taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
536F827BFFA353122AE1FB7AF3B2F5FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774885/files/figure.png	https://doi.org/10.5281/zenodo.14774885	Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth.	Figure 5. Shells, operculum, and radulae of Pseudamnicola araujoi. A, B, holotype (MNCN 15.05/200505H). C–J, paratypes (UGSB 17299). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon.I, central radular teeth. J, outer marginal teeth.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFA353122AE1FB7AF3B2F5FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774887/files/figure.png	https://doi.org/10.5281/zenodo.14774887	Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland.	Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFA353122AE1FB7AF3B2F5FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774891/files/figure.png	https://doi.org/10.5281/zenodo.14774891	Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth.	Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFA353122AE1FB7AF3B2F5FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774893/files/figure.png	https://doi.org/10.5281/zenodo.14774893	Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland.	Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFA353122AE1FB7AF3B2F5FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774875/files/figure.png	https://doi.org/10.5281/zenodo.14774875	Figure 2. Morphological variation among Pseudamnicola species. Plots in the left and right columns display the results for the unsupervised model-based clustering and canonical variate analysis (CVA), respectively, based on four different datasets: A, principal components of shell shape; B, shell measurements; C, D, female and male genitalia measurements. The number of analyzed individuals (N) is indicated on the left side. Symbols and numbers on the right represent species assignments and correspond to those used in Figure 1. Abbreviations shown in images are explained in the Materials and methods section.	Figure 2. Morphological variation among Pseudamnicola species. Plots in the left and right columns display the results for the unsupervised model-based clustering and canonical variate analysis (CVA), respectively, based on four different datasets: A, principal components of shell shape; B, shell measurements; C, D, female and male genitalia measurements. The number of analyzed individuals (N) is indicated on the left side. Symbols and numbers on the right represent species assignments and correspond to those used in Figure 1. Abbreviations shown in images are explained in the Materials and methods section.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFA0530C29E0FD6BF054F176.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774891/files/figure.png	https://doi.org/10.5281/zenodo.14774891	Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth.	Figure 7. Shells, operculum, and radulae of Pseudamnicola argentariensis. A, holotype (MNCN 15.05/200500H). B–J, paratypes (UGSB 11999). A–D, shells. E, F, operculum (E, inner side; F, outer side). G, protoconch. H, portion of radula ribbon. I, central radular teeth. J, inner and outer marginal teeth.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFA0530C29E0FD6BF054F176.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774893/files/figure.png	https://doi.org/10.5281/zenodo.14774893	Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland.	Figure 8. Anatomy of Pseudamnicola argentariensis. A–H, paratypes (UGSB 11999). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis.H, prostate gland.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFA0530C29E0FD6BF054F176.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774895/files/figure.png	https://doi.org/10.5281/zenodo.14774895	Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988).	Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988).	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFA0530C29E0FD6BF054F176.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774897/files/figure.png	https://doi.org/10.5281/zenodo.14774897	Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland.	Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFBE53092866F9EAF0E1F494.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774895/files/figure.png	https://doi.org/10.5281/zenodo.14774895	Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988).	Figure 9. Shells, operculum, and radulae of Pseudamnicola bodoni. A, holotype (MNCN 15.05/200502H). B, paratype (UGSB 988). C, shell (UGSB 5697). D, shell (UGSB 16628). E, F, operculum (E, inner side; F, outer side) (UGSB 988). G, protoconch (UGSB 988). H, portion of radula ribbon (UGSB 988). I, central radular teeth (UGSB 988). J, inner and outer marginal teeth (UGSB 988).	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFBE53092866F9EAF0E1F494.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774897/files/figure.png	https://doi.org/10.5281/zenodo.14774897	Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland.	Figure 10.Anatomy of Pseudamnicola bodoni. A–H, paratypes (UGSB 988). A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis.G, penis. H, prostate gland.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFBB530B2995FC42F3CBF145.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774901/files/figure.png	https://doi.org/10.5281/zenodo.14774901	Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P).	Figure 11. Shells, operculum, and radulae of Pseudamnicola megastomus. A, B, holotype (MNCN 15.05/200501H). C, paratype (MNCN 15.05/200501P). D, shell (MNCN 15.05/95107).E, operculum (SEM photograph; outer side) (MNCN 15.05/200501P).F, protoconch (MNCN 15.05/200501P).G, portion of radula ribbon (MNCN 15.05/200501P). H, central radular teeth (MNCN 15.05/200501P). I, outer marginal teeth (MNCN 15.05/200501P).	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFBB530B2995FC42F3CBF145.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774903/files/figure.png	https://doi.org/10.5281/zenodo.14774903	Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland.	Figure 12. Anatomy of Pseudamnicola megastomus. A–H, paratypes (MNCN 15.05/200501P).A, ctenidium and osphradium. B, stomach. C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle. F, head of male and penis. G, penis. H, prostate gland.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFBB530B2995FC42F3CBF145.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774875/files/figure.png	https://doi.org/10.5281/zenodo.14774875	Figure 2. Morphological variation among Pseudamnicola species. Plots in the left and right columns display the results for the unsupervised model-based clustering and canonical variate analysis (CVA), respectively, based on four different datasets: A, principal components of shell shape; B, shell measurements; C, D, female and male genitalia measurements. The number of analyzed individuals (N) is indicated on the left side. Symbols and numbers on the right represent species assignments and correspond to those used in Figure 1. Abbreviations shown in images are explained in the Materials and methods section.	Figure 2. Morphological variation among Pseudamnicola species. Plots in the left and right columns display the results for the unsupervised model-based clustering and canonical variate analysis (CVA), respectively, based on four different datasets: A, principal components of shell shape; B, shell measurements; C, D, female and male genitalia measurements. The number of analyzed individuals (N) is indicated on the left side. Symbols and numbers on the right represent species assignments and correspond to those used in Figure 1. Abbreviations shown in images are explained in the Materials and methods section.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFBB530B2995FC42F3CBF145.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774883/files/figure.png	https://doi.org/10.5281/zenodo.14774883	Figure 4. Species validation by IBPP based on the species-tree topology from DELINEATE as the guide tree. Posterior probabilities of classification support are shown as three coloured boxes above branches for analyses with morphological linear measurements alone (left box), with mitochondrial and nuclear gene sequences (middle box), and with all linear measurements and gene sequences (right box). Tips group the populations belonging to a particular species and are connected to our proposed taxonomic name.These are followed by the number of DNA-based methods that support each depicted species grouping.Pie charts on the right provide the percentage of individuals classified within a delimited species according to the canonical variate analysis of shell shape, shell linear measurements, and measurements of the female and male genitalia.According to DNA-sequence data, populations marked with an asterisk were lumped into a single species, but given their high morphological differentiation, they are now considered three separate species.Abbreviation: ND, no data available.	Figure 4. Species validation by IBPP based on the species-tree topology from DELINEATE as the guide tree. Posterior probabilities of classification support are shown as three coloured boxes above branches for analyses with morphological linear measurements alone (left box), with mitochondrial and nuclear gene sequences (middle box), and with all linear measurements and gene sequences (right box). Tips group the populations belonging to a particular species and are connected to our proposed taxonomic name.These are followed by the number of DNA-based methods that support each depicted species grouping.Pie charts on the right provide the percentage of individuals classified within a delimited species according to the canonical variate analysis of shell shape, shell linear measurements, and measurements of the female and male genitalia.According to DNA-sequence data, populations marked with an asterisk were lumped into a single species, but given their high morphological differentiation, they are now considered three separate species.Abbreviation: ND, no data available.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFBB530B2995FC42F3CBF145.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774887/files/figure.png	https://doi.org/10.5281/zenodo.14774887	Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland.	Figure 6. Anatomy of Pseudamnicola araujoi. A–H, paratypes (UGSB 17299). A, ctenidium and osphradium. B, stomach.C, partial nervous system.D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, G, head of male and penis. H, prostate gland.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFB95304299BF913F372F1D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774905/files/figure.png	https://doi.org/10.5281/zenodo.14774905	Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth.	Figure 13. Shells, operculum, and radulae of Pseudamnicola reticulatus. A, holotype (MNCN 15.05/200503H). B–J, paratypes (MNCN 15.05/200503P; UGSB 5971). A–D, shells. E, F, operculum (E, inner side; F, outer side).G, protoconch. H, portion of radula ribbon.I, central radular teeth.J, inner and outer marginal teeth.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFB95304299BF913F372F1D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774907/files/figure.png	https://doi.org/10.5281/zenodo.14774907	Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland.	Figure 14. Anatomy of Pseudamnicola reticulatus. A–H, paratypes (UGSB 5971). A, ctenidium and osphradium. B, stomach. C, partial nervous system. D, pallial oviduct. E, bursa copulatrix and seminal receptacle.F, head of male and penis. G, penis.H, prostate gland.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFB6530629F3F88FF1FCF7E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774909/files/figure.png	https://doi.org/10.5281/zenodo.14774909	Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth.	Figure 15. Shells, operculum, and radulae of Pseudamnicola sardiniensis. A, holotype (MNCN 15.05/200504H). B–F, paratypes (UGSB 15501). A, B, shell.C, protoconch.D, portion of radula ribbon.E, central radular teeth. F, inner and outer marginal teeth.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
536F827BFFB6530629F3F88FF1FCF7E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14774911/files/figure.png	https://doi.org/10.5281/zenodo.14774911	Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland.	Figure 16. Anatomy of Pseudamnicola sardiniensis. A–F, paratypes (UGSB 15501). A, ctenidium and osphradium.B, stomach. C, partial nervous system. D, head of male and penis. E, penis.F, prostate gland.	2024-03-05	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Boulaassafer, Khadija;Khalloufi, Noureddine;Hauffe, Torsten			
