taxonID	type	description	language	source
585B879BFF9AB65D53DA14B8FE389A71.taxon	biology_ecology	E. Gardens, pastures, cultivated soils, forests, floodplains and near freshwater (Šapkarev 1978 a). D. Demir Kapija (Čelavec izvir) (Karaman 1971); Gazi Baba (Šapkarev 1970); Mavrinci (Šapkarev 1971); Bitola, Bratin dol, Dolno Divjaci, Kruševo, Lerin, Nikola Karev, Prilep, Železnec (Šapkarev 1972 a); Belovodica, Drenovo, Rakle, Rosoman (Šapkarev 1974); Brajčino, Kriveni, Ljubojno, Oteševo, Gorno Perovo, Podmočani, Pretor, Resen, Sirhan, Stenje (Šapkarev 1976); Bukovo, Labuništa, Ohrid, Trebeništa, Velgošti, Vevčani (Šapkarev 1977); Dojran, Gevgelija, Marvinci, Miletkovo (Šapkarev 1980); Demir Kapija (Mršić 1991). Z. Peregrine species native to the Palearctic.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9AB65D53DA14B8FE389A71.taxon	materials_examined	New records. HNHM / 17491 1 ex., Peštani, bottom of the rock on the lakeshore, 41 ° 0.581 ' N, 20 ° 48.360 ' E, 712 m, Hedera bush, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi. HNHM / 17502 3 ex., Ohrid, south of the city, 41 ° 5.617 ' N, 20 ° 48.059 ' E, 693 m, willow trees on the lakeshore, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9AB65D53DA1712FB3D9D88.taxon	biology_ecology	E. Pastures and meadows of hills and mountains (Šapkarev 1974, 1976). D. Demir Kapija (Karaman 1969; Šapkarev 1974; Mršić 1991); Krastavac near Tunel (Karaman 1969, 1971); Galičica (Šapkarev 1976; Zicsi 1982). Z. Broad range Balkanic endemic species registered in North Macedonia and Albania (Szederjesi 2023).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9AB65D53DA1148FB9B9F60.taxon	biology_ecology	E. Pastures and meadows of hills and mountains (Šapkarev 1977 a, 1978 a). D. Pološki Manastir (Šapkarev 1977 a). Z. Narrow range Balkanic endemic species (known only from the type locality).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9AB65D53DA12E0FB9B9ED8.taxon	biology_ecology	E. Pastures and meadows of hills and mountains (Šapkarev 1977 b, 1978 a). D. Kožuf Mountain (Šapkarev 1977 b). Z. Narrow range Balkanic endemic species (known only from the type locality).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9DB65A53DA14F0FD0698E9.taxon	biology_ecology	E. Meadows (Šapkarev 1971; 1978 a). D. Rašče near Skoplje (Šapkarev 1971). Z. Narrow range Balkanic endemic species (known only from the type locality).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9DB65A53DA1632FE939CD9.taxon	biology_ecology	E. Arable land, beech and oak forests, river banks, humid meadows and pastures (Šapkarev 1978 a; Szederjesi 2019). D. Dedeli, Kalkovo (Ude 1922); Nakolec near Prespa, Studenčište near Ohrid (Černosvitov 1931); Dedeli, Kalkovo, Demir Kapija (Karaman 1969); Volkovo, Petrovec, Skoplje, Karadzica, Matka, Kisela Voda, Nerezi, Raduša (Šapkarev 1970); Vrutok, Gostivar, Tetovo, Radiovce, Jegunovce, Vratnica, Bojane (Šapkarev 1972 a); Skočivir, Konjarska reka, Zhvan, Dolno Divjaci, Bitola, Kruševo, Lerin, Nikola Karev, Prilep (Šapkarev 1972 b); Lakavica (Šapkarev 1973); Belovodica, Drenovo, Rakle, Rosoman, Vataša, Besvica, Kavadarci, Demir Kapija, Negotino, Vozarci, Šeškovo, Palikura, Dradnja (Šapkarev 1974); Brajčino, Kriveni, Ljubojno, Oteševo, Gorno Perovo, Podmočani, Pretor, Resen, Sirhan, Stenje (Šapkarev 1976); Bukovo, Labuništa, Ohrid, Trebeništa, Velgošti, Vevčani (Šapkarev 1977 a); Dojran, Gevgelija, Marvinci, Miletkovo (Šapkarev 1980); Makedonska Kamenica, Peščevo, Mitrašinci, Virče, Budinarci, Ratevo, Rusinovo, Negrevo, Crnik, Kalimanci, Smojmirovo, Vladimirovo (Šapkarev 1983); Demir Kapija, Kosturino, Krklja, Krstov Dol, Staro Nagoričane, Rašče, Strmoš (Mršić 1991); Bašibos, Bogdanci, Dedeli (Szederjesi 2019). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9DB65A53DA1632FE939CD9.taxon	materials_examined	New record. HNHM / 17494 3 ex., at the road between Kavadarci and Prilep, ~ 2 km west of Trojaci, 41 ° 21.960 ' N, 21 ° 43.020 ' E, 678 m, dry gorge with bushy vegetation, 25.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9DB65A53DA11BAFB999E1F.taxon	biology_ecology	E. Moist soils, floodplains, and humid meadows (Šapkarev 1978 a; Szederjesi 2013). D. Vataša, Murtino, Bašibos, Zleševo, Prdejci, Gevgelija, Neprošteno, Mavrinci, Jegunovce, Radiovce (Šapkarev 1971); Rečani, Jegunovce, Radiovce, Neprošteno (Šapkarev 1972 a); Murtino (Šapkarev 1973); Vataša (Šapkarev 1974); Vataša, Marvinci, Prdejci (Šapkarev 1980); Rabrovo (Szederjesi 2013). Z. Atlanto-Mediterranean.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9CB65B53DA14F0FE939BE7.taxon	biology_ecology	E. Arable land, beech and oak forests, river banks, humid meadows, and pastures (Šapkarev 1978 a; Szederjesi 2013; 2019). D. Crničani, Konopište, Bošava, Radoviš, Valandovo, Furka (Šapkarev 1971); Radoviš, Strumica (Šapkarev 1973); Demir Kapija, Konopište (Šapkarev, 1974); Brajčino (Šapkarev 1976); Lakavica, Radoviš (Šapkarev 1979); Furka, Crničani, Valandovo (Šapkarev 1980); Trabovište (Šapkarev 1983); Demir Kapija, Kolešino (Szederjesi 2013); Čalakli (Szederjesi 2019). Z. Trans-Aegean.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9CB65B53DA14F0FE939BE7.taxon	materials_examined	New record. HNHM / 17499 5 ex., Kjesendre, at 1.3 km on the road to Galište, 41 ° 23.423 ' N, 21 ° 50.454 ' E, 680 m, at a small stream in a young beech-oak-maple shrub, 25.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9CB65B53DA1763FF759DC0.taxon	biology_ecology	E. Forests, meadows, cultivated soils, stream in grazed pasture (Šapkrev 1978 a; Szederjesi 2019). D. Bojane, Miletkovo (Šapkarev 1972 a); Radoviš, Murtino (Šapkarev 1973); Demir Kapija, Rakle, Palikura (Šapkarev 1974); Ohrid (Šapkarev 1977 a); Dolni Balvan, Kardifakovo, Lozovo (Šapkarev 1979); Dojran, Marvinci, Miletkovo (Šapkarev 1980); Peščevo, Delčevo (Šapkarev 1983); Dzvegor (Mršić 1991); Bogdanci (Szederjesi 2019). Z. Trans-Aegean.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9CB65B53DA1080FB2B9EBB.taxon	biology_ecology	E. River banks, meadows, pastures, caves, beech and oak forests, cultivated soils (Šapkarev 1978 a; Szederjesi 2019). D. Dedeli (Ude 1922; Karaman 1971); Krastavac (Karaman 1969); Dedeli, Gazi Baba, Raduša, Dolno Divjaci, Jegunovce, Miletino, Neprošteno (Šapkarev, 1970); Galičica (Zicsi 1972); Nikola Karev, Prilep, Rastojca, Raven, Tetovo, Bratin dol, Lerin (Šapkarev 1972 b); Murtino, Strumica, Bansko, Novo Selo, Radoviš (Šapkarev 1973); Besvica, Vataša, Rakle, Rosoman, Debrishte, Demir Kapija (Šapkarev 1974); Brajčino, Resen, Kriveni, Ljubojno, Oteševo, Sirhan, Stenje, Pretor, Makazi, Izbišta (Šapkarev, 1976); Bukovo, Galičica, Ohrid, Trebeništa, Velgošti, Vevčani, Gorenci, Trpetsa, Kališta, Gorno Lakocherej (Šapkarev 1977 a); Dolni Balvan, Gorni Balvan, Kopačevo, Lakavica, Nemaninci (Šapkarev 1979); Bašibos, Crničani, Dojran, Furka, Gevgelija, Marvinci, Sirčino, Kožuf Mountain (Šapkarev 1980); Makedonska Kamenica, Peščevo (Šapkarev, 1982); Berovo, Vladimirovo, Makedonska Kamenica, Peščevo, Mitrašinci, Virce, Budinarci, Ratevo, Rusinovo, Negrevo, Crnik, Kalimanci, Smojmirovo (Šapkarev 1983); Čiflik, Makedonska Kamenica, Negrevo, Mitrašinci, Crnik, Smojmirovo Virce (Karamam 1987); Krstov Dol, Krklja, Staro Nagoričane (Mršić 1991); Bašibos (Szederjesi 2019). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9FB65853DA14F0FEE8988F.taxon	biology_ecology	E. Pastures, oak forests, subalpine meadows (Šapkarev 1980). D. Šar Mountain (Elak, Lešnica) (Šapkarev 1980). Z. Alpine-Dinaric.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9FB65853DA164BFE939DC0.taxon	biology_ecology	E. Pastures, anthropogenic soils, gardens, forests (Šapkarev 1977 a, Šapkarev 1983). D. Dedeli, Kaluckovo (Ude 1922); Nakolec near Prespa, Studenčište near Ohrid (Černosvitov 1931); Dedeli, Kaluckovo (Karaman 1969); Kos Bunar, Šipkovica, Plačkovica, Murtino, Strumica, Bansko, Novo Selo, Radoviš, Damjan (Šapkarev 1973); Bukovo, Labuništa, Ohrid, Trebeništa, Velgošti, Gorenci, Galičica, Vevčani, Velestovo, Trebeništa, Studenčišta, Trpejca, Mešeišta, Leskoec, Rača (Šapkarev 1977 a); Mavrovo (Trnica) (Šapkarev 1978); Dolni Balvan, Gorni Balvan, Kopačevo, Lakavica, Nemaninci, Buchishte, Junuzlija, Kardifakovo, Krupište, Lezovo, Pishica, Selce (Šapkarev 1979); Pehčevo, Makedonska Kamenica, Mitrašinci, Berovo, Virče, Budinarci, Ratevo, Rusinovo, Negrevo, Smojmirovo, Vladimirovo, Kalimantsi (Šapkarev 1983). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9FB65853DA164BFE939DC0.taxon	materials_examined	New record. HNHM / 17497 1 ex., Kjesendre, at 1.3 km on the road to Galište, 41 ° 23.423 ' N, 21 ° 50.454 ' E, 680 m, at a small stream in a young beech-oak-maple shrub, 25.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9FB65853DA1148FE669E2C.taxon	biology_ecology	E. Mixed deciduous forest, under the bark of fallen logs, near a stream (Šapkarev 1977 a; Šapkarev 1980; Szederjesi 2013; 2019). D. Kališta (Černosvitov 1931; Karaman 1969); Sv. Naum, Trnica (Mavrovo) (Šapkarev 1972 b); Ostrovo near Sv. Naum, Jesenice (Šapkarev 1977 a); Mavrovo (Trnica) (Šapkarev 1978 b); Lukovo, Visoka Čuka (Kožuf Mts.) (Šapkarev 1980); Veles, Gostivar (Szederjesi 2013); Bašibos (Belasica Mts.) (Szederjesi 2019). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9FB65853DA1148FE669E2C.taxon	materials_examined	New records. HNHM / 17487 5 ex., Nežilovo, Jakupica Mts., 0.5 km SE of the Babuna Spring, 41 ° 41.248 ' N, 21 ° 25.224 ' E, 1200 m, rocky beech forest, 24.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi. HNHM / 17496 1 ex., Nežilovo, Jakupica Mts., Babuna Spring, 41 ° 41.397 ' N, 21 ° 24.979 ' E, 1360 m, beech forest, 24.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi. HNHM / 17509 2 ex., between Trpejca and Carina, E slope of Galičica Mts., 40 ° 58.047 ' N, 20 ° 51.585 ' E, 1380 m, old beech forest, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9EB65953DA14F0FABD9A99.taxon	biology_ecology	E. Wooded areas, gardens, under logs; habitats with high organic matter (Šapkarev 1978 a; Szederjesi 2013, 2019); D. Dolno, Kosovrasti (Karaman 1969); Skopska Crna Gora (Šapkarev 1970); Visoka Čuka (Kožuf Mts.), Čeples, Skoplje, Dojran, Botun (Šapkarev 1972 c); Skoplje, Vrutok, Ohrid, Vratnica, Kruševo (Šapkarev 1972 d); Čiflik, Demir Kapija, Drenovo, Konopište (Šapkarev 1974); Botun, Ohrid, Studenčišta, Stogovo (Šapkarev 1977); Gevgelija, Bašibos (Šapkarev 1980); Makedonska Kamenica, Peščevo, Čiflik, Berovo, Vladimirovo, Rusinovo, Crnik (Šapkarev 1983); Osogovo, Hamzali, Kožuf Mts. (Mršić 1991); Tetovo, Popova Šapka (Šar Mts.), Sv. Naum near Ohrid Lake, Reka Mts. (Szederjesi 2013); Bašibos, Stari Dojran (Szederjesi 2019). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9EB65953DA14F0FABD9A99.taxon	materials_examined	New record. HNHM / 17510 2 ex., between Trpejca and Carina, E slope of Galičica Mt., 40 ° 58.047 ' N, 20 ° 51.585 ' E, 1380 m, old beech forest, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9EB65953DA1001FB789F0D.taxon	biology_ecology	E. Mixed forests and pastures (Šapkarev 1980) D. Vodno (Ude 1922); Krastavac near Demir Kapija, Vodno (Karaman 1969); Vrteška (Pčinja river) (Šapkarev 1970); Strumica (Šapkarev 1973); Bitola, Demir Kapija (Šapkarev 1974); Dojran, Kosturino (Šapkarev 1980); Kalimanci, Pehčevo, Ponikva (Šapkarev 1983); Gazi Baba (Šapkarev 1984); Dojran, Demir Kapija, (Karaman & Stojanović 1996); Gazi Baba, Vodno, Katlanovo (Šapkarev 1994); Katlanovo-Rudnik, Beli Kočani, Probištip, Staro Nagoričane (Mršić 1991). Z. Vardar endemic species registered in North Macedonia, Serbia (Stojanović et al. 2018; Popović et al. 2022) and Greece (Szederjesi et al. 2017).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9EB65653DA12CDFD5499AC.taxon	biology_ecology	E. Meadows of hills and mountains (Stojanović et al. 2016). D. Gostivar (Stojanović et al. 2016). Z. Vardar endemic species registered in North Macedonia and Serbia (Trakić et al. 2016; Stojanović et al. 2018; Stojanović-Petrović et al. 2020).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF9EB65653DA12CDFD5499AC.taxon	discussion	Remarks. This taxon was originally described as All. robusta serbica Karaman, 1983. According to Popović et al. (2022), several species treated as Serbiona (eg., S. robusta) by Mršić (1991) belong to the genus Cernosvitovia, thus we place this species to Cernosvitovia as well.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF91B65653DA152CFC3E9B3A.taxon	biology_ecology	E. Pasture (Šapkarev 1973). D. Strumica (Šapkarev 1973); Kosturino, Bjelasnica (Mršić 1991). Z. Vardar endemic species registered in North Macedonia and Serbia (Popović et al. 2022).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF91B65653DA16A6FDCE9A92.taxon	biology_ecology	E. Humid meadows (Mršić 1991). D. Treska (Mršić 1991). Z. Vardar endemic species registered in North Macedonia and Serbia (Popović et al. 2020, 2022).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF91B65753DA107EFD4F9848.taxon	biology_ecology	E. Pastures and meadows of hills and mountains (Šapkarev 1991). D. Plešenci, Krstov Dol, Kriva Palanka (Šapkarev 1972 d; Mršić 1991; Šapkarev 1991). Z. Vardar endemic species (known only from the type locality).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF91B65753DA107EFD4F9848.taxon	discussion	Remarks. The taxonomical status of this taxon is quite confusing. The description was published by Šapkarev (1972 d) as Allolobophora sp. however, he subjected the naming to a condition: “ if it is a new taxon, I would name it as udei ”, thus this name is not available (ICZN Art. 15.1.). The first descriptions that made the name available are from Mršić (1991) and Šapkarev (1991), as Serbiona udei and Allolobphora dofleini udei respectively. Blakemore (2008), due to the inaccessibility of publications of Šapkarev (1972 d), saw that Mršić (1991) included Allolobophora udei Sapkarev in the synonymy list (as nomen nudum). Also, wrongly, because that name was not available, Mršić should have written Allolobophora sp. Šapkarev, 1972: 120. Hence, Blakemore (2008) provided Serbiona joncesapkarevi a replacement name for All. udei) and Serbiona josapi a replacement name for All. dОfleini udei), to avoid homonymy of Allolobophora parva udei Ribaucourt, 1886. However, Allolobophora udei Šapkarev, 1972 is not an available name therefore can not fall into homonymy nor synonymy. The same mistake was made by Popovic et al. (2024). According to the International Code of Zoological Nomenclature (ICZN Art. 24.1., 50.1.), we accept Serbiona udei Mršić, 1991 as the senior synonym and first valid combination, with the first complete description of the species. Thus, Allolobophora dofleini udei Šapkarev, 1991, Serbiona josapi Blakemore, 2008 and Allolobophora joncesapkarevi Blakemore, 2004 are homotypic synonyms of S. udei, as all four namesare based on the same type specimens. Since Popović et al. (2022) revealed that several morphologically similar species treated as Serbiona by Mršić (1991) (eg., S. dofleini, S. robusta, S. serbica, S. paratuleskovi) belong to the genus Cernosvitovia, we place S. udei to Cernosvitovia as well. This is not the only case of unclear taxonomic status of species from the Balkans. The intricate history of the Cernosvitovia crainensis (Mršić, 1989) and Aporrectodea macvensis Šapkarev in Mršić, 1991 was explained by Nicholas & Csuzdi (2018).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF90B65753DA15D1FE669AEC.taxon	biology_ecology	E. Forest, humid soil, pasture (Šapkarev 1977 a, 1983; Szederjesi 2013). D. Jablanica (near Golemo Vevčansko Lake) (Šapkarev 1971); Galičica (Zicsi 1972); Jablanica (Šapkarev 1977 a); Trabotivište, Sasa, Ponikve (Šapkrev 1983); Osogovske Mts. (Carev vrh) (Mršić 1991); Ogražden Mts. (Szederjesi 2013). Z. Balkanic-Alpine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF90B65753DA15D1FE669AEC.taxon	materials_examined	New records. HNHM / 17492 2 ex., between Trpejca and Carina, pass of Galičica Mt., 40 ° 57.333 ' N, 20 ° 49.099 ' E, 1595 m, subalpine pasture with Crataegus, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi. HNHM / 17495 1 ex., Nežilovo, Jakupica Mts., Babuna Spring, 41 ° 41.397 ' N, 21 ° 24.979 ' E, 1360 m, beech forest, 24.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi. HNHM / 17508 2 ex., between Trpejca and Carina, E slope of Galičica Mt., 40 ° 58.047 ' N, 20 ° 51.585 ' E, 1380 m, old beech forest, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF90B65753DA106DFEDB9C4E.taxon	biology_ecology	E. Beech forest (Szederjesi 2013). D. Šar Mts. (Ceri Pašina), Lešnica, Elak (Šapkrev 1971); Lešnica, Jelak, Popova Šapka, Ceri Pašina (Šapkarev 1972 a); Gostivar (Szederjesi 2013). Z. Broad range Balkanic endemic species distributed in Romania, Bosnia-Herzegovina, North Macedonia, Montenegro and Albania (Sekulić et al. 2022; Szederjesi 2023).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF90B65753DA110AFAD99F94.taxon	biology_ecology	E. Forests, meadows, leaf litter, adapted to acid sandy soils (Šapkarev 1978 a; Szederjesi 2013). D. Nidze Mts. (Šapkarev 1972 b); Trpejca (Šapkarev 1977 a); Sasa (Mršić 1991); Galičica Mts., Gorno Jelovce, Ogražden Mts., Peštani (Szederjesi 2013). Z. Trans-Aegean.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF90B65753DA110AFAD99F94.taxon	materials_examined	New record. HNHM / 17488 2 ex., Nežilovo, Jakupica Mts., 0.5 km SE of the Babuna Spring, 41 ° 41.248 ' N, 21 ° 25.224 ' E, 1200 m, rocky beech forest, 24.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF90B65453DA1375FAB69814.taxon	biology_ecology	E. Stream banks (Karaman 1969). D. Near Čelavec River (Karaman 1969). Z. According to Csuzdi & Zicsi (2003) this species belongs to Eastern-Alpine species occurring in Austria, Hungary, Slovakia, Czech Republic, Poland and Ukraine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF90B65453DA1375FAB69814.taxon	materials_examined	New record. HNHM / 17503 1 ex., Demir Kapija, at the entrance of the Demir Kapija Canyon, 41 ° 24.171 ' N, 22 ° 15.504 ' E, 135 m, macchia, 28.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF90B65453DA1375FAB69814.taxon	discussion	Remarks. The new record represents the second occurrence of D. auriculata from North Macedonia.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF93B65453DA15F5FE389DE0.taxon	biology_ecology	E. Forest soils, meadows and river banks (Šapkarev 1978 a; Szederjesi 2019). D. Doline Nikola, Kobelija (Ude 1922); Kaluckova, Ohrid, Kosel, Prespa, Kališta, Šar Mts., Debar, Kolibarci near Sv. Naum (Černosvitov 1931; Karaman 1969; 1971); Skopska Crna Gora, Karadžica (Šapkarev 1970); Bukovo, Ceri Pašina, Tearce, Dolno Divjaci, Duv, Nidze Mts., Kruševo, Jelak, Vrutok, Gostivar, Tetovo, Radiovce, Jegunovce, Vratnica, Bojane, Gorno Divjaci, Volkovo, Skočivir, Pelister, Dolna Lešnica, Lavci, Lisec, Ljuboten (Šapkarev 1972 a); Plačkovica, Šipkovica, Kos Bunar, Bansko (Šapkarev 1973); Drenovo, Kavadarci, Demir Kapija, Konopište (Šapkarev 1974); Brajčino, Kriveni, Ljubojno, Oteševo, Sapunčica, Sirhan (Šapkarev 1976); Lukovo, Mali Vlaj, Manastirec, Godivje, Galičica, Ogražden Mts., Ohrid, Trebeništa, Velgošti, Vevčani, Gorenci, Trpetsa, Kališta, (Šapkarev 1977 a); Petrovo (Visoka čuka), Bogdanci, Kožuf Mts., (Mrška River, Asan česma, Flora-Dve Uši, Golema poljana, Lukovo, Moskova voda, Visoka česma, Visoka čuka) (Šapkarev 1980); Pehčevo, Ratevo, Rusinovo, Sasa, Ponikve (Šapkarev 1983); Jablanica Mts. (Labuništa), Šar Mts. (Vešala), Kožuf Mts. (near Ski Kožuf) (Szederjesi 2019). Z. Circum-Mediterranean species.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF93B65453DA15F5FE389DE0.taxon	materials_examined	New records. HNHM / 17486 2 ex., Valandovo, Hotel Izvor, 41 ° 19.633 ' N, 22 ° 33.328 ' E, 258 m, Platanus gallery along a stream, 28.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi. HNHM / 17498 1 ex., Kjesendre, at 1.3 km on the road to Galište, 41 ° 23.423 ' N, 21 ° 50.454 ' E, 680 m, at a small stream in a young beech-oak-maple shrub, 25.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi. HNHM / 17500 1 ex., Ohrid, south of the city, 41 ° 5.617 ' N, 20 ° 48.059 ' E, 693 m, willow trees on the lakeshore, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF93B65453DA1239FAF59E2A.taxon	biology_ecology	E. under stone on stream bank (Šapkarev 1970). D. Skopska Crna Gora (Šapkarev 1970). Z. Central European.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF92B65553DA15D6FE389AC5.taxon	biology_ecology	E. Cultivated and humid soil, under stones, brooks in alpine grasslands and beech forests, stream bank, springs (Šapkarev 1977 a; Szederjesi 2013); D. Ohrid, Vratnik, Velestovo, Mali Vlaj, Velgošiti, Gari, Ostrovo, Gorno Lakočerej, Gorenci, Ramne (Šapkarev 1977 a); Gorenci, Gari (Šapkarev 1978 b); Peštani, Resen (Krani), Svet Naum, Pelister Mts. (Nižepole) (Szederjesi 2013). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF92B65553DA15D6FE389AC5.taxon	materials_examined	New records. HNHM / 17490 1 ex., Peštani, bottom of the rock on the lakeshore, 41 ° 0.581 ' N, 20 ° 48.360 ' E, 712 m, Hedera bush, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi. HNHM / 17506 3 ex., Ohrid, Studenčišta, 41 ° 6.137 ' N, 20 ° 48.888 ' E, 700 m, park with plane trees, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF92B65553DA1786FB6C9D9C.taxon	biology_ecology	E. Beech forest with a brook (Szederjesi 2013). D. Ogražden Mt. (Szederjesi 2013). Z. Broad range Balkanic endemism. This species was recorded in Greece, Bulgaria and North Macedonia (Szederjesi 2013).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF92B65553DA117CFC5E9F31.taxon	biology_ecology	E. Caves and pastures (Šapkarev 1971; Karaman, 1972 a). D. Kožuf Mts. (Šapkarev 1971, 1980); Nerodova Cave, Elasov Do (Karaman 1972 a). Z. Broad range Balkanic endemic species distributed in North Macedonia, Serbia (Stojanović et al. 2018) and Montenegro (Stojanović & Milutinović 2013).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF92B65553DA12D2FEDC9EC8.taxon	biology_ecology	E. Pasture (Mršić 1991). D. Osogovska Mts. (Mršić 1991). Z. Narrow range Balkanic endemism (known only from the type locality).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF95B65253DA14F0FB5C9B5B.taxon	biology_ecology	E. Forest, meadows under fallen leaves, fallen trunks (Šapkarev 1978; Szederjesi 2013). D. Šar Mts., Ohrid, Kališta (Černosvitov 1931; Karaman 1971); Ljuboten, Duv, Jelak (Šapkarev 1972 a); Pehčevo, Kožuf Mts. (Moskova voda) (Šapkarev 1983); Osogovske Mts. (Mršić 1991); Jakupica Mt. (Bogomila), Šar Mts. (Tetovo-Popova Šapka) (Szederjesi 2013). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF95B65253DA14F0FB5C9B5B.taxon	materials_examined	New record. HNHM / 17504 1 ex., ~ 1 km N of Izbišta, at the road between Resen and Ohrid, 41 ° 8.400 ' N, 21 ° 0.048 ' E, 950 m, oak forest, 25.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF95B65253DA1607FE649A86.taxon	biology_ecology	E. Mountain pastures (Šapkarev 1972 c). D. Crni Vrh (Probištip), Nidže Mts. (Šapkarev 1972 c). Z. Broad range Balkanic endemic species distributed in North Macedonia and Greece (Szederjesi et al. 2017).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF95B65253DA1607FE649A86.taxon	discussion	Remarks. Szederjesi (2017) noticed that the subspecies Dendrobaena olympica peristerica described by Šapkarev (2001) has identical taxonomic characters as D. olympica. Therefore D. olympica peristerica is regarded as a synonym of D. olympica.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF95B65253DA1043FD0A9C1F.taxon	biology_ecology	E. Stream banks, under stones (Šapkarev 1971). D. Bošava River, Demir Kapija (Karaman 1969); Skopska Crna Gora (Šapkarev 1971). Z. Central European.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF95B65253DA11FCFDAE9F31.taxon	biology_ecology	E. Pasture (Šapkarev 1976). D. Velmej in Ilinska Bigla Mts. (Šapkarev 1976). Z. Narrow range Balkan endemism (known only from the type locality).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF95B65353DA12D2FE5399AC.taxon	biology_ecology	E. Beech forest with a brook (Šapkarev 1983). D. Sasa (Šapkarev 1983). Z. Narrow range Balkan endemism (known only from the type locality).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF95B65353DA12D2FE5399AC.taxon	discussion	Remarks. Šapkarev (1983) published and described this taxon as Dendrobaena sp., however, he subjected the naming to a condition: “ if further studies prove that this is a new species, then it should be named as sasensis ”, thus this name remained not available. Mršić (1991) was the first, who publish the description of this taxon under the name Dendrobaena sasensis Šapkarev, 1983, which made the name available, although he attributed the description entirely to Šapkarev (1983).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF94B65353DA152DFC009A4B.taxon	biology_ecology	E. Spring lake, grassland and pine forest (Šapkarev 1978 a; Szederjesi 2013). D. Skoplje (Šapkarev 1970); Železnic (Šapkarev 1972 b); Botun, Gari, Godivje, Gorenci, Mali Vlaj, Ohrid, Velestovo, Velgošti, Vevčani (Šapkarev 1972 d); Botun, Godivje, Gorenci, Mali Vlaj, Ohrid, Velestovo, Velgošti, Vevčani, Rača, Studenčišta, Sv. Ilija, Sv. Petka (Šapkarev 1977 a); Peštani, Sv. Naum (Szederjesi 2013). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF94B65353DA152DFC009A4B.taxon	materials_examined	New record. HNHM / 17507 2 ex., Ohrid, Studenčišta, 41 ° 6.137 ' N, 20 ° 48.888 ' E, 700 m, park with plane trees, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF94B65353DA17DFFC009C27.taxon	biology_ecology	E. Anthropogenic biotopes, beech forests, meadows, and stream banks (Šapkarev 1971; Szederjesi 2013, 2019). D. Gazi Baba, Konjare near Pčinja River (Šapkarev 1971); Vratnica (Šapkarev 1972 a); Lavci, Bitola, Kisela Voda, Kruševo, Lerin (Šapkarev 1972 b); Konopište, Bošava (Šapkarev 1974); Leva River (Šapkarev 1976); Velestovo, Ohrid (Šapkarev 1977 a); Makedonska Kamenica (Šapkarev 1983); Reka Mts. (Vrben) (Szederjesi 2013); Demir Kapija (Szederjesi 2019). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF94B65353DA17DFFC009C27.taxon	materials_examined	New record. HNHM / 17505 3 ex., Ohrid, Studenčišta, 41 ° 6.137 ’ N, 20 ° 48.888 ’ E, 700 m, park with plane trees, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF94B65353DA11A3FEE59F85.taxon	biology_ecology	E. Mixed forest, old beech forest, oak forest, valley meadow, streem, and pastures (Šapkarev 1971, 1973; Szederjesi 2013). D. Ljuboten (Šapkarev 1971); Plačkovica (Šapkarev 1973); Osogovski Mts. (Sasa) (Szederjesi 2013). Z. Central European.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF94B65053DA130EFE21986C.taxon	biology_ecology	E. Still and running water (Šapkarev 1962) D. Prespansko jezero (Sirhan) (Černosvitov 1931; Šapkarev 1962). Z. Circum-Mediterranean.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF97B65053DA15ECFEB29BA6.taxon	biology_ecology	E. Mud of fresh waters (Šapkarev 1963; Karaman 1969, 1971). D. Sv. Naum, Ohrid (Černosvitov 1931); Ohrid Lake (Šapkarev 1963, 1977 a; Karaman 1969, 1971;). Z. Narrow range Balkan endemism (known only from the type locality).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF97B65053DA1723FC209D1E.taxon	biology_ecology	E. Mud of fresh waters (Karaman 1969). D. Ohrid Lake (Černosvitov 1931; Karaman 1969; Šapkarev 1977 a). Z. Narrow range Balkan endemism (known only from the type locality).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF97B65053DA10FBFBBE9E86.taxon	biology_ecology	E. Limicolous / hydrophilic, in moist, damp habitats, rivers, streams or pond banks (Šapkarev 1978 a; Szederjesi 2019). D. Ohrid, Rabrovo, Struga, Sv. Naum, Šar Mts. (Černosvitov 1931); Ohrid Lake, Prespa Lake (Šapkarev 1956, 1963, 1964); Crni kamen, Debar, Demir Kapija, Dolina Nikole, Markov kamen, Rabrovo, Šar Mts. (Karaman 1969, 1971); Katlanovo, Kitka, Raduša, Skoplje, Skopska Crna Gora, Taor, Vodno, Konjare, Butel (Šapkarev 1970); Bitola, Gostivar, Sv. Naum, Tetovo, Ceri pašina, Čelopek, Dolenci, Dolno Divjaci, Duv, Gorno Divjaci, Jegunovce, Jelak, Lešnica, Kačanik, Kruševo, Ljuboten, Miletino, Popova Šapka, Skočivir, Tearce (Šapkarev 1972 c); Babuna, Barovo, Bošava, Čelovec, Demir Kapija, Drenovo, Kavadraci, Konopište, Raec, Rakle, Rosoman, Trojaci-Pletvar (Šapkarev 1974); Brajčino, Gjavato, Kriveni, Ljubojno, Oteševo, Pretor, Sirhan, Staro Selo (Kriveni) (Šapkarev 1976); Belčišta, Botun, Velgošti, Galičica, Godivje, Kališta, Labuništa, Mali Vlaj, Mešeišta, Ograždenik, Ohrid, Pesočan, Petrino, Radožda, Struga, Sv. Naum, Velmej, Vevčani (Šapkarev 1977 a); Debar, Gari, Gorenci (Šapkarev 1978 b); Bašibos, Bogdanci, Dojran, Gevgelija, Kosturino, Kožuf Mts., Miletkovo, Petrovo, Sermelin, Valandovo (Šapkarev 1980); Makedonska Kamenica, Pehčevo (Šapkarev 1983); Bošava, Ohrid Lake (Mršić 1991); Belasica Mts, Bašibos (Szederjesi 2019). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF97B65053DA10FBFBBE9E86.taxon	materials_examined	New record. HNHM / 17489 10 ex., Peštani, bottom of the rock on the lakeshore, 41 ° 0.581 ' N, 20 ° 48.360 ' E, 712 m, Hedera bush, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF96B65153DA14B8FE559BAE.taxon	biology_ecology	E. Streams, limnic habitats are mostly overgrown with vegetation (Šapkarev 1983). D. Rašče (Černosvitov 1931); Rašče, Izvor Toplec near Dojran (Karaman 1969, 1972 b); Skopska Crna Gora (Šapkarev 1970); Štip (Šapkarev 1979); Konsko, Sirično, Sobri, Dojran (Šapkarev 1980), Virče (Šapkarev 1983). Z. Broad range Balkanic endemic species distributed in North Macedonia, Serbia (Stojanović et al. 2018) and Montenegro (Stojanović & Milutinović 2013).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF96B65153DA14B8FE559BAE.taxon	discussion	Remarks. Szederjesi et al. (2022) included several taxa from the Balkans (including balcanicus balcanicus) in Imetescolex for which DNA sequences were not available, but their morphology and geographic distribution suggest their inclusion in this genus.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF96B65153DA17F3FE069D29.taxon	biology_ecology	E. In humid soils rich in organic matter, mostly in forests under the litter (Šapkarev 1972 a). D. Ljuboten (Šapkarev 1972 a). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF96B65153DA10AAFEC59CA8.taxon	biology_ecology	E. Humid soil (Šapkarev 1972 d). D. Bogomila (Šapkarev 1972 d). Z. According to Csuzdi et al. (2011) L. improvisus belongs to the Carpathian endemic species. However, Šapkarev (1972 d, 1978 a) and Stojanović et al. (2018) reported L. improvisus from North Macedonia and Serbia, which indicates that the zoogeographical status of this species has to be reviewed.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF96B64E53DA1228FCA398F8.taxon	biology_ecology	E. In forest and cultivated soils, meadows, streams, and pastures (Šapkarev 1978 a; Szederjesi 2019). D. Vodno (Ude 1922); Zvončica near Debar (Černosvitov 1931); Zvončica near Debar, Bošava River (Karaman 1969, 1971); Dračevo, Katlanovo, Kitka, Lisiče, Novo Selo, Raduša, Skoplje, Skopska Crna Gora, Taor, Vodno (Šapkarev 1970); Bitola, Gostivar, Sv. Naum, Tetovo, Ceri Pašina, Čelopek, Dolenci, Dolno Divjaci, Duv, Gorno Divjaci, Jegunovce, Jelak, Lešnica, Kačanik, Kruševo, Ljuboten, Miletino, Popova Šapka, Skočivir, Tearce (Šapkarev 1972 c); Bansko, Radoviš, Strumica, Barovo (Šapkarev 1973); Belovodica, Čiflik, Demir Kapija, Drenovo, Gorna Bošava, Klisura, Konopište, Pletvar, Rakle, Rosoman, Stobi (Šapkarev 1974); Brajčino, Gjavato, Kriveni, Leva River, Ljubojno, Perovo, Podmočani, Pretor, Resen, Sapunčica-Brajčino, Sirhan, Staro Selo (Šapkarev 1976); Ramne (Velestovo), Studenčišta, Sv. Naum, Trebeništa, Velestovo, Velgošti, Velmej, Vevčani, Vidobišta (Šapkrev 1977 a); Debar, Gorenci, Dolen Balvan, Krupište, Lezovo, Manastirec, Mavrovo (Trnica), Nemaninci (Šapkarev 1978 b); Bašibos, Bogdanci, Dojran, Gevgelija, Kosturino, Kožuf Mts., Marvinci, Rabrovo, Sermelin, Siričino (Šapkarev 1980); Makedonska Kamenica, Pehčevo (Šapkarev 1982); Mačevo, Bregalnica, Pehčevo, Čiflik, Makedonska Kamenica, Mitrašinci, Berovo, Budinarci, Sasa, Negrevo, Crnik, Smojmirovo, Vladimirovo (Šapkarev 1983); Veles, Demir Kapija, Kriva Palanka, Sasa, Smolari, Staro Nagoričane, Dzvegor (Delčevo), Belasica Mts. (Bašibos), Valandovsko Polje (Čalakli), Bogdanci (Szederjesi 2019). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF89B64E53DA1599FC7B9A5F.taxon	biology_ecology	E. Meadows (Šapkarev 1977 a). D. Ohrid (Šapkarev 1977 a). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF89B64E53DA1599FC7B9A5F.taxon	materials_examined	New record. HNHM / 17501 1 ex., Ohrid, south of the city, 41 ° 5.617 ' N, 20 ° 48.059 ' E, 693 m, willow trees on the lakeshore, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF89B64E53DA17C4FF6A9DFF.taxon	biology_ecology	E. Pastures (Šapkarev 1972 c). D. Jegunovce (Šapkarev 1972 c); Mavrovo (Trnica) (Šapkarev 1978 b). Z. Balkanic-Alpine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF89B64E53DA1215FC9D9E8F.taxon	biology_ecology	E. Humid meadows (Šapakrev 1971, 1977 a). D. Golema Vevčansko Lake (Šapkarev 1971). Z. Illyric species distributed in North Macedonia, Italy, Austria, Croatia, Albania and Greece (Hackenberger Kutuzović & Hackenberger Kutuzović 2013; Szederjesi et al. 2017; Szederjesi 2023).	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF88B64F53DA14F0FAF59B51.taxon	biology_ecology	E. Meadows, pastures, deciduous (beech) forests, humid soils near river and stream banks (Šapkarev 1972 a; Szederjesi 2013) D. Jezerski prolaz (Karaman 1971); Čelopek, Jegunovce, Ljuboten, Radiovce (Šapkarev 1972 a); Murtino, Novo Selo, Bansko (Šapkarev 1973); Stanišer, Brajčino, Oteševo, Perovo (Šapkarev 1976); Dojran, Siričino (Šapkarev 1980); Ohrid Lake (Sv. Naum), Prespa Lake (Oteševo) (Szederjesi 2013). Z. Trans-Aegean.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF88B64F53DA16FAFB859C8B.taxon	biology_ecology	E. Oak and beech forests, meadows, pastures, cultivated soil, river banks, and streams (Šapkarev 1970, 1974; Szederjesi 2019). D. Crni kamen, Debar, Demir Kapija, Dolina Nikole, Markov kamen, Rabrovo, Šar Mts. (Karaman 1969, 1971); Katlanovo, Kitka, Raduša, Skoplje, Skopska Crna Gora, Taor, Vodno, Konjare, Butel (Šapkarev 1970); Bitola, Gostivar, Sv. Naum, Tetovo, Ceri Pašina, Čelopek, Dolenci, Dolno Divjaci, Duv, Gorno Divjaci, Jegunovce, Jelak, Lešnica, Kačanik, Kruševo, Ljuboten, Miletino, Popova Šapka, Skočivir, Tearce (Šapkarev 1972 c); Babuna, Barovo, Bošava, Čelovec, Demir Kapija, Drenovo, Kavadarci, Konopište, Raec, Rakle, Rosoman, Trojaci-Pletvar (Šapkarev 1974); Brajčino, Gjavato, Kriveni, Ljubojno, Oteševo, Pretor, Sirhan, Staro Selo (Kriveni) (Šapkarev 1976); Belčišta, Botun, Velgošti, Galičica, Godivje, Kališta, Labuništa, Mali Vlaj, Mešeišta, Ograždenik, Ohrid, Pesočan, Petrino, Radožda, Struga, Sv. Naum, Velmej, Vevčani (Šapkarev 1977 a); Debar, Gari, Gorenci (Šapkarev 1978 b); Bašibos, Bogdanci, Dojran, Gevgelija, Kosturino, Kožuf Mts., Miletkovo, Petrovo, Sermelin, Valandovo (Šapkarev 1980 b); Makedonska Kamenica, Pehčevo, Mačevo, Sasa, Budinarci, Negrevo, Crnik, Smojmirovo, Vladimirovo, Rusinovo, Berovo, Mitrašinci (Šapkarev 1983); Bošava, Ohrid Lake (Mršić 1991); Bogdanci, Valandovsko Polje (Čalakli) (Szederjesi 2019). Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF88B64F53DA16FAFB859C8B.taxon	materials_examined	New records. HNHM / 17493 1 ex., between Trpejca and Carina, pass of Mt. Galičica, 40 ° 57.333 ' N, 20 ° 49.099 ' E, 1595 m, subalpine pasture with Crataegus, 26.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi. HNHM / 17511 1 ex., Kjesendre, at 3.7 km on the road to Galište, 41 ° 23.046 ' N, 21 ° 50.922 ' E, 920 m, beech forest with rocks, 25.04.2019, leg. L. Dányi, Z. Fehér, Gy. Makranczy, T. Szederjesi.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF88B64C53DA121FFEE099AC.taxon	biology_ecology	E. Humid meadows, forests, floodplains (Šapkarev 1971, 1977 a). D. Studenčišta near Ohrid (Šapkarev 1971, 1977). Z. Trans-Aegean.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF88B64C53DA121FFEE099AC.taxon	discussion	Remarks. Allolobophora (s. l.) leoni has long been of uncertain taxonomic status within the genus Allolobophora. Mršić & Šapkarev (1988) classified leoni in the genus Panoniona, due to the structure of the nephridial bladders. On the other hand, All. chlorotica was found in a distant clade together with the species Allolobophora dubiosa (Örley, 1881) and Allolobophora molleri Rosa, 1889, the mentioned species forming the backbone of the genus (Domínguez et al. 2015). In the most recent phylogenetic study, Navarro et al. (2023) confirmed the validity of the genus Panoniona.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF8BB64C53DA15F5FEDB9A46.taxon	biology_ecology	E. Humid meadows (Karaman 1969). D. Miletkovo near Vardar (Ude 1922); Miletkovo (Karaman 1969, 1971). Z. Southern-Alpine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF8BB64C53DA17CAFB089DFA.taxon	biology_ecology	E. Hilly meadows, humid soil (Mršić 1991). D. Rašče (Mršić 1991). Z. Central European.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF8BB64C53DA1098FCBF9F3D.taxon	biology_ecology	E. Beech forest and humid meadows (Šapkarev 1977 a). D. Novo Selo, Radoviš (Šapkarev 1973); Besvica, Demir Kapija (Šapkarev 1974); Jablanica, Trpejca (Šapkarev 1977 a); Demir Kapija (Miravci) (Šapkarev 1980); Galičica (Šapkarev 1983); Kosturino (Mršić 1991). Z. Trans-Aegean.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF8BB64D53DA132EFBD79980.taxon	biology_ecology	E. Mud of fresh and brackish waters (Ude 1922). D. Valandovo (Ude 1922); Studenčište near Ohrid (Černosvitov 1938); Ohrid (Šapkarev 1962); Sirhan, Stenje, Perovo (Šapkarev 1976); Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
585B879BFF8AB64D53DA15D1FE8B9BCB.taxon	biology_ecology	E. Anthropogenic habitats, humid soil (Karaman 1971). D. Dojran, Bogdanci, Demir Kapija (Karaman 1971); Z. Peregrine.	en	Popović, Filip J., Trakić, Tanja B., Szederjesi, Tímea, Hackenberger, Davorka K., Sekulić, Jovana M. (2025): Checklist of earthworms of North Macedonia (Clitellata: Megadrili). Zootaxa 5589 (1): 210-234, DOI: 10.11646/zootaxa.5589.1.17, URL: https://doi.org/10.11646/zootaxa.5589.1.17
