identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
7C3D87FBB617902C71E4FC76C3A1FDA2.text	7C3D87FBB617902C71E4FC76C3A1FDA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chironomidae NEWMAN 1834	<div><p>FAMILY:  CHIRONOMIDAE NEWMAN, 1834</p><p>SUBFAMILY:  CHIRONOMINAE NEWMAN, 1834</p><p>TRIBE:  TANYTARSINI ZAVŘEL, 1917</p><p>SUBTRIBE:  TANYTARSINA ZAVŘEL, 1917</p></div>	https://treatment.plazi.org/id/7C3D87FBB617902C71E4FC76C3A1FDA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zakrzewska, Marta;Singh, Hukam;Wagner-Wysiecka, Ewa;Giłka, Wojciech	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB617902C71C2FB5BC062FC38.text	7C3D87FBB617902C71C2FB5BC062FC38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gujaratomyia GILKA & ZAKRZEWSKA 2018	<div><p>GUJARATOMYIA GIŁKA &amp; ZAKRZEWSKA, 2018</p><p>Ty p e s p e c i e s: G u j a r a t o m y i a m i r i p e s G i ł k a &amp; Zakrzewska, 2018.</p></div>	https://treatment.plazi.org/id/7C3D87FBB617902C71C2FB5BC062FC38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zakrzewska, Marta;Singh, Hukam;Wagner-Wysiecka, Ewa;Giłka, Wojciech	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB617902C722BFF5DC5DFFB6B.text	7C3D87FBB617902C722BFF5DC5DFFB6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gujaratomyia miripes Gilka & Zakrzewska 2018	<div><p>GUJARATOMYIA MIRIPES GIŁKA &amp; ZAKRZEWSKA, 2018</p><p>(FIGS 1A–F, 11A, B)</p><p>Material examined</p><p>Adult male, inventory no. BSIP Tad-610 (left: antenna, wing, foreleg, tarsi of mid- and hindlegs and tarsus of right foreleg damaged; Fig. 1A, B) preserved in an 8.5 mm × 4.5 mm × 3.5 mm piece of amber (early  Eocene, ~54 Mya, Tadkeshwar mine, Gujarat state, India; IR spectrum no. 14258 IAA; Fig. 11A). Holotype and paratype, inventory no. BSIP Tad-607 a and b, syninclusions (see Zakrzewska et al., 2018); IR spectrum no. 14259 IAA (Fig. 11B)  .</p></div>	https://treatment.plazi.org/id/7C3D87FBB617902C722BFF5DC5DFFB6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zakrzewska, Marta;Singh, Hukam;Wagner-Wysiecka, Ewa;Giłka, Wojciech	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB611902A7004FD0CC3FAFA34.text	7C3D87FBB611902A7004FD0CC3FAFA34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus van der Wulp 1874	<div><p>TANYTARSUS VAN DER WULP, 1874</p><p>Type species:  Tanytarsus signatus (van der Wulp, 1858) .</p><p>TANYTARSUS FORFEX GIŁKA &amp; ZAKRZEWSKA SP.</p></div>	https://treatment.plazi.org/id/7C3D87FBB611902A7004FD0CC3FAFA34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zakrzewska, Marta;Singh, Hukam;Wagner-Wysiecka, Ewa;Giłka, Wojciech	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB61090277187F952C35BFC57.text	7C3D87FBB61090277187F952C35BFC57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus ramus Gilka & Zakrzewska 2020	<div><p>TANYTARSUS RAMUS GIŁKA &amp; ZAKRZEWSKA SP. NOV.</p><p>(FIGS 4A–F, 5A–F, 11D)</p><p>LSID: urn:lsid:zoobank.org:act: D8802738-369F- 4D19-823F-379C31FFA80B</p><p>Type material</p><p>Holotype, inventory no. BSIP Tad-521: adult male (tarsi of left foreleg and of both mid- and hindlegs missing; Fig. 4A, B) in a 7 mm × 5 mm × 2.5 mm piece of amber preserved in a cubicoid plastic mass (early  Eocene, ~54 Mya, Tadkeshwar mine, Gujarat state, India; IR spectrum no. 14257 IAA, Fig. 11D).</p><p>Etymology</p><p>From Latin  ramus, branch or twig, refering to the frail, simply branched median volsellae. Noun in apposition.</p><p>Diagnosis</p><p>Wing vein Sc extending well beyond RM; FCu placed well distal to RM; distance between ends of R</p><p>1</p><p>–R</p><p>2 + 3</p><p>and R 2 + 3 –R 4 + 5 unequal, VR C ~5. Tibial combs and spurs stout, combs fan shaped, only one comb with spur on mid-leg, each comb with spur on hindleg. Anal tergite with three spinulae in a row, placed in pit surrounded by crest-like ridges. Anal point slender, long, reaching apex of inferior volsella. Superior volsella pipe shaped. Stem of median volsella twig shaped, branched into two simple, slender lamellae. Inferior volsella parallel sided.</p><p>Description</p><p>Adult male (N = 1). Total length ~ 2.1 mm; wing length ~840 μm.</p><p>Head (Fig. 4B): Eyes bare, slightly produced in dorsomedian position. Frontal tubercles not observed. Antenna with 13 flagellomeres, AR unmeasurable, plume fully developed. Length of palpomeres 2–5 (in micrometres): ~25, 105, 105 and 169. At least 12 strong clypeals.</p><p>Thorax: Dc at least nine, Pa at least one, other thoracic setae unobservable.</p><p>Wing (Fig. 4C): Anal lobe relatively well developed, broadest at two-thirds of length, width 265 μm, length/ width ratio 3.17. Sc extending well beyond RM. RM slightly oblique relative to M and R. R 1 and R 2 + 3 arcuate in distal section. FCu placed well distal to RM; VR Cu 1.35. Veins ending as follows (in order from base to tip): An, Sc, Cu 1, R 1, R + 2 3, M + 3 4, R + 4 5, M + 1 2; distance between ends of R 1 –R 2 + 3 and R 2 + 3 –R 4 + 5 unequal; VR C ~5. Wing membrane covered with macrotrichia in distal half.</p><p>Legs: Tibia of foreleg with stout spur ~25 μm long. Tibial combs of mid leg fan shaped, teeth ≤ 25 μm long, only one comb bearing slender spur ~55 μm long (Fig. 4D, E); combs of hindleg broadly separated, fan shaped; one comb broad, with teeth ≤ 25 μm long, bearing spur ~50 μm long; second comb slender, teeth ≤ 30 μm long, bearing spur ~60 μm long (Fig. 4F). For the leg segment lengths and leg ratios, see Table 3.</p><p>Hypopygium (Fig. 5A–F): Gonostylus slightly longer than gonocoxite, ~85 μm long, tapering towards pointed apex. Anal tergite semicircular, bearing three spinulae arranged in a row, placed in rounded, longitudinal pit, its crest-like ridges tapering towards base of anal point; several setae on each side. Anal point long, reaching apex of inferior volsella, slender, parallel sided with blunt tip; severalstrongposterolateralsetaeoneachside(Fig.5A,B). Superior volsella pipe shaped, with oval basal part, distinctly narrowed at mid-length, distal part elongate with apex blunt; several short setae at median margin of basal part (Fig. 5B, D). Digitus not observed. Stem of median volsella ~20 μm long, posteromedially directed, twig shaped, branched into two simple, slender lamellae (Fig. 5C, E, F). Inferior volsella reaching half the length of the gonostylus, nearly parallel sided, with distal half armed, bearing numerous stout setae.</p><p>Remarks</p><p>Tanytarsus ramus differs from all known species of the genus by a set of distinct characters as follows: the VR C ratio is nearly five (Fig. 4C), the anal tergite has a couple of spinulae placed in a row in a large pit, and the anal point is slender and extraordinarily long (Fig. 5A, B); the pipe shaped superior volsella should be considered a key feature (Fig. 5B, D), whereas the digitus, typical of  Tanytarsus, is not observed (probably absent). The most peculiar structure diagnostic of  T. ramus is the hypopygial median volsella (Fig. 5E, F) of a shape perceived as a plesiomorphy (see Discussion: Evolution of the diagnostic structures).</p></div>	https://treatment.plazi.org/id/7C3D87FBB61090277187F952C35BFC57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zakrzewska, Marta;Singh, Hukam;Wagner-Wysiecka, Ewa;Giłka, Wojciech	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB61C90277276FB11C492FC13.text	7C3D87FBB61C90277276FB11C492FC13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stempellina Thienemann & Bause 1913	<div><p>STEMPELLINA THIENEMANN &amp; BAUSE, 1913</p><p>Type species:  Stempellina bausei (Kieffer, 1911) .</p><p>STEMPELLINA STEBNERI GIŁKA &amp; ZAKRZEWSKA SP.</p></div>	https://treatment.plazi.org/id/7C3D87FBB61C90277276FB11C492FC13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zakrzewska, Marta;Singh, Hukam;Wagner-Wysiecka, Ewa;Giłka, Wojciech	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB61C90277075FAE0C421FD74.text	7C3D87FBB61C90277075FAE0C421FD74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus SPP.	<div><p>TANYTARSUS SPP.</p><p>(FIG. 6A–F)</p><p>Material examined</p><p>Ta n y t a r s u s s p. 1, o n e a d u l t m a l e i n a   n 8.5 mm × 6 mm × 3.5 mm piece of amber (inventory no. BSIP Tad-533; Fig. 6A, B);  Tanytarsus sp. 2, one adult male in a 6.5 mm × 5 mm × 4 mm piece of amber (inventory no. BSIP Tad-883 a; Fig. 6C, D), syninclusion:  Coleoptera (inventory no. BSIP Tad-883 b);  Tanytarsus sp. 3, one adult male in a 6 mm × 3 mm × 1 mm piece of amber preserved in a cubicoid plastic mass (inventory no. BSIP Tad-884; Fig. 6E, F);  Early Eocene, ~54 Mya, Tadkeshwar mine, Gujarat state, India  .</p><p>Remarks</p><p>The above three specimens are ascribed to the genus  Tanytarsus on the basis of the head, wing and hypopygium characters (cf. Stebner et al., 2017). However, most of the diagnostic structures are weakly observable, deformed or insufficiently preserved to define the individuals definitively to species level. Nevertheless, these males differ distinctly from each other and from those designated as new species described in this paper, thus they most probably belong to another species. Therefore, with presumably five species,  Tanytarsus is the most species-rich genus among the Cambay  Tanytarsini . The specific diversity proportions within the tribe may thus be similar relative to the extant fauna: five  Tanytarsus / eight  Tanytarsini in Cambay amber vs. ~360  Tanytarsus /~700  Tanytarsini extant species (Giłka W, Zakrzewska M, unpublished data).</p><p>SUBTRIBE:  ZAVRELIINA SAETHER, 1977</p></div>	https://treatment.plazi.org/id/7C3D87FBB61C90277075FAE0C421FD74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zakrzewska, Marta;Singh, Hukam;Wagner-Wysiecka, Ewa;Giłka, Wojciech	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB61890237010FBEAC3EDFC29.text	7C3D87FBB61890237010FBEAC3EDFC29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stempellinella Brundin 1947	<div><p>STEMPELLINELLA BRUNDIN, 1947</p><p>Type species:  Stempellinella saltuum (Goetghebuer, 1921) .</p><p>STEMPELLINELLA POLLEX GIŁKA &amp; ZAKRZEWSKA SP.</p></div>	https://treatment.plazi.org/id/7C3D87FBB61890237010FBEAC3EDFC29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zakrzewska, Marta;Singh, Hukam;Wagner-Wysiecka, Ewa;Giłka, Wojciech	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB6069039723CF880C389F9FF.text	7C3D87FBB6069039723CF880C389F9FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stempellina Thienemann & Bause 1913	<div><p>In  Tanytarsini, the median volsella is an appendage of the male gonocoxite, consisting of a stem that bears lamellae of highly diverse shapes (Saether, 1980). The median volsellae, with a couple of exceptions (i.e. when reduced, cf. Ekrem &amp; Reiss, 1999), are thus the most important diagnostic structures in  Tanytarsini . In  T. ramus, the MVo is simple, consisting of a twig shaped stem, branched into two lamellae (Fig. 5E, F). If we treat  T. ramus as one of the oldest known tanytarsines,</p>19. Ultimate palpomere with strong, stiff apical seta; anal tergite with longitudinal crest-like hump bearing median setae (Giłka, 2011: figs 1, 2) ................................  Tanytarsus fereci (Baltic region: Gulf of Gdańsk) Ultimate palpomere without stiff apical seta; anal tergite without longitudinal hump, median setae absent (Giłka et al., 2013: fig. 7; Zakrzewska &amp; Giłka, 2015a: fig. 8; Zakrzewska et al., 2016: fig. 5) ......20 20. Gonostylus straight, with long seta on apex; superior volsella finger like, distinctly curved at mid-length and directed medially; median volsella sickle shaped, with slender arcuate lamellae (Zakrzewska et al., 2016: fig. 5) .....................................................................  Tanytarsus crocota (Baltic region: Gulf of Gdańsk) Gonostylus curved at mid-length or boomerang like, with subapical tooth-like process at most; superior volsella bean shaped or with nipple-like apical extension, straight, directed posteromedially or posteriorly; median volsella never sickle shaped, bearing leaf shaped (foliate) lamellae (Giłka et al., 2013: fig. 7; Zakrzewska &amp; Giłka, 2015a: fig. 8; Zakrzewska et al., 2016: fig. 6).............................................................21 21. Stem of median volsella bulb shaped (Giłka et al., 2013: fig. 7) ...................................................................... .............................................................................................  Tanytarsus congregabilis (Baltic region: Rovno) Stem of median volsella elongated, straight (Zakrzewska &amp; Giłka, 2015a: fig. 8) ........................................ ........................................................................  Tanytarsus PROTOGREGARIUS (Baltic region: Gulf of Gdańsk) 22. Gonostylus distinctly longer than gonocoxite (Fig. 8; Seredszus &amp; Wichard, 2007: fig. 18)  Stempellina … 23  Gonostylus shorter than or as long as gonocoxite (Fig. 10; Seredszus &amp; Wichard, 2007: fig. 19; Zakrzewska &amp; Giłka, 2014: fig. 4; Zakrzewska &amp; Giłka, 2015a: fig. 4; Zakrzewska et al., 2016: fig. 8) ....  Stempellinella ........................................................................................................................................................................ 24 23. Anal point spinulae absent; superior volsella twig shaped, strongly elongated: length/width ratio nearly ten (Seredszus &amp; Wichard, 2007: fig. 18) .....................  Stempellina exigua (Baltic region: Gulf of Gdańsk) Anal point spinulae present; superior volsella cylindrical, apically rounded, slightly elongated: length/ width ratio nearly two (Fig. 8) ...........................................................  Stempellina stebneri (India: Cambay) 24. Inferior volsella with stout process on median margin (Fig. 10) .................................................................... .............................................................................................................  Stempellinella pollex (India: Cambay) Inferior volsella without process on median margin ...................................................................................25 25. Gonostylus with apical nipple-like process; anal point with spinulae; superior volsella directed posteriorly; inferior volsella with beak-like protrusion (Zakrzewska &amp; Giłka, 2014: fig. 4) .............................................. ....................................................................  Stempellinella ivanovae (Baltic region: Gulf of Gdańsk, Rovno) Gonostylus without apical nipple-like process; anal point without spinulae; superior volsella directed medially; inferior volsella without beak-like protrusion (Seredszus &amp; Wichard, 2007: fig. 19; Zakrzewska &amp; Giłka, 2015a: fig. 4; Zakrzewska et al., 2016: fig. 8) ................................................................................ 26 26. Antennal flagellum consisting of ten flagellomeres; median volsella shorter than superior volsella (Seredszus &amp; Wichard, 2007: fig. 19) ......................  Stempellinella bicorna (Baltic region: Gulf of Gdańsk) Antennal flagellum consisting of 13 flagellomeres, flagellomeres 11–13 or 12–13 partly fused (Zakrzewska &amp; Giłka, 2015a: fig. 3; Zakrzewska et al., 2016: fig. 7); median volsella longer than superior volsella (Zakrzewska &amp; Giłka, 2015a: fig. 4; Zakrzewska et al., 2016: fig. 8) ......................................................... 27 27. Median volsella (stem + lamellae) distinctly shorter than gonostylus, bearing wide pectinate and foliate lamellae (Zakrzewska &amp; Giłka, 2015a: fig. 4) .........  Stempellinella electra (Baltic region: Gulf of Gdańsk) Median volsella (stem + lamellae) and gonostylus of similar length, bearing slender foliate lamellae, single lamella with strongly elongated filiform tip (Zakrzewska et al., 2016: fig. 8) ..................................... .......................................................................................  Stempellinella fibra (Baltic region: Gulf of Gdańsk)<p>we then assume that this type of volsella might display one of the simplest/oldest types of MVo structure in  Tanytarsini (plesiomorphy), comparable to that known from the extinct genus  Archistempellina Giłka</p><p>&amp; Zakrzewska, 2013 described from Baltic amber (cf. Giłka et al., 2013; Zakrzewska &amp; Giłka, 2014).</p></div>	https://treatment.plazi.org/id/7C3D87FBB6069039723CF880C389F9FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zakrzewska, Marta;Singh, Hukam;Wagner-Wysiecka, Ewa;Giłka, Wojciech	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
