taxonID	type	description	language	source
5D1FB5691A0234567AB9859F8C39FCDD.taxon	discussion	Comment. Burckhardt et al. (2021) erected a new subfamily, Amorphicolinae, for the genus Amorphicola based on the clade grouping and phylogenetic topology of the mitogenome analyses of Percy et al. (2018). The new genus described here was also included in the phylogenetic analyses of Percy et al. (2018), but the genus was not described at the time of the revised classification and the definition of subfamily Amorphicolinae was based solely on Amorphicola. Here, the subfamily definition is broadened to reflect characteristics of both genera in Amorphicolinae, as follows: Adult. Amorphicola and Danieliana share head characteristics as described in Burckhardt et al. (2021) except the antenna length which is 1 – 1.5 x head width in Amorphicola and longer, 1.5 – 1.6 x head width, in Danieliana; in addition, there are differences in the relative lengths of the antennal segments which are either with segment 3 longer than segments 7 or 8 (in Amorphicola) or segment 3 shorter than segment 7, and segments 4 – 6 and 8 subequal (in Danieliana). Thorax and leg characters are as described in Burckhardt et al. (2021), except metatibia is either without (in Amorphicola) or with a small genual spine (in Danieliana), and metatibia apex either bearing 4 irregularly spaced, sclerotised, apical spurs (in Amorphicola) or 5 grouped (1 + 3 + 1) sclerotised apical spurs (in Danieliana). Forewing and hindwing as described in Burckhardt et al. (2021). Male proctiger as described in Burckhardt et al. (2021); paramere complex, in profile either axe or hammer-shaped with several sclerotised peg setae on the inner face (in Amorphicola) or long and sinuous with many long, extended setae on the inner face (in Danieliana). Fifth instar immature. As described in Burckhardt et al. (2021).	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A0234567AB9859F8C39FCDD.taxon	description	Systematics. The majority of adult and all immature characteristics for Amorphicolinae given in Burckhardt et al. (2021) are shared by Amorphicola and Danieliana. The main shared characteristics are in the general structure of the adult head, thorax, legs, male proctiger, and elongate male subgenital plate, and in the immature structure and chaetotaxy. The main differences between the genera being antenna length, relative lengths of antennal segments, size of forewing cells cu 1 and m 1, presence / absence of a genual spine on metatibia and number of sclerotised apical metatibial spurs, and the shape of the paramere. Some of the morphological characters of Danieliana that differentiate it from Amorphicola, suggest an affiliation with Ciriacreminae Enderlein, 1910 (see comment under generic description below), particularly the genus Isogonoceraia Tuthill, 1964, which includes two species in South America (White & Hodkinson 1980; Burckhardt & Queiroz 2012) and one in Micronesia (Tuthill 1964). Isogonoceraia is also the only member of Ciriacreminae known to have caesalpinoid legume (referring to Caesalpinieae and Cassieae clades of Fabaceae) host plants (Ouvrard 2022), which is the same host plant group as Danieliana. Characteristics shared between some members of Amorphicolinae and Ciriacreminae may reflect shared ancestry (e. g., in the root of subfamilies Amorphicolinae, Ciriacreminae and Psyllinae Latreille, 1807), as the two subfamilies are phylogenetically close, or they may be homoplasious and converged in the two groups.	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A0134567AB985368C38F9B9.taxon	materials_examined	Type genus: Amorphicola Heslop-Harrison, 1961 Morphological description as given in Burckhardt et al. (2021). Included genus: Amorphicola Heslop-Harrison, 1961 Material examined. Amorphicola amorphae, 7 males, 6 females, 5 immatures, White Oak Conservation Park, ~ 30 km north of Jacksonville, ex Amorpha fruticosa, 27 May 2005, coll. FL 17 - 05, DM Percy leg. (DMPC).	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A0134567AB986FA8B1BF891.taxon	materials_examined	Type genus: Danieliana gen. nov. Included genus: Danieliana gen. nov.	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A0134507AB987128C66FD99.taxon	materials_examined	Type species: Danieliana manmiaoyangae sp. nov., by present designation. Adult structure. Mid-sized psyllid. Head, in profile, moderately inclined at 30 – 45 ° from longitudinal body axis (Figs. 1 H, 1 M). Thorax moderately arched (Fig. 1 M). Vertex longer than genae, separated from genae by transverse suture; genae forming diverging conical processes contiguous at base; coronal suture fully developed. Antenna 10 - segmented, filiform, equal to or more than 1.5 times as long as head width, segment 3 shorter than segment 7, segments 4 – 6 and 8 subequal (Figs. 1 I, 1 J), a single subapical rhinarium on each of segments 4, 6, 8, and 9; terminal seta longer than segment 10 (Fig. 1 G). Clypeus slightly flattened ventrally, hardly visible in lateral view, distal proboscis segment short, distinctly exceeding procoxae (Figs. 1 F 1 H). Thorax moderately arched (Figs. 1 H, 1 M), width subequal to head width; pronotum ribbon-shaped; mesopraescutum in longitudinal body axis about as long as mesoscutum; propleurites with subequal epimeron and episternum. Legs moderately long (Fig. 1 M); metacoxa with moderately large, thorn-shaped meracanthus, length of metafemur and metatibia subequal, base of metatibia with short genual spine, apex bearing 5 (1 + 3 + 1) sclerotised apical spurs, basal metatarsus with 2 apical spurs and slightly shorter than apical metatarsus (Fig. 1 K). Forewing widest in apical half, broadly rounded apically (Figs. 1 A, 1 C); membrane semitransparent, covered in densely spaced surface spinules in all cells, but with mostly spinule free areas bordering veins (Fig. 1 L); three typical radular spine clusters, one each at apical margin of cells cu 1, m 2 and m 1, and a fourth less prominent cluster at the apical margin of cell r 2 (Fig. 1 L); costal break developed; pterostigma broad but short; vein R longer than M + Cu; anal break close to apex of vein Cu 1 b. Hindwing ~ 0.8 times length of forewing, membranous, costal setae grouped, vein M + Cu developed. Male proctiger one segmented, more or less parallel sided (Fig. 2 A); male subgenital plate elongate, about twice as long as high (Fig. 2 A); paramere in lateral view simple, long, and slender (Figs. 2 B, 2 C), with long simple setae on the interior and posterior surfaces, and short simple setae on the exterior surface (Fig. 2 C). Female terminalia short (Fig. 2 K); proctiger apex acute in dorsal view (Fig. 2 G), anal ring narrowly oval and composed of a double row of cells (Fig. 2 J); subgenital plate shorter than proctiger, apex blunt in ventral view (Fig. 2 H); ovipositor small with valvulae dorsalis low (Fig. 2 I). Immature structure. Body elongate-oval in dorsal view with wing pads protruding (Fig. 3 A). Antenna 7 - segmented, with last segment showing numerous partial divisions, longest segment 7 th, shortest segment 4 th, single subapical rhinarium on 3 rd and 5 th segments, and two on the basal half of 7 th segment, with small scattered simple setae and two long stout setae apical and subapical on terminal segment (Figs. 3 C, 3 D). Head and thorax with scattered long and shorter capitate setae and very short rod-like setae, and a single ocular capitate seta (Fig. 3 B). Forewing pads lacking humeral lobes, both forewing and hindwing pads bearing long marginal capitate setae, and shorter capitate setae on the surface (Fig. 3 H). Legs bearing capitate and simple setae (Fig. 3 I), tarsal arolium longer than claws, fan-shaped with petiole and unguitractor, claws well developed (Fig. 3 E). Caudal plate developed, semi-circular and rounded apically, with 4 + 4 marginal sectasetae and long capitate setae (Fig. 3 F). Anus in ventral position, circumanal ring heart-shaped, consisting of a single row of pores (Fig. 3 G).	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A0134507AB987128C66FD99.taxon	biology_ecology	Biology. Immatures are free living on young foliage.	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A0134507AB987128C66FD99.taxon	discussion	Comment. Previously, Amorphicola, with two species, was the only genus in subfamily Amorphicolinae. Danieliana gen. nov. is sufficiently divergent from the North American Amorphicola based on both morphological and molecular data for these genera to be recognized as separate tribes within subfamily Amorphicolinae. Danieliana gen. nov. is similar to Amorphicola in general body and wing structure as given in Burckhardt et al. (2021). It differs from Amorphicola in having broader forewings with higher cells cu 1 and m 1, head with longer, upturned genae, longer antenna with a short 3 rd segment (shorter than 7 th and subequal to 4 th, 5 th, 6 th and 8 th), metatibia with a genual spine and 5 versus 4 apical spurs, paramere long and slender. Some of these differences suggest Danieliana gen. nov. shares traits characteristic of the related subfamily, Ciriacreminae (Burckhardt et al. 2021). Ciriacreminae includes 19 genera most of which occur on Fabaceae hosts (Ouvrard 2022). In particular, there is some resemblance to Isogonoceraia Tuthill, 1964, particularly Isogonoceraia venusta Tuthill, 1964 with which Danieliana shares the broad forewing, longer antenna with near equal length of most flagellar sections (Fig. 1 J), large pterostigma (wide but short), presence of small genual spine, short female genitalia and somewhat in the shape of the female proctiger. It does not share the elongate vertex, contiguous genae, and peculiar shape of the aedeagus apex. Isogonoceraia includes two described species, with one species in Brazil and one species in the Mariana Islands (Micronesia). Isogonoceraia is also the only member of Ciriacreminae with caesalpinoid legume hosts, a trait also shared with Danieliana Systematics. Danieliana gen. nov. was included, as an undescribed genus, in the mitogenome analysis of Percy et al. (2018) (referred to as “ Genus unnamed (Taiwan) ” in Table 3) where it was found to group strongly (97 %) in Group U with Amorphicola Heslop-Harrison, 1961. Amorphicola includes two species, both distributed in North America on hosts in the legume genus Amorpha (Fabaceae) (Tuthill 1943; Halbert & Burckhardt 2020; Ouvrard 2022). Group U was recognized as a new subfamily, Amorphicolinae, in Burckhardt et al. (2021) to accommodate Amorphicola, which is considered well characterised by its paramere morphology and by the Amorpha (Fabaceae) host associations. The phylogenetic placement and strong support grouping Danieliana gen. nov. with Amorphicola in Percy et al. (2018) implies it was tacitly assumed included in subfamily Amorphicolinae. The morphological differences between the two genera are also outlined in the revised subfamily definition (above) to accommodate Danieliana in subfamily Amorphicolinae.	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A0134507AB987128C66FD99.taxon	etymology	Etymology. The genus name honours Daniel Burckhardt for his prodigious contribution to our knowledge of psyllids. Note on species description. Danieliana gen. nov. is described as a monotypic genus. The species description below provides details of species-specific characteristics not supplied in the generic description above.	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A07345D7AB9821A88E6FEE4.taxon	description	(Figs. 1, 2, 3, 4) Adult colour and structure. General body colour bright green, to yellow-green (Figs. 4 A, 4 C); forewing infused orange-yellow, in some individuals fuscous margins to apical cells. Antenna yellow-green with darker apices on segments 6 – 8, and segments 9 – 10 dark. Head and thorax structure as given in generic description, both with scattered short simple setae and longer simple setae on genae, surfaces rugose. Antenna structure as in generic description, with scattered simple setae. Forewing with vein Rs slightly to moderately sinuate (Figs. 1 A, 1 C). Male proctiger in lateral view, parallel sided with constriction only below apex (Fig. 2 A). Male subgenital plate elongate, about twice as long as high, broader basally and tapering apically, in posterior view slightly vase-shaped, well rounded ventrally (Fig. 2 E). Paramere in lateral view simple, long, slender and slightly sinuate (Figs. 2 B, 2 C), in dorsal view with interiorly directed ridge (Figs. 2 E, 2 F), with long simple setae on the interior and posterior surfaces, and short simple setae on the exterior surface, and one particularly long seta directed anteriorly from the apex (Fig. 2 C). Proximal aedeagus segment inflated distally (Fig. 2 A, 2 D), distal aedeagus segment with bulbous, rounded apex bearing a posteriorly directed flange (Fig. 2 D). Female terminalia short (Fig. 2 K), in lateral view proctiger post anal ring concave, descending steeply to an upturned apex bearing long setae dorsally (Fig. 2 L), apex acute in dorsal view (Fig. 2 G), anal ring narrowly oval and composed of a double row of pores (Fig. 2 J); ovipositor valvulae dorsalis small and low, with a single blunt serration on valvulae ventralis (Fig. 2 I); subgenital plate short, apex blunt in ventral view (Fig. 2 H). Adult measurements (mm) and ratios. (7 males, 3 females). Males 2.7 – 3, females 3 – 3.4 (measured from apex of genae to wing apex). WL: 2.01 – 2.56; WW: 0.89 – 1.14; HW: 0.73 – 0.78; AL: 1.11 – 1.20; PB: 0.09 – 0.11; WL: WW: 2.12 – 2.34; WL: RsL: 1.63 – 1.72; CUR: 1.18 – 1.42; MR: 0.46 – 0.55; HM: HCU: 1.88 – 2.16; HW: VW: 1.63 – 1.76; VL: VW: 0.51 – 0.57; AL: HW: 1.49 – 1.59; HW: HT: 1.63 – 1.74; HT: HF: 0.98 – 1.07. Male terminalia: MP: 0.51 – 0.53; PL: 0.38 – 0.39; AEL: 0.24 – 0.26; PL: HW: 0.51 – 0.52; MP: PL: 1.31 – 1.37; PL: AEL: 1.50 – 1.63; AEL: AELH: 2.40 – 3.00; PL: SH: 1.63 – 1.90. Female terminalia: FP: 0.53 – 0.56; FSP: 0.32 – 0.33; RL: 0.21 – 0.23; OVH: 0.02 – 0.03; EL: 0.28; EW: 0.12; FP: RL: 2.43 – 2.52; FP: HW: 0.71 – 0.72; FP: SP: 1.7 – 1.75; EL: EW: 2.33. Immature colour and structure. 5 th instar yellow, green, or blue-green, typically with dark brown to black sclerites, caudal plate, wing pads, and notably, antennal segments 5 th and 7 th (Fig. 4 B). Younger instars usually yellow or orange (Fig. 4 A), some with darker sclerites. 5 th instar structure and chaetotaxy as in generic description. Immature measurements (mm) and ratios. 5 th instar (n = 5): BL 1.69 – 1.99; BW 1.12 – 1.34; WPL 0.55 – 0.59; CPL 0.42 – 0.45; CPW 0.77 – 0.87; RW 0.20 – 0.21; HW 0.65 – 0.73; AL 0.89 – 0.93; BL: BW 1.31 – 1.60; HW: AL 0.73 – 0.80; CPW: RW 3.85 – 4.35. Egg colour and structure. Orange-yellow (Fig. 4 D), broadly ovoid, smooth, with no apparent surface sculpturing or patterning, with a short sub-basal pedicel on the underside, and a long tail (Fig. 2 M).	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A07345D7AB9821A88E6FEE4.taxon	biology_ecology	Host plant. Caesalpinia crista (Fabaceae). Biology. Immatures free-living, and the species is possibly multivoltine (see Discussion). Note. Despite Caesalpinia crista (Fabaceae) being widespread and not uncommon in SE Asia, Australasia and South Asia, Danieliana manmiaoyangae is the first psyllid to be described from this plant.	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A07345D7AB9821A88E6FEE4.taxon	discussion	Comment. This is the only species currently known for the genus. Systematics. In the mitogenome analysis of Percy et al. (2018), this species is recovered in Group U (subfamily Amorphicolinae) as a strongly supported clade (97 %) together with Amorphicola amorphae. Genetic resources. Genbank: MG 988747 (DPTAI 79) for cytochrome oxidase 1 (COI), and MG 989051 (DPTAI 7910), OR 886079 (DPTAI 79) for cytochrome b (cytB).	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A07345D7AB9821A88E6FEE4.taxon	distribution	Distribution. Only known from one area (Pingtung) in southern Taiwan.	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A07345D7AB9821A88E6FEE4.taxon	materials_examined	Type material. Holotype male (slide mounted), Pingtung, Taiwan, N 21.95014, E 120.82314, 189 m, ex Caesalpinia crista, 30 th January 2010, coll. DPTAI 79 - 10, DM Percy leg. (NHMUK). Paratypes: 20 males (6 slide mounted, 14 in ethanol), 12 females (3 slide mounted, 9 in ethanol), 16 immatures (10 slide mounted, 6 in ethanol), as for holotype (NHMUK, DMPC). Other material examined. Additional material collected in 2023 by Yi-Chang Liao is the subject of the photographs in Fig. 4, collection details as follows: 10 males, 11 females, 25 immatures; Pingtung Co., Hengchun; 21 ° 57 ’ 11.9 ” N, 120 ° 49 ’ 26.7 ” E; 19 Jan. 2023; YC Liao leg.; Caesalpinia crista; NCHU, dry mounted or stored in ethanol.	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
5D1FB5691A07345D7AB9821A88E6FEE4.taxon	etymology	Etymology. The species name honours Man-Miao Yang in a tribute to her extraordinary contribution to our knowledge of psyllids.	en	Percy, Diana M. (2025): A new psyllid genus from Taiwan, Danieliana gen. nov. (Hemiptera: Psylloidea: Psyllidae). Zootaxa 5727 (1): 101-112, DOI: 10.11646/zootaxa.5727.1.7, URL: https://doi.org/10.11646/zootaxa.5727.1.7
