taxonID	type	description	language	source
5B7587A9A10AFFE5FEA0FE63FD3F0676.taxon	type_taxon	Type species: Malthinus marginatus Latreille, 1806.	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10AFFE4FF1EFDF1FEF4037A.taxon	description	Figs 1, 3, 4, 10, 11.	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10AFFE4FF1EFDF1FEF4037A.taxon	materials_examined	MATERIAL. Holotype, ♂, South Ossetia, Dzau District, between Edisa and Zgubir, 42 ° 32 ′ 22 ″ N, 44 ° 10 ′ 55 ″ E, 1760 m, 17.06.2023, leg. M. V. & S. V. Nabozhenko (ICM); paratypes, 1 ♂ and 2 ♀♀, Georgia, Bakuriani, [1800 m], 16 – 18. VII. 1928, D. Romashov (ZMMU); 1 ♀, Georgia, env. Bakuriani, gorge nr Mitarbi (to Kochta Mt.), 29. VII. 1928, D. Romashov (ICM).	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10AFFE4FF1EFDF1FEF4037A.taxon	description	DESCRIPTION. Male. Dark brown to black; palps, except ultimate palpomeres, antennomere 1, knees, pronotal posterior angles and anterior margin narrowly testaceous; elytral middle two thirds pale brown; elytral apices yellow (Fig. 1). Head transverse, without eyes about as wide as pronotum. Eyes relatively small, spherical, interocular distance ca 2.2 times greater than eye diameter. Vertex in scarce punctuation. Ultimate maxillary and labial palpomeres ca 2 times longer than wide. Antennae filiform, almost attaining to apices of folded wings; antennomere 3 ca 1.5 times longer than pedicel (antennomere 2) and ca 1.4 times shorter than antennomere 4; antennal pubescence short and semi-erect (Fig. 1). Pronotum transverse, ca 1.4 times wider than long, slightly widening anteriorly, with somewhat concave sides, rounded posterior and blunt laterally developed anterior angles, distinctly concave anteriorly and convex posteriorly (Fig. 1). Elytra elongate, ca. 2.4 times longer than wide at humeri, parallel-sided, leaving posterior third of folded wings uncovered; elytral pubescence uniform, short and sub-erect. Scutellum transverse, narrowing distally, broadly rounded at apex (Fig. 1). Legs long and slender; femora and tibiae narrow and straight, tibiae noticeably shorter than femora; hind tarsomere length ratio 5.5: 3.4: 1.7: 1: 2.7; tarsomere 4 deeply cleft; all claws simple (Fig. 1). Ultimate sternite elongate, gradually narrowing distally and slightly widened at apex, strongly curved in lateral view; ultimate tergite elongate, slightly narrowed distally, almost truncate at apex and slightly curved before apex in lateral view; penultimate tergite elongate, distally widened, with convex sides; third from end tergite with acute lateral process (Figs 3, 4). Aedeagus oval, with prominent parameres, noticeably widened and densely pubescent at apex; laterophyses with acute dents at base, pointed and bent at apex in lateral view (Figs 10, 11). Length (from head to apices of folded elytra): 3.6 – 3.7 mm; width (at humeri): 0.8 – 0.85 mm. FEMALE. Unknown.	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10AFFE4FF1EFDF1FEF4037A.taxon	etymology	ETYMOLOGY. The new species is named after Drs M. V. & S. V. Nabozhenko (Makhachkala, Dagestan) who collected the type specimen.	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10AFFE4FF1EFDF1FEF4037A.taxon	diagnosis	DIAGNOSIS. Malthodes nabozhenkorum sp. n. is similar to M. kasantsevi Wittmer, 1992, known from the alpine zone of mountains of Krasnodar Krai and Abkhasia, differing in the widened distally ultimate sternite (Fig. 3) and presence of teeth at the base of aedeagal laterophyses (Figs 10, 11), vs narrow parallel-sided ultimate sternite in its distal part and absence of teeth at the base of laterophyses in M. kasantsevi (Figs 5, 12, 77, 110).	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10BFFE4FEFDF8E5FABD028B.taxon	description	Figs 2, 6, 7, 13, 14.	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10BFFE4FEFDF8E5FABD028B.taxon	materials_examined	MATERIAL. Holotype, ♂, S Russia, Krasnodar Krai, Anapa Distr., env. Sukko, 28 - 30. IV. 2018, leg. E. A. Khachikov leg. (ICM); paratype, 1 ♂, same label (ICM).	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10BFFE4FEFDF8E5FABD028B.taxon	description	DESCRIPTION. Male. Dark brown to black; antennomere 1 and antennomere 2 proximally, knees and anterior tibiae apically testaceous; elytral middle third pale brown; elytral apices yellow (Fig. 2). Head subquadrate, about as wide as pronotum. Eyes moderately large, spherical, interocular distance about as long as eye diameter. Vertex in scarce punctuation. Ultimate maxillary and labial palpomeres ca 3.7 times longer than wide. Antennae filiform, almost attaining to apices of folded wings; antennomeres 2 and 3 subequal in length and ca 1.3 times shorter than antennomere 4; antennal pubescence relatively short and semi-erect (Fig. 2). Pronotum transverse, ca 1.2 times wider than long, widening anteriorly, with small acute posterior and truncate laterally developed anterior angles, convex anteriorly and posteriorly (Fig. 2). Elytra elongate, ca. 2.3 times longer than wide at humeri, parallel-sided, leaving posterior third of folded wings uncovered; elytral pubescence uniform, short and sub-erect. Scutellum transverse, slightly narrowing distally, broadly rounded at apex (Fig. 2). Legs long and slender; femora and tibiae narrow and straight, tibiae noticeably shorter than femora; hind tarsomere length ratio 4.5: 2.5: 1.8: 1: 1.5; tarsomere 4 deeply cleft; all claws simple (Fig. 2). Ultimate sternite elongate, gradually narrowing distally, widened and deeply emarginate at apex, somewhat curved in lateral view; ultimate tergite elongate, slightly narrowed distally, with roundish incision at apex; penultimate tergite transverse, sub-rectangular; third from end tergite simple (Figs 6, 7). Aedeagus roundish, with rounded distally dorsal plate; laterophyses in distal half relatively broad, parallel-sided, with almost rectangular bulges at base, in lateral view slanted outwardly, but not hooked at apex; penis strongly curved, in lateral view (Figs 13, 14). Length (from head to apices of folded elytra): 3.0 mm; width (at humeri): 0.5 – 0.6 mm. FEMALE. Unknown.	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10BFFE4FEFDF8E5FABD028B.taxon	etymology	ETYMOLOGY. The new species is named after the type locality.	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10BFFE4FEFDF8E5FABD028B.taxon	diagnosis	DIAGNOSIS. Malthodes sukkoensis sp. n. resembles M. pseudobesucheti Wittmer, 1970, from the mountains of Krasnodar Krai and Abkhasia, differing in the relatively narrowly incised at apex ultimate sternite, shorter and not widened at apex ultimate tergite, more rounded dorsal plate of the aedeagus and not hooked apically, in lateral view, laterophyses (Figs 6, 7, 13, 14), vs broadly widened at apex ultimate sternite, longer and widened at apex ultimate tergite, more narrowed distally dorsal plate of the aedeagus and apically hooked laterophyses, in lateral view, in M. pseudobesucheti (Figs 8, 9, 15, 48, 49, 87, 88).	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A10BFFE4FEFDF8E5FABD028B.taxon	discussion	REMARKS. The paratype lacks head and pronotum.	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A102FFE8FCBBFE63FD7906BD.taxon	type_taxon	Type species: Malthinus marginatus Latreille, 1806.	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
5B7587A9A102FFE8FCBBFE63FD7906BD.taxon	discussion	Malthodes lobicollis (Reitter, 1888) described from ‘ Circassien’ based on several females [Reitter, 1888], whereas all the taxonomy of the genus is based on the morphological characters of males (shape and structures of the terminal abdominal segments and aedeagi), cannot be placed amidst its congeners properly. Therefore it is included in the list only as incertae sedis. The following three species are excluded from the list of the Greater Caucasian Malthodes: Malthodes debilis Kiesenwetter, 1852. Distributed from West to East Europe [Delkeskamp, 1977; Wittmer, 1992; Kazantsev, Brancucci, 2007]. One specimen is reported from the Caucasus: ’ Parabotch Forest, Ki [z] l., Tersk distr. ’ [Wittmer, 1992]. The taxon is excluded from the list, because the locality where the only ‘ Caucasian’ specimen was registered (Kizlyar distr., northern Dagestan), does not belong to the Greater Caucasus. Malthodes quadristilus Wittmer, 1992. The taxon is excluded from the list, as its type locality (Terekli-Mekteb, northern Dagestan — Wittmer, 1992) does not belong to the Greater Caucasus. The second known specimen of M. quadristilus (from Krasnodar Krai — Kazantsev, 2011; 2022), after being re-studied, has proven to belong to the closely related M. orientalicus Wittmer, 1970. Malthodes tordi Wittmer, 1970. Malthodes tordi, described from Turkey (Trabzon) [Wittmer, 1970], was later indicated for Abkhazia and Adjara [Wittmer, 1992]. However, the Abkhazian ‘ M. tordi ’ proved to be a related, but different species — M. jaromiri Švihla, 2002 [Švihla, 2002], while the re-examination of the Adjara specimens, from the ‘ M. tordi ’ series identified by Wittmer, showed that they also belong to another species, M. vladimiri Kazantsev, 2021 [Kazantsev, 2021 b]. Therefore, M. tordi is excluded from the regional list as well.	en	Kazantsev, S. V. (2024): New species of Malthodes Kiesenwetter, 1852 (Coleoptera: Cantharidae) from the Greater Caucasus, with notes on distribution of the genus in the area. Russian Entomological Journal 33 (1): 79-96, DOI: 10.15298/rusentj.33.1.08, URL: https://doi.org/10.15298/rusentj.33.1.08
