identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5F5487DB944D762AFF0AFA6219D265E2.text	5F5487DB944D762AFF0AFA6219D265E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Muiravea voanio Bahder, Stroinski & Bartlett 2025	<div><p>Muiravea voanio Bahder, Stroiński &amp; Bartlett, sp. nov.</p><p>(Figs 2–7)</p><p>Type locality. Madagascar, Antsinanana Region, Toamasina.</p><p>Etymology. The specific name “voanio ” is Malagasy word for coconut, referencing the primary host plant (Fig. 2), and is used as a proper noun in apposition and is indeclinable.</p><p>Diagnosis. Small, pale-bodied with light fuscous markings on forewings, very narrow stridulatory plate on hindwing margin. Large subantennal shelf present. Head and thorax without fuscous markings. Gonostyli with large, bifid dorsal process and aedeagus with slender, laterally curved process arising at apex on dorsal margin</p><p>Description. Color. Base color of body pale, ivory white, slight yellow wash over head and thorax, legs slightly more yellow than main body, forewings pale, translucent, veins white, fuscous patch at midpoint of Pcu+A 1, random light fuscous spots at midpoint, light fuscous band extending from costal margin to hind margin at distal ¾ point, apices with light fuscous patches; abdomen yellow-orange, terminalia ivory white (Fig. 3).</p><p>Structure. Body length (incl. wings) males: 3.9–4.2 mm (n = 10), females: 4.2–4.5 mm (n = 10) (Table 3).</p><p>Head. Vertex without sensory pits, acarinate; in dorsal view subtriangular, concave at anterior margin, angulate, concave at posterior margin, angulate, about as wide at posterior margin as long at midpoint, lateral margins straight; in lateral view, rounded, extending slightly above and beyond eye margin. Frons without sensory pits; in frontal view, narrow dorsally, lateral margins touching at midpoint, expanding just below ventral margin of eyes, expanding to frontoclypeal suture (Fig. 4). Compound eyes with distinct ventral incision; ocelli absent. Subantennal shelf arising near posterior margin of genae, extending to just beyond midpoint of genae, ridge large and foliaceous with setae on external margin. Antennal scape very short and ringlike; second antennal segment ovoid, approximately twice as long as wide, anterior margin curved, posterior margin nearly linear (Fig. 4B). Clypeus weakly carinate at midpoint, lateral margins acarinate, rostrum extending to midcoxae, with apical segment approximately long as wide.</p><p>Thorax. Pronotum in dorsal view moderately convex at anterior margin, rounded, moderately concave at posterior margin, tricarinate, lateral carinae terminating at approximately 1/3 distance from dorsal margin on lateral margins, forming shallow foveae (Fig. 4). Mesonotum approximately wide as long; weakly tricarinate, lateral carinae gently curved mesad, extending from anterior to posterior margin, mid carina extending from anterior to posterior margin (Fig. 4); lateral angles placed after midlength. Hindlegs lacking lateral spines on tibia, metatarsal spinulation 5-5-4.</p><p>Forewing elongate, about 2.7 times longer than broad (at the level of end of clavus), membranous, broad spatulate; costal and claval margin subparallel, posterior margin strongly arcuate with breaking point at the level of MP 1.1 vein. Sensory pits present on common stem of ScP+R and MP and on basal part of stem ScP+R after forking of MP vein and on basal part of Pcu vein. Basal cell small and elongate. ScP+R+MP arising from basal cell as a common stem with first fork close to basal cell (distinctly before fusion of claval veins). Forking sequence of longitudinal veins (proximal to distal) ScP+R→CuA→MP; all before tip of clavus. Longitudinal vein branching pattern RA unbranched, RP 2-branched, MP 6-branched (MP 1+2 with 4 terminals, MP 3+4 with 2 terminals), CuA 2- branched: CuA 1 blind not reaching margin, CuA 2 reaching margin. Terminals of RA, RP and MP veins ending on posterior margin, CuA terminals ending on postclaval margin. Median area apically the largest, radial and cubital area apically occupying similar area. Cell C1 longest about 12 times longer than wide; cell C3 shorter and widest than C1, about 5.9 times longer than wide; C5 shortest about 6.8 times longer than wide. Cell length sequence C1→ C3→C5. Clavus extending to midpoint. Tegmen with single apical line of transverse crossveins; icu transverse crossvein present, reaching postclaval margin; radial and median cell each with two transverse crossveins rp-mp and mp-cua. Clavus closed (Pcu+A 1 reaching wing margin distinctly before CuP) without any transverse crossveins; claval veins Pcu and A 1 joined together about the middle of the clavus. Tegula smooth, without carinae. Hindwing hyaline, shorter than forewing (approximately ¾ the length). ScP+RA ending with single terminal about middle of costal margin; RP unbranched, MP single, CuA with 3 terminals, forking after middle of wing, CuP single, Pcu single, A 1 with 2 terminals, A 2 single, blind (not reaching margin) distinctly thickened, apically enlarged; stridulatory plate with outer margin weakly concave; crossveins r-m and m-cu present at apical third of hindwing.</p><p>Genitalia. In lateral view, pygofer narrow, strongly sinuate at anterior and posterior margin, reaching dorsal margin, broadest at ventral margin, narrowing immediately, expanding just before midpoint; in ventral view, medioventral process absent (Fig. 6). Gonostyli lateral view broadly spatulate, rounded at apex, large, bifid process on dorsal margin, distal bifurcation with apex curved ventrad, angled laterad, basal bifurcation with apex truncated, curved caudad (Fig. 6). In ventral view, narrow at base, expanding just past midpoint, forming rounded, inner projections, constricting at distal ¾ point, rounded at apex, outer margins irregularly sinuate (Fig. 6). Aedeagus simple, short, cylindrical, moderately sclerotized process arising at apex on dorsal margin (A1), slender, angled cephalad, curved to left lateral side (Fig. 7). Endosoma complex with two large expanded flange-like processes (E1 &amp; E2); E1 arising on right side, deeply trifucated, forming three hooked, dorsal processes (E1a, E1b and E1c), E1a longest and most robust, angled dorsad, curved caudad, E1b smallest process, angled and curved the same as E1a, less than ¼ the size of E1a and E1c almost as long as E1a, approximately half the width at the base, angled and curved the same as E1a and E1b, fourth process (E1d) arising on E1 along ventral margin, more sclerotized than rest of E1, curving across midline, angled to left lateral side; E2 arising on left lateral side, less sclerotized that E1 at base, becoming more sclerotized in distal half, distal half constricts, forming long, slender process, angled cephalad, curved ventrad (Fig. 7). Anal segment short, irregularly sinuate on dorsal and ventral margins, apex rounded, not extending beyond apex of gonostylus (Fig. 6).</p><p>Remarks. The novel taxon, M. voanio sp. nov. is placed within Muiravea based on the wing characters outlined by Banaszkiewicz &amp; Szwedo (2005), namely the unbranched RA and CuA not reaching the wing margin. The genus Muiravea is monotypic and was erected for the single species from Sierra Leone (West Africa) – Patara hargeavesi Muir, 1930 . The novel taxon differs significantly from M. hargreavesi in overall color with the latter being dark in color and M. voanio sp. nov. being pale. Also, M. hargreavesi differs from M. voanio sp. nov. in that the former has a blind CuA2 and the latter has a blind CuA1. The armature of the aedeagus among the two species differs significantly. One aspect of the genitalia listed as a generic feature, the presence of three processes at apex of the anal segment are absent in M. voanio sp. nov. but with all other similarities (including overall similar form of the genitalia), this does not appear enough to exclude it from the genus, thus requiring amending of the genus diagnosis so that this feature is not included. In the future, as more material becomes available for analysis and specimens of M. hargreavesi are obtained for examination, the novel taxa described herein could be placed in a new genus based on the differences observed. However, based on the material and information available, placing the novel taxa within Muiravea is the most prudent.</p><p>Plant associations. Cocos nucifera L. ( Arecaceae Bercht. &amp; J. Presl) (Table 4)</p><p>Type material. Holotype, male: “ Madagascar, Atsinanana, Toamasina / Hotel Darifify / 27.I.2023, Coll.: B.W.Bahder / Host: Cocos nucifera // Holotype / Muiravea voanio ♂ /” (FLREC).</p><p>Paratypes: 9 males, 10 females, same locality as holotype (FLREC), 1 male and 1 female in MIIZ . 2 males “ Madagascar, Atsinanana, Toamasina / Coconut plantation 10km N / 28.I.2023, Coll.: B.W.Bahder / Host: Cocos nucifera // Holotype / Muiravea voanio ♂ / “</p><p>Distribution. Central-eastern Madagascar (Atsinanana Region, Toamasina I District, Toamasina commune: 18°6’54.88”S, 49°24’16.75”E).</p></div>	https://treatment.plazi.org/id/5F5487DB944D762AFF0AFA6219D265E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bahder, Brian W.;Stroiński, Adam;Bartlett, Charles R.;Randretsiferana, Safidinirina Armande;Razafindrakotomamonjy, Andrianatenaina;Pilet, Fabian;Hasinjaka, Rasolondalao Harin’Hala	Bahder, Brian W., Stroiński, Adam, Bartlett, Charles R., Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala (2025): New derbid planthoppers in the genus Muiravea (Hemiptera: Fulgoromorpha: Derbidae) from palms in eastern Madagascar, a key to the genus and an updated molecular phylogeny of available Malagasy Otiocerinae. Zootaxa 5642 (2): 147-162, DOI: 10.11646/zootaxa.5642.2.3, URL: https://doi.org/10.11646/zootaxa.5642.2.3
5F5487DB944D7621FF0AFB021D43672D.text	5F5487DB944D7621FF0AFB021D43672D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Muiravea Banaszkiewicz & Szwedo 2005	<div><p>Genus Muiravea Banaszkiewicz &amp; Szwedo, 2005</p><p>Type species. Patara hargreavesi Muir, 1930</p><p>Amended Diagnosis (Modified from Banaszkiewicz &amp; Szwedo 2005). Tegmen with single terminal of RA, vein CuA bifurcate then anastomosing, closing anterior cubital area of one cell. At least one of the CuA veins blind, not reaching wing margin. Subantennal shelf on gena well developed, with distinct row of setae on lateral margin.</p><p>Key to species of Muiravea (adult males)</p><p>1. Body fuscous, antennae elongate/ovoid, three processes at apex of anal tube (Banaszkiewicz &amp; Szwedo 2005, fig. 10)................................................................................. M. hargreavesii (Sierra Leone)</p><p>–. Body pale/white, three processes at apex of anal segment absent (Fig. 6c)......................................... 2</p><p>2. Tegulae, lateral margins of pronotum, frons and vertex fuscous (Fig. 8), aedeagal process serpentine and angled dorsad (Fig. 11)................................................................. M. maintysorokai sp. nov. (Madagascar)</p><p>–. Preceding parts not fuscous (Fig. 3), aedeagal process slender, angled laterally (Fig. 7)..... M. voanio sp. nov. (Madagascar)</p></div>	https://treatment.plazi.org/id/5F5487DB944D7621FF0AFB021D43672D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bahder, Brian W.;Stroiński, Adam;Bartlett, Charles R.;Randretsiferana, Safidinirina Armande;Razafindrakotomamonjy, Andrianatenaina;Pilet, Fabian;Hasinjaka, Rasolondalao Harin’Hala	Bahder, Brian W., Stroiński, Adam, Bartlett, Charles R., Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala (2025): New derbid planthoppers in the genus Muiravea (Hemiptera: Fulgoromorpha: Derbidae) from palms in eastern Madagascar, a key to the genus and an updated molecular phylogeny of available Malagasy Otiocerinae. Zootaxa 5642 (2): 147-162, DOI: 10.11646/zootaxa.5642.2.3, URL: https://doi.org/10.11646/zootaxa.5642.2.3
5F5487DB9447762EFF0AFB9D1C886425.text	5F5487DB9447762EFF0AFB9D1C886425.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Muiravea maintysorokai Bahder, Stroinski & Bartlett 2025	<div><p>Muiravea maintysorokai Bahder, Stroiński &amp; Bartlett, sp. nov.</p><p>(Figs 8–11)</p><p>Type locality. Madagascar, Atsinanana Region, Analalava Forest Reserve.</p><p>Etymology. The specific name is Malagasy referencing the dark/fuscous tegulae with “soroka” meaning shoulder (tegulae) and “mainty” meaning black.</p><p>Diagnosis. Small, pale-bodied, fuscous tegulae and lateral margins of pronotum. Large sub antennal shelf present. Gonostyli with large, bifid dorsal process and aedeagus with slender, serpentine process arising at apex on dorsal margin.</p><p>Description. Color. Base color of body pale, ivory white, head with fuscous patch at dorsal margin of frons, extending to lateral margins of vertex, fuscous bands on lateral margins of pronotum and fuscous tegulae; abdomen yellow-orange, terminalia ivory white (Fig. 8).</p><p>Structure. Body length (without wings) male: 1.9 mm (n = 1) (Table 3).</p><p>Head. Vertex subtriangular in dorsal view, concave at anterior margin, angulate, concave at posterior margin, angulate, about as wide at posterior margin as long at midpoint, lateral and margins lightly sinuate; in lateral view, rounded, extending slightly above and beyond eye margin, sub antennal ridge arising at posterior margin, extending just beyond midpoint of genae. In frontal view, frons narrowed dorsally, lateral margins touching at midpoint, expanding just below ventral margin of eyes, expanding to frontoclypeal suture (Fig. 9). Subantennal shelf arising near posterior margin of genae, extending to just beyond midpoint of genae, shelf large and foliaceous, with setae on external margin.</p><p>Thorax. Pronotum in dorsal view moderately convex at anterior margin, rounded, moderately concave at posterior margin, tricarinate, lateral carinae terminating at approximately 1/3 distance from dorsal margin on lateral margins, forming shallow foveae (Fig. 9). Mesonotum approximately wide as long; weakly tricarinate, lateral carinae gently curved mesad, extending from anterior to posterior margin, mid carina extending from anterior to posterior margin (Fig. 4). Hindlegs lacking lateral spines on tibia, metatarsal spinulation 5-5-4.</p><p>Genitalia. In lateral view, pygofer narrow, strongly sinuate at anterior and posterior margin, reaching dorsal margin, broadest at ventral margin, narrowing immediately, expanding just before midpoint; in ventral view, medioventral process absent (Fig. 10). Gonostyli in lateral view broadly spatulate, rounded at apex, large, bifid process on dorsal margin, distal bifurcation with apex curved ventrad, angled laterad, basal bifurcation with apex truncated, curved caudad (Fig. 10). In ventral view, narrow at base, expanding just past midpoint, forming rounded, inner projections, constricting at distal ¾ point, rounded at apex, outer margins irregularly sinuate (Fig. 10). Aedeagus simple, short, cylindrical, moderately sclerotized process arising at apex on dorsal margin (A1), slender, angled cephalad, dorsad, snake-like in appearance (Fig. 11). Endosoma complex with two large expanded flange-like processes (E1 &amp; E2); E1 arising on right side, deeply trifucated, forming three processes (E1a, E1b and E1c), E1a longest and most robust, angled dorsad, curved caudad, apex serrated, E1b smallest process, angled dorsad, knob-like, and E1c same orientation at E1a, approximately half the size, apex pointed; E2 arising on left lateral side, less sclerotized that E1 at base, becoming more sclerotized in distal half, distal half constricts, forming long, slender process (E2a), angled cephalad, curved ventrad (Fig. 11) and an elongate lobe projecting off anterior margin (E2b). Anal segment short, irregularly sinuate on dorsal and ventral margins, apex rounded, not extending beyond apex of gonostylus (Fig. 10).</p><p>Remarks. The novel taxon, M. maintysorokai sp. nov. is placed within Muiravea based on similarity in general body form and genitalia features to M. voanio sp. nov. Unfortunately, the wings of M. maintysorokai sp. nov. were damaged in the field and could not be used for generic placement or comparison to M. voanio sp. nov. However, the similar genitalia and the strong molecular support (see below), aid in placing this taxon despite lacking wings.</p><p>Plant associations. Dypsis sp. ( Arecaceae Bercht. &amp; J. Presl) (Table 4)</p><p>Type material. Holotype, male: “ Madagascar, Atsinanana, Mahavelona / Analalava Forest Reserve / 28.I.2023, Coll.: B.W.Bahder / Host: Dypsis sp. // Holotype / Muiravea maintysorokai ♂ /” (FLREC).</p><p>Distribution. Central-eastern Madagascar (Atsinanana Region, Toamasina II District, Mahavelona (Foulpointe) commune, Analalava Forest Reserve).</p><p>Sequence data and analysis. Sequence data was generated for the three loci selected (18S rRNA, D9-D10 expansion region of 28S rRNA and 5’ region of COI) for both M. maintysorokai sp. nov. and M. voanio sp. nov. Accession numbers for both taxa and respective loci are presented in Table 2. Both novel taxa resolved adjacent to each other in all independent analyses for respective markers with strong bootstrap support (99, 100 and 85 respectively for 18S, 28S and COI (Fig. 12). The difference between the two taxa for 18S and 28S was approximately 0.1% and 10.6% different for COI. The consensus tree based on concatenate data for all three loci demonstrated strong bootstrap support (100) for the novel taxa resolving adjacent to each other and strong bootstrap support (98) for Muiravea (based on the taxa analyzed) resolving adjacent to Patara Westwood (Fig. 13).</p></div>	https://treatment.plazi.org/id/5F5487DB9447762EFF0AFB9D1C886425	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bahder, Brian W.;Stroiński, Adam;Bartlett, Charles R.;Randretsiferana, Safidinirina Armande;Razafindrakotomamonjy, Andrianatenaina;Pilet, Fabian;Hasinjaka, Rasolondalao Harin’Hala	Bahder, Brian W., Stroiński, Adam, Bartlett, Charles R., Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala (2025): New derbid planthoppers in the genus Muiravea (Hemiptera: Fulgoromorpha: Derbidae) from palms in eastern Madagascar, a key to the genus and an updated molecular phylogeny of available Malagasy Otiocerinae. Zootaxa 5642 (2): 147-162, DOI: 10.11646/zootaxa.5642.2.3, URL: https://doi.org/10.11646/zootaxa.5642.2.3
