taxonID	type	description	language	source
5368996BFFD9767CFF5FF6F7FB336365.taxon	description	(Figs 1 – 7, 8 A)	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD9767CFF5FF6F7FB336365.taxon	materials_examined	Material examined. Holotype (female on card): INDIA: Karnataka, Madikeri, Kabbinakad, Chingaara estate, 10. i. 2025, ex. Larva of Agrilus sp. on Villebrunea integrifolia Gaud., coll. Hemanth, accession number: NIM / NBAIR / HEYD / HEYD / 10125 - H (NIM). Paratypes: same data as holotype, accession numbers: NIM / NBAIR / HEYD / HEYD / 10125 - P 1 (NIM) ‒ NIM / NBAIR / HEYD / HEYD / 10125 - P 3 (NIM) (2 ♀ and 1 ♂ on three cards). Karnataka, Melinajuganahalli, Doddaballapura, 20. vii. 2025, ex. Larva of Agrilus sp. on Albizia amara (Roxb.) Boivin., coll. Hemanth, accession numbers: NIM / NBAIR / HEYD / HEYD / 20725 - P 4 (NIM) ‒ NIM / NBAIR / HEYD / HEYD / 20725 - P 16 (NIM) (7 ♀ and 6 ♂ on thirteen cards).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD9767CFF5FF6F7FB336365.taxon	diagnosis	Diagnosis. Female, body length 4.4 – 7.5 mm; face with iridescent lanceolate setae (Fig. 2 A); POL 0.95 – 1.20 × OOL; gena uniformly reticulate (Fig. 2 C); antenna with clava subequal to combined length of preceding two segments (Fig. 2 B); mesosoma moderately compressed dorsoventrally (Figs 1 B, 8 A); lateral panels of pronotum angled at about 45 ° to the dorsal surface, with a punctate region as in Fig. 2 F; axillae uniformly reticulate (Fig. 2 F); dorsellum smooth, shiny and punctate dorsally (Fig. 3 A); propodeum distinctly reticulate-punctate (Fig. 3 A) to variably sculptured (Fig. 5 B, C), plical region with median carina distinct only in posterior half, shallowly merging with plical region in anterior half (Figs 3 A, 5 B, C); fore leg with profemur distinctly enlarged with distinct basal ventral angle and minute serrations, protibia with stout spines on dorsal margin (Fig. 2 E); fore wing basal cell with basal setal line present, infuscation near STV broad (Fig. 3 C); petiole conspicuous, 1.4 – 1.5 × longer than wide, Gt 1 and Gt 2 with posterior margins distinctly incised medially (Fig. 3 D); ovipositor sheath 1.9 – 2.0 × length of metatibia (Figs 1 B, 8 A).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD9767CFF5FF6F7FB336365.taxon	description	Description. FEMALE (holotype): body length 6.0 mm, ovipositor sheath length 2.4 mm. Color: mandible black; tegula blackish brown; scape bicolored, rest of the antenna uniformly blackish brown; legs black with basal and apical margins of femora and tibiae faintly yellowish, protarsus honey brown, meso- and metatarsi whitish with apex slightly darker; head greenish with bluish and golden metallic luster, face darker than gena and with lanceolate setae on face silvery and iridescent; mesosoma mostly with greenish blue metallic luster; pronotum anteriorly, mesoscutum in median anterior half between notauli, axillae in anterior half and scutellum with strong dark coppery metallic luster; under strong illumination lateral panels of pronotum, sides of mesoscutum, axillae in posterior half, prepectus, and mesopleuron and metapleuron in posterior half with bluish metallic luster; lateral panel of pronotum posteriorly with dark brownish coloration (Fig. 2 F); dorsellum with golden green metallic luster; propodeum with bluish and golden green metallic luster; wings hyaline and slightly iridescent; fore wing with discal setae and thick tuft of setae at base of parastigma and MV black; fore wing veins with MV and parastigma pale brown, SMV (excluding parastigma), PMV and STV darker; metasoma mostly black with faint purple tinge, with metallic luster on Gt 4 – Gt 6; petiole testaceous and ovipositor sheaths black. Head: mostly reticulate, in frontal view 1.17 × wider than long; face above scrobe, around median ocellus, distinctly shallowly reticulate, scrobe reticulate-strigose, bell shaped; distance between interantennal region 1.66 × distance from torulus to nearest eye margin; parascrobal region, interantennal region and lower face with lanceolate setae; head in dorsal view 1.75 × wider than long; POL subequal to OOL (Fig. 2 D); temple narrow; malar sulcus distinct (Fig. 2 C); antenna with fl 1 transverse, fl 2 shorter than fl 3, fl 3 equal to combined length of fl 1 and fl 2, fl 3 and fl 4 almost equal to each other, rest of the segments decreasing in length towards apex, clava subequal to combined length of preceding two segments (Fig. 2 B). Mesosoma: reticulate-punctate, moderately dorsoventrally compressed (Figs 1 B, 8 A); pronotum long, tent like with lateral panels angled at about 45 ° to the dorsal surface, and smooth and shiny except for a punctate region as in Fig. 2 F; mesoscutum 1.2 × as wider as long, notauli complete and weakly indicated; scutellum 1.12 × as wide as long, reticulation closer and with a differentiated axillula; scutellum without frenum, frenal arm indicated laterally; dorsellum shiny and punctate dorsally (Fig. 3 A), metanotum faintly reticulate laterally; propodeum (Fig. 3 A) with plical region distinctly reticulate-punctate, slightly irregularly rugose anteriorly, with median carina distinct only in posterior half and shallowly merging with cuticle surface in anterior half, and with distinct, almost parallel plicae; propodeal callus anteriorly with two interconnected smooth and shiny broad fovea, with reticulate-punctate region and setation as in Fig. 3 A; propodeal spiracles inside a smooth and shiny area, with distance between spiracles 1.2 × medial length of propodeum; prepectus triangular and setose; mesopleuron ventrally setose, distinctly reticulate-punctate with acropleuron smooth, shiny and faintly aciculate, upper mesepimeron turning aciculate posteriorly (Fig. 3 B); metapleuron punctate and setose; fore wing hyaline except for infuscation near STV, the infuscation narrow anteriorly and broadening towards the posterior wing margin, 3.3 × as long as wide, with costal cell setose, basal cell bare with basal setal line present and cubital setal line incomplete, speculum present (Fig. 3 C), marginal fringe minute, and MV 1.3 × PMV, PMV 2.2 × STV; profemur and metacoxa with aciculate sculpture represented by longitudinally impressed lines; fore leg with profemur distinctly enlarged, ventral margin basally angled, with minute serrations (Fig. 2 E), protibia with distinct stout spines on dorsal margin (one at distal margin) and with protibial spur curved, and basitarsus slightly curved and more conspicuous than rest of tarsal segments; midleg with mesotibial spur longest of all spurs; hind leg with metacoxa with moderately long setae, and metatibia with two unequal spurs. Metasoma: only slightly longer than mesosoma (1.1 ×); petiole oval shaped, 1.45 × as long as wide (Fig. 3 D); Gt 1 – Gt 3 smooth and shiny, Gt 1 and Gt 2 with posterior margins distinctly incised medially; Gt 4 – Gt 6 reticulate, reticulation on Gt 4 weak and faintly visible; ovipositor sheath shorter than metasoma, in lateral aspect 1.9 × length of metatibia (Figs 1 B, 8 A). MALE. Similar to female but body length 4.65 mm; POL 1.1 × OOL (Fig. 4 B); antenna with clava slightly longer (1.2 ×) than combined length of preceding two segments (Fig. 4 C); metasoma with petiole distinctly elongate, 3 × as long as wide (Fig. 4 H), remaining segments 3.5 × medial length of petiole; Gt 1 with posterior margin medially incised, Gt 2 concealed under Gt 1 with posterior margin straight (Fig. 4 G). Variation. Females: body length 4.4 – 7.5 mm (n = 10, including holotype); POL 0.95 – 1.20 × OOL; propodeum with plical region distinctly reticulate-punctate (Fig. 3 A) or with reticulation incompletely / completely reduced with irregular transverse rugae (Fig. 5 B, C), propodeal callus below the level of spiracles distinctly reticulate-punctate (Fig. 3 A) or irregularly rugose with reticulation reduced (Fig. 5 B, C); fore wing 3.1 – 3.3 × as long as wide, basal cell with cubital setal line incomplete to complete, MV 1.2 – 1.4 × PMV, PMV 2.1 – 2.3 × STV; petiole 1.4 – 1.5 × as long as wide, ovipositor sheaths 1.9 – 2 × length of metatibia. Males: body length 2.2 – 5.5 mm (n = 7); all legs including coxae but excluding tarsi blackish (Fig. 1 C) to uniformly testaceous (Fig. 1 D); POL 1.10 – 1.65 × OOL; propodeum with median carina distinct either in posterior half (Fig. 4 D) or in posterior two thirds (Fig. 5 A); fore wing with infuscation near STV broad (Fig. 4 F) or nearly rectangular (Fig. 5 D) to distinctly reduced or absent (Fig. 5 F); petiole 2.85 – 3.10 × as long as wide.	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD9767CFF5FF6F7FB336365.taxon	biology_ecology	Biology. Larval parasitioid of Agrilus spp. (Coleoptera: Buprestidae) infesting dead branches of Villebrunea integrifolia Gaud. (Urticaceae) (Fig. 7 A) and Albizia amara (Roxb.) Boivin. (Fabaceae) (Fig. 7 D). Individuals of H. longipetiolata (Fig. 8 A) were first recorded emerging from branch cuttings of V. integrifolia infested with Agrilus sp. 1 (Fig. 7 C) collected from Madikeri, Karnataka. A total of three females and one male of H. longipetiolata and more than thirty individuals of Agrilus sp. 1 emerged from the branch cuttings. During the second encounter with the same species in July 2025, several individuals of H. longipetiolata (Fig. 6 A, B) were observed inspecting dead branches of A. amara in Doddaballapura, Karnataka. During subsequent visits to the same location, females of H. longipetiolata were seen ovipositing (Fig. 6 D) and several males were observed approaching females (Fig. 6 C) on dead branches of A. amara. The branches had a large number “ D ” shaped exit holes (Fig. 6 I) characteristic of Agrilus beetles and one individual of Agrilus sp. 2 (Figs 6 G, 7 F) was observed emerging from the dead branch. Upon collecting the infested branch, part of it was carefully split open and several pupae of H. longipetiolata along with host remains (Fig. 6 F) and adults of Agrilus sp. 2 were recovered. A total of fifteen individuals (ten females and five males) of H. longipetiolata and more than twenty adults of Agrilus sp. 2 emerged from the sampled branch cuttings of A. amara.	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD9767CFF5FF6F7FB336365.taxon	distribution	Distribution. India: Karnataka (Doddaballapura and Madikeri).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD9767CFF5FF6F7FB336365.taxon	etymology	Etymology. The species name (adjective) refers to the remarkable and distinctly long petiole of the male, which is comparatively longer than most other species of Heydenia.	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD9767CFF5FF6F7FB336365.taxon	discussion	Comments. Females of H. longipetiolata can be readily differentiated from other Indian species (H. indica, H. gibsoni, H. kashmirensis and H. tuberculata) and Chinese species (H. angularicoxa, H. coomoni Xiao & Huang, 2002, H. scolyti Yang, 1996 and H. testacea) in having the mesosoma moderately compressed dorsoventrally, propodeum with plical region distinctly reticulate-punctate or with reticulation incompletely to completely reduced with irregular transverse rugae, with median carina distinct in posterior half and shallowly merging with the cuticle surface in the anterior half, propodeal callus below the level of spiracles distinctly reticulate-punctate (Fig. 3 A) or irregularly rugose with reticulation reduced (Fig. 5 B, C), fore wing with basal setal line present, profemur distinctly enlarged with basal ventral angle, and ovipositor sheath 1.9 – 2 × the length of metatibia. Females are similar to those of H. trinodis Bouček, 1988, described from Papua New Guinea, in having a flat scutellum. They differ from H. trinodis in having the axillae reticulate, the plical region with the median carina distinct in its posterior half and shallowly merging with the surface in its anterior half, the petiole 1.4 – 1.5 × as long as wide, and the ovipositor sheaths 1.9 – 2.0 × the length of metatibia (in H. trinodis the axillae shiny and almost smooth, plical region with sharp median carina, petiole about 2 × as wide as long, and ovipositor sheaths about 1.2 × length of metatibia). Females differ from those of H. longicollis Cameron, 1912, described from Australia, H. pretiosa, described from Europe, and H. unica Cook & Davis, 1891, described from USA, in having a longer ovipositor sheath, 1.9 – 2.0 × the length of metatibia (in H. longicollis, H. pretiosa and H. unica the ovipositor sheath is rather short, 0.1 – 0.3 × the length of the metatibia). Individuals are separated from H. mateui Hedqvist, 1967, described from Chad, in having the plicae distinct and complete on the propodeum, the fore wing with the costal cell setose, and the metatibia with two unequal spurs (in H. mateui, propodeum with plicae reduced, indicated posteriorly, fore wing with costal cell bare, and metatibia with a single spur). Males of H. longipetiolata can be differentiated from those of H. natalensis, Westwood, 1874, described from South Africa, in having the antennal flagellomeres uniformly black, and the petiole 3 × as long as wide (in H. natalensis, antennae with fl 5 – fl 5 lighter in color than rest of the segments, and petiole very short). Males are similar to those of H. madagascariensis, Hedqvist, 1961, described from Madagascar, in possessing a long petiole, but can be differentiated by having the propodeal plical region with the median carina distinct in its posterior half to two thirds, the fore wing basal cell with a basal setal line, and the infuscation near the stigmal vein small (in H. madagascariensis, the propodeal plical region with a complete median carina, fore wing with basal cell bare, and infuscation near STV broad, reaching the posterior wing margin).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD0767CFF5FF7D6FBBF66BF.taxon	description	(Figs 8 B, 9 E, 10)	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD0767CFF5FF7D6FBBF66BF.taxon	materials_examined	Material examined. INDIA: Karnataka, Doddaballapura, 26. vi. 2024, coll. Hemanth (3 ♀ on separate cards); Karnataka, Chikkaballapur, Gudibande, 19. vii. 2024, coll. Hemanth (1 ♀ and 1 ♂ on separate cards).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD0767CFF5FF7D6FBBF66BF.taxon	diagnosis	Diagnosis. FEMALE. Body length 2.6 – 4.2 mm; face with iridescent lanceolate setae (Fig. 10 A); POL 2 × OOL; gena reticulate with distinct smooth and shiny area below outer margin of eye (Fig. 10 C), extending to vertex (Fig. 10 B); antenna with clava subequal to combined length of preceding three segments (Fig. 10 C); lateral panels of pronotum angled at about 90 ° to the dorsal surface (Fig. 10 D) and with a punctate region as in Fig. 10 E; axillae uniformly reticulate (Fig. 10 D); dorsellum smooth and shiny (Fig. 10 F); propodeal plical region with distinct median carina and plicae forming well defined hexagonal area (Fig. 10 F); profemur with ventral margin distally and protibia with dorsal margin having distinct spines (Fig. 10 G); fore wing with basal cell bare, infuscation near STV narrow, almost triangular, and with a small spot below (Fig. 10 I); metasoma with petiole wider than long, widening posteriorly (Fig. 10 H), Gt 1 and Gt 2 with posterior margins distinctly incised medially (Fig. 10 J); ovipositor sheath 1.6 × length of metatibia (Fig. 9 E). MALE. Closely similar to female but body length 2.2 – 3.6 mm and POL 1.4 × OOL.	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD0767CFF5FF7D6FBBF66BF.taxon	description	Variation. POL 1.45 – 2.00 × OOL (females).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD0767CFF5FF7D6FBBF66BF.taxon	biology_ecology	Biology. Complete biology unknown. In the present study the specimens were reared from dead branch cuttings of Pongamia pinnata (L.) Pierre. (Fabaceae) and an undetermined tree.	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD0767CFF5FF7D6FBBF66BF.taxon	distribution	Distribution. India (Karnataka) (new record), Sri Lanka (Sureshan 2009).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD07667FF5FF371FB0062F9.taxon	description	(Figs 8 C, 9 A, B, 11)	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD07667FF5FF371FB0062F9.taxon	materials_examined	Material examined. INDIA: Karnataka, Doddaballapura, 10. i. 2024, coll. Hemanth (1 ♀ on card); Karnataka, Chikkaballapur, Nandi Grama, 05. vi. 2025, coll. Rohit Pattar (1 ♀ and 1 ♂ on separate cards); Karnataka, Ramanagara, 05. vii. 2025, coll. Hemanth (1 ♂ on card); Karnataka, Tumkur, 22. vii. 2025, coll. Hemanth (1 ♀ on card).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD07667FF5FF371FB0062F9.taxon	diagnosis	Diagnosis. FEMALE. Body length 2.4 – 4.6 mm; face with iridescent lanceolate setae (Fig. 11 A); POL 1.4 × OOL; gena uniformly reticulate (Fig. 11 C); antenna with clava subequal to combined length of preceding three segments (Fig. 11 A, C); lateral panels of pronotum angled at about 90 ° to the dorsal surface (Fig. 11 D) and with a punctate area as in Fig. 11 E; axillae uniformly reticulate (Fig. 11 D); dorsellum smooth and shiny (Fig. 11 F); propodeal plical region with distinct median carina and plicae forming well-defined hexagonal area (Fig. 11 F); profemur with ventral margin distally and protibia with dorsal margin having distinct spines (Fig. 11 G); fore wing with basal cell bare, infuscation near STV almost rectangular (Fig. 11 I); metasoma with petiole bell shaped, about as long as wide (Fig. 11 H), Gt 1 and Gt 2 with posterior margins distinctly incised medially (Fig. 11 J); ovipositor sheath 1.6 × length of metatibia (Fig. 9 B). MALE. Similar to female but body length 2.1 – 2.9 mm; POL 1.45 × OOL; metasoma with petiole narrow, slightly longer than wide, Gt 1 with posterior margin incised medially and Gt 2 with posterior margin almost straight.	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD07667FF5FF371FB0062F9.taxon	biology_ecology	Biology. Complete biology unknown. Khanday et al. (2021) described H. kashmirensis based on specimens collected from Ulmus villosa (Ulmaceae) infested with Sinoxylon sp. (Coleoptera: Curculionidae) collected from Jammu & Kashmir. In the present study it was reared from dead branch cuttings of Cassia seimea Lam. (Fabaceae) and Gliricidia sepium (Jacq.) Walp. (Fabaceae).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFD07667FF5FF371FB0062F9.taxon	distribution	Distribution. India: Jammu & Kashmir (Khanday et al. 2021), Karnataka (present study).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFCB7667FF5FF72AFC2A6427.taxon	description	(Figs 8 D, 9 C, D, 12)	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFCB7667FF5FF72AFC2A6427.taxon	materials_examined	Material examined. INDIA: Karnataka, Gudibande, Chikkaballapur, 19. vii. 2024, coll. Hemanth (1 ♀ on card); Karnataka, Kolar, Kailasagiri, 12. ix. 2023, coll. Hemanth (1 ♂ on card); Karnataka, Ramanagara, 05. vii. 2025, coll. Hemanth (2 ♀ and 1 ♂ male on separate cards); Himachal Pradesh, Hamirpur, 20. xi. 2024, coll. Adarsh (1 ♀ female on card).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFCB7667FF5FF72AFC2A6427.taxon	diagnosis	Diagnosis. FEMALE. Body length 1.9 – 3.4 mm; face with a transverse patch of thick white setae (Fig. 12 A) (without iridescent lanceolate setae); POL 2.5 × OOL; gena reticulate (Fig. 12 C); antenna with clava subequal to combined length of preceding four segments (Fig. 12 A, C); lateral panels of pronotum smooth (Figs 12 E, 9 D) and angled at about 90 ° to the dorsal surface (Fig. 12 D); axillae smooth and shiny with weak, shallow reticulation (Fig. 12 D); dorsellum smooth and shiny (Fig. 12 F); propodeal plical region with median carina projecting upwards in the middle (Figs 9 D, 12 E) and with two oblique carinae together forming a ‘ Y’ shape (Fig. 12 F); profemur with ventral margin distally and protibia with dorsal margin lacking distinct spines (Fig. 12 G); fore wing with basal cell bare, infuscation near STV broadly rectangular with a circular spot below (Fig. 12 I); metasoma with petiole distinctly wider than long (Fig. 12 H), 1.6 – 2.0 × as wide as long, Gt 1 with posterior margin straight and Gt 2 with posterior margin medially produced (Fig. 12 J); ovipositor sheath 0.5 × length of metatibia (Fig. 9 D). MALE. Similar to female but body length 1.6 – 2.0 mm; infuscation near STV broad but not rectangular (ending nearly before the anterior half); metasoma with petiole only slightly wider than long, 1.2 – 1.3 × as wide as long and Gt 1 distinctly narrowly elongated (tubular).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFCB7667FF5FF72AFC2A6427.taxon	description	Variation. POL 3.0 – 3.4 × OOL (females), POL 3.0 – 3.1 × OOL (males).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFCB7667FF5FF72AFC2A6427.taxon	biology_ecology	Biology. Yang (1996) reported the species to be parasitic on larvae of Sphaerotrypes coimbatorensis Stebbing, 1906 (Coleoptera: Curculionidae) on Juglans regia L. (Juglandaceae), and on Scolytus seulensis Murayama, 1930 and S. japonicus Chapuis, 1875 (Coleoptera: Curculionidae) on Prunus persica L. Batsch. (Rosaceae). In the present study, specimens (from Karnataka) were reared from branch cuttings of Pongamia pinnata (L.) Pierre. (Fabaceae).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFCB7667FF5FF72AFC2A6427.taxon	distribution	Distribution. China (Jiangsu, Shaanxi) (Yang 1996), India (new records: Himachal Pradesh and Karnataka).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
5368996BFFCB7667FF5FF72AFC2A6427.taxon	discussion	Comments. The specimens newly collected from India match the description of H. testacea by Yang (1996) in having the head and mesosoma testaceous, the toruli located distinctly below the lower orbit of the eyes, and with a transverse patch of thick white setae below, the fore wing with a small spot close to the broad infuscation below the stigmal vein (fig. 157 in Yang 1996), the propodeal plical region with the median carina projecting upwards in the middle and with two oblique carinae together forming a ‘ Y’ shape (fig. 20 in Yang 1996), and the ovipositor sheath short, in dorsal view 0.1 × the length of the metasoma. Our specimens differ from that described for H. testacea only in having different POL: OOL measurements (POL 3.0 – 3.4 × OOL).	en	Kumar, Hemanth, Gupta, Ankita, Sharma, Adarsh (2025): Description of a new species of Heydenia Förster, 1856 (Hymenoptera: Heydeniidae) from Southern India with two new distributional records. Zootaxa 5711 (1): 93-113, DOI: 10.11646/zootaxa.5711.1.4, URL: https://doi.org/10.11646/zootaxa.5711.1.4
