taxonID	type	description	language	source
003017B3D2DB5128B4F4C91AFDB452A6.taxon	description	Figs 7, 10	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
003017B3D2DB5128B4F4C91AFDB452A6.taxon	etymology	Etymology. This species name is derived from the Latin elongatus (prolonged), in reference to the very long, narrow hypandrium.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
003017B3D2DB5128B4F4C91AFDB452A6.taxon	diagnosis	Diagnosis. This species is distinguished by one stout anterior subapical seta on the fore femur and males of this species are distinguished by their very long hypandrium (1.1 mm) (Figs 7, 10) and lacking stout posteroventral setae on the apical fourth of the fore femur.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
003017B3D2DB5128B4F4C91AFDB452A6.taxon	description	Description. Wing length (n = 1): 4.6 mm. Male. Bright blue pruinescence laterally, with dark brown vertex of head, scutum and dorsum of abdomen. Head rounded; compound eyes large, rounded. Face convex, narrowest at middle, ca. 1.5 × width of antennal sockets, entirely clothed in bright whitish blue pruinescence; lower margin of face with slight cleft with small carina or swelling; gena one-fourth as wide as height of eye. Vertex and ocellar triangle dark brown, lower postocciput and gena lightly pruinescent. Ocellar setae longest among cranial setae; occiput with 1 pair of setae posterior to ocellar triangle; above ‘ neck’ row or crown of supracervical setae, setae in middle longest, decreasing in length laterally; postocular with row of short setae along margin of eye, upper 6 setae dark, stouter than lower setae. Arista stout, shorter than width of eye; slightly tapered apically. Palpus cylindrical, length less than one-third height of eye; labellum with ventral triangular projection. Pleura and lateral margin of scutum, including postpronotal lobe, extending toward 1 st dc and notopleuron with bright blue pruinescence (viewed laterally); scutellum, postalar ridge and region medial to postalar light brown. Thoracic setae shorter than width of eye; several acr anterior to 1 st dc; 5 pairs dc; 1 slender short pprn, one-half length of dc; 1 slender presut spal; 2 npl; 1 psut spal; 1 pal; 1 pair sctl and several pale marginal setulae; several pale setulae on postpronotal lobe and notopleural depression. Laterotergite with patch of pale setae; prothoracic episternum, posteromedial margin of katepisternum and hind margin of metepisternum with long pale setulae. Wing narrow, infuscate; narrow, faint, elliptical pterostigma present. One basal costal seta, two-thirds length of posterior dc. Row of setulae from base of R to just beyond origin of Rs, basal setulae longer than apical setulae, subequal to basal costal seta; cell dm produced apically; R 4 + 5 curved; R 4 and R 5 divergent throughout. Halter knob dark with base of shaft orange-brown. Legs dark brown, except coxae with blue pruinescence concolourous with pleura. Coxae with long pale marginal setae; anterior face of fore coxae densely clothed in long, pale setae, outer basal not longer than other setae. Fore femur with 1 stout, black anterior subapical seta, shorter than width of femur; anteroventral margin with row of 4 stout, black setae on apical third and row of 6 setae on basal half, separated by wide median gap, basal setae ca. one-sixth width of femur; basal half with posteroventral row of ca. 8 long golden setae, longer than width of femur, apical half with several shorter pale setae, less than width of femur, lacking stout, black setae. Fore tibia with biserial row of short setae beneath, ca. one-third to one-half width of tibia. Foreleg tarsomeres lacking white ventral pubescence. Mid femur with long, erect anterior setae, slightly longer than width of femur. Mid and hind femora with 2 – 4 pairs of slender antero- and posteroventral preapical setae; antero- and posterodorsal bristles absent. Hind tibia generally lacking setae except for several erect, stout dorsal and ventral setae on apical fourth. Tarsi much shorter than corresponding tibiae. Empodium pulvilliform, slightly shorter than claw, more than one-half length of tarsomere. Hypandrium long and narrow, 1.1 mm, more than 3 × longer than epandrium (Figs 7, 10); anterior margin rounded. Phallus not extending beyond clasping cercus; gently curved, apex with small membranous sac; ejaculatory apodeme small relative to hypandrium, arched anteriorly. Surstylus small, much shorter than clasping cercus with pair of 4 teeth-like projections on anterior margin. Clasping cercus finger-like, gradually tapered to rounded apex; anteroventral margin with several setae, longer than width of cercus; inner face with short peg-like setae; posterodorsal margin with fine setulae. Female. Unknown.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
003017B3D2DB5128B4F4C91AFDB452A6.taxon	distribution	Distribution. This species is known only from Yunnan Province of China (Fig. 14).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
4B0C8F3B6ADF5B9C80552D8BDCD3D5EC.taxon	description	Figs 8, 11	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
4B0C8F3B6ADF5B9C80552D8BDCD3D5EC.taxon	etymology	Etymology. This species is named after its similarity to T. elongata sp. nov.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
4B0C8F3B6ADF5B9C80552D8BDCD3D5EC.taxon	diagnosis	Diagnosis. This species is distinguished by two stout anterior subapical setae on the fore femur and males of this species are distinguished by their very long hypandrium (0.75 mm) (Figs 8, 11) and presence of stout posteroventral setae on apical fourth of fore femur.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
4B0C8F3B6ADF5B9C80552D8BDCD3D5EC.taxon	description	Description. Wing length (n = 4): 3.9 – 4.6 mm. Male. Pale blue pruinescence laterally, with dark brown vertex of head, scutum and dorsum of abdomen. Head rounded; compound eyes large, rounded. Face convex, narrowest at middle, ca. 1.5 × width of antennal sockets, entirely clothed in bright whitish blue pruinescence; lower margin of face with slight cleft with small carina or swelling; gena one-fourth as wide as height of eye. Vertex and ocellar triangle dark brown, lower postocciput and gena lightly pruinescent. Ocellar setae longest among cranial setae; occiput with 1 pair of setae posterior to ocellar triangle; above ‘ neck’ row or crown of supracervical setae, setae in middle longest, decreasing in length laterally; postocular with row of short setae along margin of eye, upper 6 setae dark, stouter than lower setae. Arista stout, shorter than width of eye; slightly tapered apically. Palpus cylindrical, length less than one-third height of eye; labellum with ventral triangular projection. Pleura and lateral margin of scutum, including postpronotal lobe, extending toward 1 st dc and notopleuron with pale blue pruinescence (viewed laterally); scutellum, postalar ridge and region medial to postalar light brown. Thoracic setae shorter than width of eye; several acr anterior to 1 st dc; 5 pairs dc; 1 slender short pprn, one-half length of dc; 1 slender presut spal; 2 npl; 2 psut spal; 1 pal; 1 pair sctl and several pale marginal setulae; several pale setulae on postpronotal lobe and notopleural depression. Laterotergite with patch of pale setae; prothoracic episternum, posteromedial margin of katepisternum and hind margin of metepisternum with long pale setulae. Wing narrow, infuscate; narrow, faint, elliptical pterostigma present. One basal costal seta, two-thirds length of posterior dc. Row of setulae from base of R to just beyond origin of Rs, basal setulae not longer than apical setulae; cell dm produced apically; R 4 + 5 straight; R 4 and R 5 divergent throughout. Halter knob dark with base of shaft orange-brown. Legs dark brown, except coxae with blue pruinescence concolourous with pleura. Coxae with long pale marginal setae; anterior face of fore coxae with pale setae, outer basal not longer than other setae. Fore femur with 2 stout, black anterior subapical setae, shorter than width of femur; anteroventral margin with row of 4 stout, black setae on apical third and row of 6 setae on basal half, separated by wide median gap; ca. one-fourth width of femur; posteroventral margin with row of setae, 3 setae on apical fourth similar to anteroventral setae, remaining setae slender, increasing in length basally to ca. two-thirds width of femur. Fore tibia with biserial row of short setae beneath, ca. one-third to one-half width of tibia. Foreleg tarsomeres lacking white ventral pubescence. Mid femur with long, erect anterior setae, slightly longer than width of femur. Mid and hind femora with 2 – 4 pairs of slender antero- and posteroventral preapical setae; antero- and posterodorsal bristles absent. Hind tibia generally lacking bristles except for several erect, stout dorsal and ventral setae on apical fourth. Tarsi much shorter than corresponding tibiae. Empodium pulvilliform, slightly shorter than claw, more than one-half length of tarsomere. Hypandrium long and narrow, 0.75 mm, more than 2 × longer than epandrium (Figs 8, 11); anterior margin with small apodeme. Phallus not extending beyond clasping cercus; sinuous, apex with small membranous sac; ejaculatory apodeme small relative to hypandrium, slightly arched anteriorly. Surstylus small, much shorter than clasping cercus with pair of 4 teeth-like projections on anterior margin. Clasping cercus finger-like, gradually tapered to rounded apex; anteroventral margin with several setae, longer than width of cercus; inner face with short peg-like setae; posterodorsal margin with fine setulae. Female. Similar to male except as follows: chaetotaxy of fore femur very similar, except posteroventral setae reduced; fore tibia lacking modified setae beneath. Terminalia: undissected.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
4B0C8F3B6ADF5B9C80552D8BDCD3D5EC.taxon	distribution	Distribution. This species is known only from eastern Nepal (Fig. 14).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
26158750F2B9520CA587E5B7514544BF.taxon	description	Figs 1 – 4, 5 – 6, 9, 12 – 13, 14	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
26158750F2B9520CA587E5B7514544BF.taxon	etymology	Etymology. The species name is derived from the Greek systenos (tapering to a point) and Greek gnathos (jaw), in reference to the pointed labellum of this species, which also characterizes this group of species.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
26158750F2B9520CA587E5B7514544BF.taxon	diagnosis	Diagnosis. This species is distinguished from the other two species of this species subgroup by one (rarely two) stout anterior subapical seta (e) on the fore femur (Figs 3, 4) and males with a much shortened and stouter hypandrium (Figs 9, 12) and lacking stout posteroventral setae on the apical fourth of the fore femur (Fig. 3).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
26158750F2B9520CA587E5B7514544BF.taxon	description	Description. Wing length (n = 5): 4.7 – 5.3 mm. Male. Bright blue pruinescence laterally, with dark brown vertex of head and scutum (Fig. 5). Head rounded; compound eyes large, rounded (Figs 1, 2). Face convex, narrowest at middle, ca. 1.3 × width of antennal sockets, entirely clothed in bright whitish blue pruinescence; lower margin of face with slight cleft with small carina or swelling; gena one-fourth as wide as height of eye. Vertex and ocellar triangle dark brown, lower postocciput and gena lightly pruinescent. Ocellar setae longest among cranial setae; occiput with pair of setae posterior to ocellar triangle; above ‘ neck’ row or crown of supracervical setae, setae in middle longest, decreasing in length laterally; postocular with row of short setae along margin of eye, upper 4 – 6 setae dark, stouter than lower setae. Arista stout, shorter than width of eye; slightly tapered apically. Palpus cylindrical, length less than one-third height of eye; labellum with ventral triangular projection. Pleura and lateral margin of scutum, including postpronotal lobe, extending medially toward first dc and notopleuron with bright blue pruinescence; scutellum, postalar ridge and region medial to postalar light brown. Thoracic setae shorter than width of eye; several acr anterior to first dc; 5 pairs dc; 1 slender, short pprn, one-half length of dc; 1 slender presut spal; 2 npl; 2 psut spal; 1 pal; 1 pair sctl and several pale marginal setulae; several pale setulae on postpronotal lobe and notopleural depression. Laterotergite with patch of pale setae; proepisternum, posteromedial margin of katepisternum and hind margin of metepisternum with long pale setulae. Wing (Fig. 6) narrow, infuscate; narrow, faint, elliptical pterostigma present. One basal costal seta, two-thirds length of posterior dc. Row of setulae from base of R to just beyond origin of Rs; cell dm produced apically; R 4 + 5 slightly curved; R 4 and R 5 divergent throughout. Halter knob dark with base of shaft orange-brown. Legs dark greyish brown, except coxae with blue pruinescence concolourous with pleura. Coxae with long pale marginal setae; anterior face of fore coxae with scattered pale setae, outer basal not longer than other setae. Fore femur with 1 (rarely 2) stout, black anterior subapical seta (e), shorter than width of femur; anteroventral margin with row of very short, stout, black setae, increasing in length apically, ca. one-sixth width of femur (Fig. 3); basal half with posteroventral row of slender, pale setae, slightly shorter than width of femur, without stout posteroventral setae on the apical fourth. Fore tibia with biserial row of short, erect setae beneath, ca. one-half width of tibia. Foreleg tarsomeres lacking white ventral pubescence. Mid and hind femora with row of long, erect anterior setae, slightly longer than width of femur. Mid and hind femora with 2 – 4 pairs of slender anteroventral and posteroventral preapical setae; anterodorsal and posterodorsal setae absent. Hind tibia generally lacking setae except for several erect, stout dorsal and ventral setae on apical fourth. Tarsi much shorter than corresponding tibiae; tarsomere 1 of all legs with long ventral seta at base. Empodium pulvilliform, slightly shorter than claw, more than one-half length of tarsomere. Hypandrium broad, quadrate, distinctly longer than epandrium (less than 1.3 times); anterior margin with or without long slender apodeme (Fig. 9). Phallus extending beyond clasping cercus; gradually arched throughout; apical portion flattened in dry specimens; apex with flexible anterior sclerite, occasionally with long narrow membranous sac and longitudinal groove laterally (maybe expansion of inner sac due to clearing); ejaculatory apodeme large, arched strongly anteriorly. Surstylus small, much shorter than clasping cercus with pair of knob-like projections on anterior margin bearing setulae. Clasping cercus finger-like, gradually tapered to rounded apex; anteroventral margin with several setae, longer than width of cercus; inner face with short, peg-like setae especially on basal half. Female. Similar to male except as follows: face blue pruinescent, brownish medially; chaetotaxy of fore femur reduced, with setae of anteroventral row shorter and less stout (Fig. 4); posteroventral setae slenderer, one-fifth width of femur; fore tibia lacking modified setae beneath. Terminalia (Fig. 13): sclerites closely approximated laterally; T 8 with posterior row of long setae, dorsal setae longer than length of sclerite in dorsal view; T 9 + 10 with curved setae on apical two-thirds, with 5 apical and several inner setae more stout than remaining setae; anteroventral corner of T 9 + 10 subtriangular, prolonged obliquely.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
26158750F2B9520CA587E5B7514544BF.taxon	distribution	Distribution. This species is known from eastern Bhutan, Nepal, Yunnan Province (China) and Vietnam (Fig. 14).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2025): Revision of the Trichoclinocera systenognatha species subgroup from eastern Asia (Diptera, Empididae, Clinocerinae). Evolutionary Systematics 9 (1): 39-47, DOI: 10.3897/evolsyst.9.145184
