identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5B7587A9A10AFFE5FEA0FE63FD3F0676.text	5B7587A9A10AFFE5FEA0FE63FD3F0676.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthodes Kiesenwetter 1852	<div><p>Malthodes Kiesenwetter, 1852</p><p>Malthodes Kiesenwetter, 1852: 242 .</p><p>Type species: Malthinus marginatus Latreille, 1806 .</p></div>	https://treatment.plazi.org/id/5B7587A9A10AFFE5FEA0FE63FD3F0676	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kazantsev, S. V.	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10AFFE4FF1EFDF1FEF4037A.text	5B7587A9A10AFFE4FF1EFDF1FEF4037A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthodes nabozhenkorum Kazantsev	<div><p>Malthodes nabozhenkorum Kazantsev, sp.n.</p><p>Figs 1, 3, 4, 10, 11.</p><p>MATERIAL. Holotype, ♂, South Ossetia, Dzau District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.181946&amp;materialsCitation.latitude=42.539444" title="Search Plazi for locations around (long 44.181946/lat 42.539444)">between Edisa and Zgubir</a>, 42°32′22″N, 44°10′55″E, 1760 m, 17.06.2023, leg. M.V. &amp; S.V. Nabozhenko (ICM) ; paratypes, 1♂ and 2♀♀, Georgia, Bakuriani, [1800 m], 16–18.VII.1928, D. Romashov (ZMMU) ; 1♀, Georgia, env. Bakuriani, gorge nr Mitarbi (to Kochta Mt.), 29.VII.1928, D. Romashov (ICM) .</p><p>DESCRIPTION. Male. Dark brown to black; palps, except ultimate palpomeres, antennomere 1, knees, pronotal posterior angles and anterior margin narrowly testaceous; elytral middle two thirds pale brown; elytral apices yellow (Fig. 1).</p><p>Head transverse, without eyes about as wide as pronotum. Eyes relatively small, spherical, interocular distance ca 2.2 times greater than eye diameter. Vertex in scarce punctuation. Ultimate maxillary and labial palpomeres ca 2 times longer than wide. Antennae filiform, almost attaining to apices of folded wings; antennomere 3 ca 1.5 times longer than pedicel (antennomere 2) and ca 1.4 times shorter than antennomere 4; antennal pubescence short and semi-erect (Fig. 1).</p><p>Pronotum transverse, ca 1.4 times wider than long, slightly widening anteriorly, with somewhat concave sides, rounded posterior and blunt laterally developed anterior angles, distinctly concave anteriorly and convex posteriorly (Fig. 1).</p><p>Elytra elongate, ca. 2.4 times longer than wide at humeri, parallel-sided, leaving posterior third of folded wings uncovered; elytral pubescence uniform, short and sub-erect. Scutellum transverse, narrowing distally, broadly rounded at apex (Fig. 1).</p><p>Legs long and slender; femora and tibiae narrow and straight, tibiae noticeably shorter than femora; hind tarsomere length ratio 5.5: 3.4: 1.7: 1: 2.7; tarsomere 4 deeply cleft; all claws simple (Fig. 1).</p><p>Ultimate sternite elongate, gradually narrowing distally and slightly widened at apex, strongly curved in lateral view; ultimate tergite elongate, slightly narrowed distally, almost truncate at apex and slightly curved before apex in lateral view; penultimate tergite elongate, distally widened, with convex sides; third from end tergite with acute lateral process (Figs 3, 4).</p><p>Aedeagus oval, with prominent parameres, noticeably widened and densely pubescent at apex; laterophyses with acute dents at base, pointed and bent at apex in lateral view (Figs 10, 11).</p><p>Length (from head to apices of folded elytra): 3.6–3.7 mm; width (at humeri): 0.8–0.85 mm.</p><p>FEMALE. Unknown.</p><p>ETYMOLOGY. The new species is named after Drs M. V. &amp; S. V. Nabozhenko ( Makhachkala, Dagestan) who collected the type specimen.</p><p>DIAGNOSIS. Malthodes nabozhenkorum sp.n. is similar to M. kasantsevi Wittmer, 1992, known from the alpine zone of mountains of Krasnodar Krai and Abkhasia, differing in the widened distally ultimate sternite (Fig. 3) and presence of teeth at the base of aedeagal laterophyses (Figs 10, 11), vs narrow parallel-sided ultimate sternite in its distal part and absence of teeth at the base of laterophyses in M. kasantsevi (Figs 5, 12, 77, 110).</p></div>	https://treatment.plazi.org/id/5B7587A9A10AFFE4FF1EFDF1FEF4037A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kazantsev, S. V.	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10BFFE4FEFDF8E5FABD028B.text	5B7587A9A10BFFE4FEFDF8E5FABD028B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthodes sukkoensis Kazantsev	<div><p>Malthodes sukkoensis Kazantsev, sp.n.</p><p>Figs 2, 6, 7, 13, 14.</p><p>MATERIAL. Holotype, ♂, S Russia, Krasnodar Krai, Anapa Distr., env. Sukko, 28-30.IV.2018, leg. E.A. Khachikov leg. (ICM); paratype, 1♂, same label (ICM).</p><p>DESCRIPTION. Male. Dark brown to black; antennomere 1 and antennomere 2 proximally, knees and anterior tibiae apically testaceous; elytral middle third pale brown; elytral apices yellow (Fig. 2).</p><p>Head subquadrate, about as wide as pronotum. Eyes moderately large, spherical, interocular distance about as long as eye diameter. Vertex in scarce punctuation. Ultimate maxillary and labial palpomeres ca 3.7 times longer than wide. Antennae filiform, almost attaining to apices of folded wings; antennomeres 2 and 3 subequal in length and ca 1.3 times shorter than antennomere 4; antennal pubescence relatively short and semi-erect (Fig. 2).</p><p>Pronotum transverse, ca 1.2 times wider than long, widening anteriorly, with small acute posterior and truncate laterally developed anterior angles, convex anteriorly and posteriorly (Fig. 2).</p><p>Elytra elongate, ca. 2.3 times longer than wide at humeri, parallel-sided, leaving posterior third of folded wings uncovered; elytral pubescence uniform, short and sub-erect. Scutellum transverse, slightly narrowing distally, broadly rounded at apex (Fig. 2).</p><p>Legs long and slender; femora and tibiae narrow and straight, tibiae noticeably shorter than femora; hind tarsomere length ratio 4.5: 2.5: 1.8: 1: 1.5; tarsomere 4 deeply cleft; all claws simple (Fig. 2).</p><p>Ultimate sternite elongate, gradually narrowing distally, widened and deeply emarginate at apex, somewhat curved in lateral view; ultimate tergite elongate, slightly narrowed distally, with roundish incision at apex; penultimate tergite transverse, sub-rectangular; third from end tergite simple (Figs 6, 7).</p><p>Aedeagus roundish, with rounded distally dorsal plate; laterophyses in distal half relatively broad, parallel-sided, with almost rectangular bulges at base, in lateral view slanted outwardly, but not hooked at apex; penis strongly curved, in lateral view (Figs 13, 14).</p><p>Length (from head to apices of folded elytra): 3.0 mm; width (at humeri): 0.5–0.6 mm.</p><p>FEMALE. Unknown.</p><p>ETYMOLOGY. The new species is named after the type locality.</p><p>DIAGNOSIS. Malthodes sukkoensis sp.n. resembles M. pseudobesucheti Wittmer, 1970, from the mountains of Krasnodar Krai and Abkhasia, differing in the relatively narrowly incised at apex ultimate sternite, shorter and not widened at apex ultimate tergite, more rounded dorsal plate of the aedeagus and not hooked apically, in lateral view, laterophyses (Figs 6, 7, 13, 14), vs broadly widened at apex ultimate sternite, longer and widened at apex ultimate tergite, more narrowed distally dorsal plate of the aedeagus and apically hooked laterophyses, in lateral view, in M. pseudobesucheti (Figs 8, 9, 15, 48, 49, 87, 88).</p><p>REMARKS. The paratype lacks head and pronotum.</p></div>	https://treatment.plazi.org/id/5B7587A9A10BFFE4FEFDF8E5FABD028B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kazantsev, S. V.	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10BFFEDFCBBF954FA650583.text	5B7587A9A10BFFEDFCBBF954FA650583.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthodes Kiesenwetter 1852	<div><p>A KEY TO MALTHODES SPECIES OF THE GREATER CAUCASUS</p><p>1(20) Small, usually smaller than 3 mm. Antennomere 2 subequal in length to antennomere 3 or slightly longer (only in ............... M. bourgeoisi slightly shorter) (Figs 16–24).</p><p>2(3) Antennomere 2 slightly shorter than antennomere 3. Dark brown, pronotum light brown, darkened at sides. Elytra long, almost completely covering folded wings (Fig. 16). Ultimate sternite abruptly narrowed in distal half; ultimate tergite narrow, elongate, slightly narrowed and rounded at apex (Figs 38–40). Aedeagus — Fig. 81. Body length 2.8–3.2 mm. ....................... M. bourgeoisi (Reitter, 1888)</p><p>3(2) Antennomere 2 subequal in length to antennomere 3 or slightly longer. Elytra shorter, leaving apices of folded wings uncovered (Figs 17–24).</p><p>4(5) Upperside, including antennae, uniformly dark brown to black (Fig. 17). Ultimate sternite parallel-sided, slightly emarginate at apex; ultimate tergite short, non-emarginate apically; (Fig. 41). Aedeagus — Fig. 82. Body length 2.3–2.8 mm ..................................... ........................................ M. crassicornis (Mäklin, 1846)</p><p>5(4) Upperside, including antennae, partly yellow or light brown (Figs 18–24).</p><p>6(15) Elytra with bright yellow apical spots (Figs 18–22).</p><p>7(10) Pronotum uniformly black or dark brown, or narrowly touched with light brown at anterior and posterior margins (Figs 18, 19).</p><p>8(9) Ultimate sternite widened in distal half, with two processes; ultimate tergite parallel-sided, with broadly separated processes at apex (Figs 42, 43). Aedeagus — Fig. 83. Body length 2.7–3.1 mm (Fig. 18). ......... M. amplithorax Wittmer, 1992</p><p>9(8) Ultimate sternite narrow, deeply split at apex; ultimate tergite semi-oval, with shallow incision at apex (Figs 44, 45). Aedeagus — Fig. 84. Body length 2.3–2.8 mm (females can be slightly over 3 mm) (Fig. 19). ............................................. M. lederi Pic, 1912</p><p>10(6) Pronotum more or less broadly yellow at anterior and posterior margins (Figs 20–22).</p><p>11(12). Ultimate sternite incised at apex; ultimate tergite narrow, narrowed towards apex (Figs 46, 47). Aedeagus — Figs 85, 86. Body length 2.9–3.1 mm (Fig. 20) .................................... M. strejceki Švihla, 1990</p><p>12(11). Ultimate sternite not incised at apex (Figs 6, 7, 48, 49).</p><p>13(14) Ultimate sternite broadly widened at apex, ultimate tergite relatively long and widened at apex (Figs 8, 9, 48, 49). Aedeagal dorsal plate narrowed distally; laterophyses hooked apically, in lateral view (Figs 15, 87, 88). Body length 2.9–3.1 mm (Fig. 21). ................... M. pseudobesucheti Wittmer, 1970</p><p>14(13) Ultimate sternite relatively narrowly widened at apex, ultimate tergite relatively short and not widened at apex (Figs 6, 7). Aedeagal dorsal plate distally rounded; laterophyses not hooked apically, in lateral view (Figs 13, 14). Body length 3 mm (Fig. 2). ............................................. M. sukkoensis sp.n.</p><p>15(6) Elytra without yellow apical spots (Figs 22–24).</p><p>16(19) Pronotum uniformly dark brown to black (Fig. 22, 23).</p><p>17(16) Ultimate sternite long, narrow, strongly curved, split at apex; ultimate tergite almost perpendicular to the preceding one, deeply cleft at apex (Figs 50, 51). Aedeagus — Fig. 89. Body length 2.4–2.8 mm (Fig. 22). ......................... M. sotschienesis Wittmer, 1970</p><p>18(17) Ultimate sternite moderately long, slightly bent and shallowly emarginate at apex; ultimate tergite coaxial with the preceding one, with shallow split at apex (Figs 52, 53). Aedeagus — Figs 90, 91. Body length 2.7–2.8 mm (Fig. 23). ............. M. seregiusi Kazantsev, 2021</p><p>19(16) Pronotum partly yellow or light brown at anterior and posterior margins (Fig. 24). Ultimate sternite with relatively short and broad apical processes; ultimate tergite bent downwards before apex, widened and semi-circularly incised at apex (Figs 54, 55).Aedeagus — Fig. 92. Body length 3.0– 3.5 mm. ........ .......................................................... M. mutatus Wittmer, 1970</p><p>20(1) Relatively large, usually larger than 3.5 mm. Antennomere 2 noticeably shorter than antennomere 3 (Fig. 25–37).</p><p>21(24) Pronotum completely margined at sides (Figs 26, 27).</p><p>22(23) Third from end tergite with a dent; ultimate tergite elongate; ultimate sternite without elongate lateral processes (Figs 56–58). Aedeagus — Fig. 93. Body length 3.8–4.5 mm (Fig. 25). ........................ M. circassicus Švihla, 1980</p><p>23(22) Third from end tergite simple; ultimate tergite transverse; ultimate sternite with narrow long lateral processes (Figs 59, 60). Aedeagus — Fig. 94. Body length 4.5 mm (Fig. 26). ............................... M. kobiensis Wittmer, 1970</p><p>24(21) Pronotum margined at sides only at anterior angles (Figs 28–37).</p><p>25(34) Third from end tergite simple (Figs 61–65).</p><p>26(29) Pronotum uniformly dark brown to black, sometimes with narrow testaceous bordering at anterior and posterior margins (Fig. 27).</p><p>27(28) Pronotum uniformly black, distinctly elongate; elytra uniformly black (Fig. 27). Ultimate sternite widened and slightly cleft at apex (Figs 61, 62). Aedeagus — Fig. 95. Body length 3.5–5.0 mm.................................... M. abkhasicus Wittmer, 1979</p><p>28(27) Pronotum with narrow testaceous bordering at anterior and posterior margins, distinctly transverse. Ultimate sternite oval. Body length 4.8–5.0 mm. ................. M. jaromiri Švihla, 2002</p><p>29(26) Pronotum mostly testaceous or broadly lightened at angles (Figs 28–30).</p><p>30(31) Ventral plate of aedeagus very narrow, only slightly widened towards apex; laterophyses massive, widened and dentate at apex; interophyses short and straight (Fig. 96). Bases of tibiae testaceous; elytra uniformly dark brown; antennae light brown, relatively short, slightly reaching over elytral apices (Fig. 28). Body length 5.0– 5.5 mm. .. ............................................ M. medvedevi Wittmer, 1992</p><p>31(30) Ventral plate of aedeagus relatively broad; laterophyses narrow and long; interophyses absent (Figs 97–100).</p><p>32(33) Laterophyses in lateral view widened and bent at apex; at base, in dorsal view, flat (Figs 97, 98). Pronotum varies from uniformly testaceous to uniformly dark brown (Fig. 29). Body length 3.8–4.8 mm. ................... M. castanicollis Reitter, 1888</p><p>33(32) Laterophyses in lateral view narrow and straight; at base, in dorsal view, dentate (Figs 99, 100). Body length 3.3–5.3 mm (Fig. 30)................... M. bohaci Švihla, 2002</p><p>34(25) Third from end tergite with processes or dents (Figs 66–80).</p><p>35(48) Third from end tergite with a dent (Figs 3–5, 66–77).</p><p>36(45) Ultimate sternite and tergite with a pair of apical processes (Figs 66–76).</p><p>37(38) Ultimate sternite with long cithern-like processes (Figs 66, 67). Pronotum from uniformly black to noticeably lightened at angles (Fig. 31). Aedeagus — Figs 101, 102. Body length 4.0– 4.5 mm. .... M. lyriformis Wittmer, 1992</p><p>38(37) Ultimate sternite with relatively short and straight processes (Figs 68–72).</p><p>39(42) Aedeagus with relatively short parameres, not surpassing ventral plate in length (Figs 103–105).</p><p>40(41) Ultimate sternite weakly narrowed in the middle and not bent before apex (Figs 68–70). Aedeagus with almost truncate apex of ventral plate and non-widened before apex laterophyses (Figs 103). Elytra with bright yellow apices (Fig. 32). Body length 5.0– 5.5 mm. ......................... M. nyholmi Wittmer, 1970</p><p>41(40) Ultimate sternite distinctly narrowed in the middle and abruptly bent before apex (Figs 71, 72). Aedeagus with distally rounded ventral plate and noticeably widened before apex laterophyses (Figs 104, 105). Elytra uniformly black (Fig. 33). Body length 4.7–4.8 mm. ........................... ................................................... M. lozovoyi Kazantsev, 2021</p><p>42(39) Aedeagus with relatively long parameres, surpassing ventral plate in length (Figs 106–109).</p><p>43(44) Apical processes of ultimate sternite relatively long (Figs 73, 74). Aedeagus elongate; penis before apex convex, in lateral view (Figs 106, 107). Upperside dark brown to black; pronotal angles testaceous (Fig. 34). Body length 3.0– 3.8 mm. ............. ..................................................... M. caucasicus Wittmer, 1958</p><p>44(43) Apical processes of ultimate sternite relatively short (Figs 75, 76). Aedeagus subquadrate; penis before apex concave, in lateral view (Figs 108, 109). Habitually similar to M. caucasicus (Fig. 35). Body length 4.0 mm. ........... .............................................. M. vikhrevi Kazantsev 2021</p><p>45(36) Ultimate sternite and tergite simple (Figs 3–5, 77).</p><p>46(47) Ultimate sternite noticeably widened distally (Figs 3, 4). Pronotal angles testaceous; elytra with bright yellow apical spots (Fig. 1). Aedeagal laterophyses with dents (Figs 10, 11). Body length 3.6–3.7 mm. ......................... .................................................. M. nabozhenkorum sp.n.</p><p>47(46) Ultimate sternite parallel-sided distally (Figs 5, 77). Pronotal angles, bases of antennomeres and bases of tibiae testaceous; elytra with bright yellow apical spots (Fig. 36). Aedeagal laterophyses without dents (Figs 12, 110). Body length 3.0– 3.5 mm. ............. M. kasantsevi Wittmer, 1992</p><p>48(35) Third from end tergite with a pair of relatively broad processes at posterior edge, also bearing downward directed additional appendage at each side; ultimate sternite not emarginate at apex (Figs 78–80). Pronotal angles, bases of antennomeres 1–2 and bases of tibiae testaceous; elytra uniformly dark brown to black (Fig. 37).Aedeagus — Fig. 111. Body length 3.2–4.2 mm. ... M. orientalicus Wittmer, 1970</p></div>	https://treatment.plazi.org/id/5B7587A9A10BFFEDFCBBF954FA650583	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kazantsev, S. V.	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A102FFE8FCBBFE63FD7906BD.text	5B7587A9A102FFE8FCBBFE63FD7906BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malthodes Kiesenwetter 1852	<div><p>Annotated checklist of Malthodes species of the Greater Caucasus</p><p>Malthodes Kiesenwetter, 1852</p><p>Malthodes Kiesenwetter, 1852: 242 .</p><p>Type species: Malthinus marginatus Latreille, 1806 .</p><p>1. abkhasicus Wittmer, 1979: 141 . North-western Caucasus: Krasnodar Krai (Sochi area, Krasnaya Polyana, Lagonaki, Ubinskaya) and Abkhasia (Miussera) (Map 2). 100–2000 m, June through July.</p><p>2. amplithorax Wittmer, 1992: 26 . North-western Caucasus: Krasnodar Krai (Krasnaya Polyana, Lagonaki) (Map 1). 800–2000 m, June through July.</p><p>3. bohaci Švihla, 2002: 137 . North-western Caucasus: Krasnodar Krai (Sochi area) and Abkhasia (Bzyb valley, Hodzhal Mt) (Map 2). 500–1000 m, June.</p><p>4. bourgeoisi (Reitter, 1888): 210 ( Malchinus). Northern Caucasus: Krasnodar Krai (Sochi area), North Ossetia (Ardon valley, Tsei Gorge, Tersky Mts), Dagestan (Gunib) (Map 1). The distribution area also includes Georgia, Iran and Turkey [Wittmer, 1992]. 1800–2000 m, June through August.</p><p>= rhaphidostylus Pic, 1912: 59.</p><p>5. castanicollis Reitter, 1888: 208 . North-western Caucasus: Krasnodar Krai (Sochi area, Tkhab, Ubinskaya) and Abkhasia (env. Sukhum) (Map 2). 2000 m, June.</p><p>6. caucasicus Wittmer, 1958: 120 . Caucasus: Georgia (Kobi) and Abkhasia (Ritsa R.N. P.) (Map 2). 1500-2120 m, July.</p><p>7. circassicus Švihla, 1980: 249 . North-western Caucasus: Krasnodar Krai (Krasnaya Polyana, Utrish) (Map 2). 100– 2000 m, June.</p><p>8. crassicornis (Mäklin, 1846): 179 ( Malthinus). From West to East Europe, also in North-western Caucasus: Krasnodar Krai (Krasnaya Polyana) (Map 1). The distribution area also includes Iran and Turkey. May through July.</p><p>9. jaromiri Švihla, 2002: 138 . North-western Caucasus: Abkhasia (Kelasuri valley) (Map 2). 500 m, June.</p><p>10. kasantsevi Wittmer, 1992: 34 . North-western Caucasus: Krasnodar Krai (Krasnaya Polyana, Lagonaki) and Abkhasia (Ritsa R.N. P., Bzyb Mts) (Map 2). 1700–2500 m, subalpine and alpine zone, July.</p><p>11. kobiensis Wittmer, 1970: 62 . North-western and northern Caucasus: Krasnodar Krai (Sochi area, Krasnaya Polyana, Lagonaki, Ubinskaya), North Ossetia (Skalisty Mts), Chechnya, Abkhasia (Gagrsky Range, Bzyb valley, Hodzhal Mt, Tsumuri), also Georgia (Kobi) (Map 2). 840– 3000 m, June through August.</p><p>12. lederi Pic, 1912: 59 . Northern Caucasus: North Ossetia (Vladikavkaz, Skalisty Mts), Ingushetia, Dagestan (Gunib, Nukatl Mts); also South Ossetia (Dzau) and Georgia (Kobi) (Map 1). 1050–2200 m, June through July. First record for South Ossetia.</p><p>13. lozovoyi Kazantsev, 2021: 231 . North-western Caucasus: Abkhasia (Hodzhal Mt); also South Ossetia (Dzau) (Map 2). 1300–1800 m, June through July. First record for South Ossetia.</p><p>14. lyriformis Wittmer, 1992: 29 . North-western Caucasus: Krasnodar Krai (Krasnaya Polyana, Lagonaki) and Abkhasia (Gagrsky Mts, Ritsa R.N. P.) (Map 2). 1700–2000 m, July.</p><p>15. medvedevi Wittmer, 1992: 22 . North-western Caucasus: Krasnodar Krai (Krasnaya Polyana, Lagonaki) and Abkhasia (Hodzhal Mt) (Map 2). The distribution area also includes Georgia (Tsagveri). 800–2000 m, July.</p><p>16. mutatus Wittmer, 1970: 40 . Northern Caucasus: Teberda, North Ossetia (Tbaukhokh Mt, Skalisty Mts), Dagestan (Gunib) (Map 2). 1270–2350 m, June through July.</p><p>17. nabozhenkorum Kazantsev, sp.n. Greater Caucasus: South Ossetia. Also in central Georgia (Bakuriani). 1760–1800 m, June through July.</p><p>18. nyholmi Wittmer, 1970: 52 . North-western Caucasus: Krasnodar Krai (Ubinskaya) and Abkhasia (Sukhum, Shoudidi Mt) (Map 2). Also distributed in Georgia and Turkey. 420–2000 m, May through July.</p><p>19. orientalicus Wittmer, 1970: 85 . North-western Caucasus: Krasnodar Krai (Ubinskaya), also South Ossetia (Map 2). The distribution area also includes Azerbaijan and Turkey. 1760 m, May. First record for South Ossetia.</p><p>20. pseudobesucheti Wittmer, 1970: 61 . North-western Caucasus: Krasnodar Krai (Krasnaya Polyana) and Abkhasia (Ritsa R.N. P., Bzyb Mts) (Map 1). 1670–2200 m, June through July.</p><p>21. seregiusi Kazantsev, 2021: 288 . North-western Caucasus: Krasnodar Krai (Sochi area) (Map 1). 220–550 m, May.</p><p>22. sotschienesis Wittmer, 1970: 63 . North-western Caucasus: Krasnodar Krai (Sochi area, Krasnaya Polyana) and Abkhasia (Ritsa R.N. P., Bzyb valley) (Map 1). 550-1900 m, June through July.</p><p>23. strejceki Švihla, 1990: 200 . North-western Caucasus: Krasnodar Krai (Krasnaya Polyana, Lagonaki, Ubinskaya) and Abkhasia (Ritsa R.N. P., Bzyb Mts) (Map 1). 1500– 1900 m, June through July.</p><p>24. sukkoensis Kazantsev, sp.n. North-western Caucasus: Krasnodar Krai (S Anapa). 100 m, April.</p><p>25. vikhrevi Kazantsev, 2021: 288 . North-western Caucasus: Krasnodar Krai (Khosta, Sochi area) (Map 2). May.</p><p>Malthodes lobicollis (Reitter, 1888) described from ‘Circassien’ based on several females [Reitter, 1888], whereas all the taxonomy of the genus is based on the morphological characters of males (shape and structures of the terminal abdominal segments and aedeagi), cannot be placed amidst its congeners properly. Therefore it is included in the list only as incertae sedis.</p><p>The following three species are excluded from the list of the Greater Caucasian Malthodes:</p><p>Malthodes debilis Kiesenwetter, 1852 . Distributed from West to East Europe [Delkeskamp, 1977; Wittmer, 1992; Kazantsev, Brancucci, 2007]. One specimen is reported from the Caucasus: ’Parabotch Forest, Ki[z]l., Tersk distr.’ [Wittmer, 1992]. The taxon is excluded from the list, because the locality where the only ‘Caucasian’ specimen was registered (Kizlyar distr., northern Dagestan), does not belong to the Greater Caucasus.</p><p>Malthodes quadristilus Wittmer, 1992 . The taxon is excluded from the list, as its type locality (Terekli-Mekteb, northern Dagestan — Wittmer, 1992) does not belong to the Greater Caucasus. The second known specimen of M. quadristilus (from Krasnodar Krai — Kazantsev, 2011; 2022), after being re-studied, has proven to belong to the closely related M. orientalicus Wittmer, 1970 .</p><p>Malthodes tordi Wittmer, 1970 . Malthodes tordi, described from Turkey (Trabzon) [Wittmer, 1970], was later indicated for Abkhazia and Adjara [Wittmer, 1992]. However, the Abkhazian ‘ M. tordi ’ proved to be a related, but different species — M. jaromiri Švihla, 2002 [Švihla, 2002], while the re-examination of the Adjara specimens, from the ‘ M. tordi ’ series identified by Wittmer, showed that they also belong to another species, M. vladimiri Kazantsev, 2021 [Kazantsev, 2021b]. Therefore, M. tordi is excluded from the regional list as well.</p></div>	https://treatment.plazi.org/id/5B7587A9A102FFE8FCBBFE63FD7906BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kazantsev, S. V.	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
