric males exhibit more uniformly distributed tegmina maculae without such pronounced regional differentiation.	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
80CD9127DC8D521695E630C195288935.taxon	description	Description. Holotype. Measurements (mm). Overall length (including tegmen): 24.68; body length: 18.74; body width (tegmina not included): 9.40; tegmen length × width: 20.83 × 8.58; pronotum length × width: 7.73 × 4.14. Coloration. Body dark brown (Fig. 4 A, B). Pronotum black, outer margin with pale yellow pubescence, anterior margin yellow (Fig. 4 E). Tegmina yellowish brown, with large dark brown patches at the basal and middle region; remaining areas with evenly distributed dark brown markings. Hind wings pale yellow, bearing dark brown plaques (Fig. 4 A, B). Face black. Antennae yellow. Eyes black. Ocelli white. Ante-clypeus pale yellow, post-clypeus dark brown. Labrum yellow (Fig. 4 G). Legs dark brown. Pulvilli and arolia white. Abdomen dark brown, gradually deepening in color toward the lateral and distal end (Fig. 4 B). Body. Head: Sub-rounded, hidden under pronotum. Eyes and ocelli well-developed. Ocellar ridge slightly curved, bearing a row of slender setae along the upper edge. Interocular space narrower than the distance between ocelli, the latter narrower than the distance between antennal sockets. Clypeus developed (Fig. 4 G). Pronotum: Oval-shaped, widest near the middle. Surface densely covered with short setae and long pubescence, central portion bearing symmetrical black stripes. Anterior whitish margin extremely narrow, disconnect at the middle of the front margin (Fig. 4 E). Tegmina and hind wings: Basal half of tegmina nearly completely covered by black patches except anal area. Anal area and distal half of tegmina bearing variably sized markings (Fig. 4 A, B). Legs: Slender, front femur Type C 1. Pulvilli and arolia present (Fig. 4 B). Abdomen: Supra-anal plate transverse, pubescent, posterior margin slightly protruded medially. Paraprocts simple (Fig. 4 I). Subgenital plate with short setae, hind margin concave in the middle, the left side less prominent than the right side. Styli short and small (Fig. 4 J). Genitalia: Well-sclerotized. L 1 apically swollen and bearing two short branches; the two basal branches distinct and elongated. L 2 arcuately curved. Genital hook (L 3) medially swollen, tapering toward hooked apex. L 4 M broadly lamellate. L 4 N well-developed, pda and paa strongly curved. L 8 irregular, plate-like. R 1 M stoutly expanded terminally. R 1 L elongate and banded. R 2 divided into two chunks. R 3 broadly concave (Fig. 4 K, L). Female paratype (mm). Body length: 21.92; body width: 12.05; pronotum length × width: 8.64 × 4.87. Coloration. Terga deep reddish brown (Fig. 4 C). Vertex black. Face yellow. Antennae yellow. Ocelli yellowish white, space between ocelli, antennal sockets and post-clypeus brown to dark brown. Ante-clypeus and labrum pale yellowish brown. Post-clypeus yellowish brown (Fig. 4 H). Legs dark brown overall, tarsus slight pale. Spines on legs yellowish brown, terminal nearly black. Sterna uniformly dark brown and middle part with yellow strips (Fig. 4 D). Body. Pronotum widest near the hind margin, middle part with symmetrical black dark stripes, anterior whitish margin indistinct (Fig. 4 F). Ocelli big but not prominent, presenting white spots in a near-triangular shape. The distance between ocelli narrower than the distance between antennal sockets, both measurements less than interocular space (Fig. 4 H). Front femur Type C 1. Arolia and pulvilli absent. Nymph. Similar to the female. Ootheca. Reddish brown, surface with parallel and dense longitudinal lines. Ridges of serrated protuberances slightly small. No respiratory canals (Fig. 8 B, E).	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
80CD9127DC8D521695E630C195288935.taxon	etymology	Etymology. The species epithet is derived from the Latin term spelunc, which refers to natural caves, indicating it was found in dark subterranean environments (a few hundred meters from the cave entrance).	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
D92A881810A958AB9BEED66914A22292.taxon	description	Figs 7 A – D, 10 C	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
D1C8B03AB7B35737B7A100E83BE1B155.taxon	description	Figs 5 A – L, 8 A, D, I, J	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
D1C8B03AB7B35737B7A100E83BE1B155.taxon	diagnosis	Diagnosis. The male of this species resembles Pseudoeupolyphaga deficiens Han, Che & Wang, 2024, but differs in having a narrower anterior whitish margin in the pronotum. In addition, the abdominal tergites of females are uniformly orange-red (observation of two samples), making it easy to distinguish from most congeners.	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
D1C8B03AB7B35737B7A100E83BE1B155.taxon	description	Description. Holotype. Measurements (mm). Overall length (including tegmen): 26.30; body length: 19.48; body width (tegmina not included): 10.17; tegmen length × width: 22.77 × 9.30; pronotum length × width: 8.63 × 4.06. Coloration. Pronotum deep reddish brown, setae yellowish, anterior whitish margin yellow (Fig. 5 A, E). Tegmina pale yellow, interspersed with varying-sized blackish-brown maculae (Fig. 5 A). Eyes, vertex, and space between ocelli black. Ocelli pale yellow. Antennal sockets, and ante-clypeus yellowish brown. Post-clypeus, labrum, labial palpi and maxillary palpi blackish brown (Fig. 5 G). Legs yellowish brown, with spines ranging from reddish brown to black. Pulvilli and arolia white. Sterna pale yellow, middle and distal part gray to pale black (Fig. 5 B). Body. Head: Sub-rounded. Eyes and ocelli well-developed. Interocular space narrower than the distance between ocelli, the latter narrower than the distance between antennal sockets. Clypeus developed (Fig. 5 G). Pronotum: Elliptical, widest near the posterior margin. Surface densely covered with short setae, bearing symmetrical black stripe medially. Anterior whitish margin clearly delineated from deep reddish-brown areas (Fig. 5 E). Tegmina and hind wings: Tegmina with dark brown patch confluent basally along outer margin. Remaining areas sparsely scattered with variably sized blackish- brown maculae (Fig. 5 A). Legs: Slender, front femur Type C 1. Pulvilli and arolia present (Fig. 5 B). Abdomen: Supra-anal plate transverse, pubescent, posterior margin protruded medially. Paraprocts simple (Fig. 5 I). Subgenital plate bearing short setae, posterior margin slightly curved. Styli short and small (Fig. 5 J). Genitalia: Well-sclerotized. L 1 apically dilated, basally bifurcated. L 2 arched curved, distally dilated. Genital hook (L 3) thin and straight, curved hook section small. L 4 M broad lamellate. L 4 N developed, pda and paa broad and curved. L 8 irregular, lamellate, bearing a long clavate process. R 1 M stoutly expanded terminally. R 1 L elongate and banded. R 2 divided into two chunks, narrowly spaced, with rounded margins. R 3 broadly concave (Fig. 5 K, L). Paratype. Measurements (mm). Body length: 20.08; body width: 12.43; pronotum length × width: 9.14 × 5.28. Coloration. Terga orange-red (Fig. 5 C). Head dark brown. Antennae yellow. Ocelli white. Ante-clypeus yellowish brown. Post-clypeus dark brown. Labrum with a white patch basally, remaining portions yellowish brown (Fig. 5 H). Legs reddish brown. Spines on the leg reddish brown, terminal nearly black. Sterna yellowish brown to orange-red, subgenital plate black (Fig. 5 D). Body. The widest point of pronotum near the hind margin, middle part with symmetrical dark-black stripe, anterior yellow margin indistinct (Fig. 5 F). Ocelli indistinct, degenerated to two patches. The distance between ocelli narrower than the distance between antennal sockets, and the latter narrower than interocular space (Fig. 5 H). Front femur Type C 1. Arolia and pulvilli absent. Nymph. Similar to female, only the coloration is slightly paler. Ootheca. Reddish brown, surface with many parallel, dense longitudinal lines. Ridges of serrated protuberances with broad bases, slightly curved lateral edges and blunt tips. No respiratory canals (Fig. 8 A, D).	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
D1C8B03AB7B35737B7A100E83BE1B155.taxon	etymology	Etymology. The species epithet is derived from the Latin word vest, denoting its tegmina markings resembling a vest.	en	Ren, Yi-Ming, Han, Wei, Che, Yan-Li, Wang, Ji-Rui (2025): Four new species of the genus Pseudoeupolyphaga Qiu & Che, 2024 (Blattodea, Corydioidea, Corydiinae) from Yunnan, China. ZooKeys 1261: 337-358, DOI: 10.3897/zookeys.1261.168015
