identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8E7F87986466FFF6FF047AE9B1BC3CFB.text	8E7F87986466FFF6FF047AE9B1BC3CFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotyphlinae Fauvel 1874	<div><p>Leptotyphlinae Fauvel, 1874</p><p>Diagnosis. Very small beetles, body length about 0.8–1.9 mm; depigmented; eyes completely atrophied. Head capsule large, neck region short; gular sutures always well-defined, confluent or not in middle or anterior third of head length; mandibles strong, symmetric with one or more teeth; maxillary palpi composed from four palpomeres, palpomere 2 and 3 of different shape and size, tribe characteristic. Antennae with 10 or 11 antennomeres, scape and pedicel always large; antennal club composed from two to five antennomeres. Pronotum about as long as head or slightly shorter. Elytra reduced, slightly wider than long; lacking hind wings. Abdomen longer than head, pronotum and elytra combined; visible sternites simple, slightly impressed or with deep transverse sulcus at posterior region.  Leptotyphlinae are actually classified into six tribes which can be separated by following key.</p><p>1 All visible tergites with well-defined paratergites............................................................ 2</p><p>- Visible tergites 2–5 lacking paratergites. Australia ....................................  Australiotyphlini Pace, 2014</p><p>2 Visible sternite 5 with deep, transversal sulcus (Fig. 3), in some genera deep, transversal sulcus is present on visible sternites 2–5 ( Cephalotyphlus Coiffait, 1963,  Romanotyphlus Struyve, 2022 and  Paratyphlus Blackwelder, 1952)............... 3</p><p>- All visible sternites lacking deep, transversal sulcus, at most with shallow depression............................... 4</p><p>3 Maxillary palpi with palpomere 2 strongly dilated, distinctly larger than palpomere 3. The whole Mediterranean region, Canary Islands and Romania ...........................................................  Entomoculiini Coiffait, 1957</p><p>- Maxillary palpi with palpomeres 2 and 3 dilated, palpomere 3 larger than palpomere 2. France and Italy ...........................................................................................  Cephalotyphlini Coiffait, 1963</p><p>4 Palpomere 2 strongly dilated, much more developed than palpomere 3. Mediterranean region, Slovakia, Tanzania, The Democratic republic of Congo ....................................................  Leptotyphlini Fauvel, 1874</p><p>- Both palpomeres 2 and 3 about equally dilated.............................................................. 5</p><p>5 Fore coxal cavities rounded on its anterior margin, with or without small excision on lateral side of its anterior margin; proepisternum and prosternum fused, not separated by suture; ligula not distinct; mandibles not falciform. Europe, Chile, Mexico (undescribed species) and Venezuela ........................................  Metrotyphlini Coiffait, 1963</p><p>- Fore coxal cavities on its anterior margin with well-defined excision which is prolonged anteriad by suture separating proepisternum from prosternum; ligula distinct, represented with pair of well visible, widely-separated projections; mandibles falciform. The most widely distributed tribe, known from the mediterranean region, Romania, Serbia, Slovenia, South Africa, USA (California, Idaho, Florida, Alaska), Belize, Cuba, Guatemala and Chile ................  Neotyphlini Coiffait, 1963</p></div>	https://treatment.plazi.org/id/8E7F87986466FFF6FF047AE9B1BC3CFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFF6FF047AFDB6183A2C.text	8E7F87986461FFF6FF047AFDB6183A2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus Coiffait 1955	<div><p>Allotyphlus Coiffait, 1955</p><p>Allotyphlus Coiffait, 1955: 67 .</p><p>Type species:  Leptotyphlus aspericeps Scheerpeltz, 1931 . Original designation.</p><p>Allotyphlus Coiffait: Coiffait, 1957: 61 (in key); Coiffait, 1959: 310 (redescription, key to species); Pace, 1977b: 197 (key to species of Apennine Mountains); Pace, 1996: 124 (redescriptions, key to Italian species); Struyve, 2022: 10 (in key).</p><p>Syn.  Tropidotyphlus Scheerpeltz, 1956: 459 . Type species:  Leptotyphlus aspericeps Scheerpeltz, 1931 . Original designation. Synonymy in Coiffait (1959: 310).</p><p>sbg. Moreotyphlus Coiffait, 1959: 312. Type species:  Allotyphlus spartensis Coiffait, 1959 . Original designation.</p><p>Diagnosis. Gular sutures confluent in anterior part of head length, from confluent point continuing as single, straight, short suture or again separated to two divergent sutures; antennae with 11 antennomeres, antennomeres 5 larger than 4 and 6, antennal club with 3 antennomeres; labrum dentate with anterior margin rectilinear; tarsal formula 3-3-3; aedeagus asymmetric, parameres of different shape, with distal setae.</p><p>Remarks. Large genus with 25 valid species names assigned in two subgenera. Five species are known from Italy (Trentino-Alto Adige, Veneto, Puglia), three species from Turkey, all from the province Burdur and 17 species from Greece, 10 from mainland (Epirus, Peloponnese, Teygetos, Eastern Macedonia and Thrace) and 7 from islands (Crete, Kefaloniá, Kérkira, Lefkáda).</p></div>	https://treatment.plazi.org/id/8E7F87986461FFF6FF047AFDB6183A2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFF6FF047802B1BC3FA0.text	8E7F87986461FFF6FF047802B1BC3FA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Entomoculiini Coiffait 1957	<div><p>Entomoculiini Coiffait, 1957</p><p>Diagnosis. Mandibles lacking premolar tooth; antennae with 10 or 11 antennomeres; all visible tergites with well-defined paratergites; deep, basal transversal sulcus at least on visible abdominal sternite 5, except for  Romanotyphlus and  Paratyphlus where it is present also on visible sternites 2–4, later genus known from northern Africa, Balearic Islands, Iberian Peninsula and Sardinia); maxillary palpi with palpomere 2 strongly dilated, distinctly larger than palpomere 3.</p><p>Remarks.  Entomoculiini with seven genera and 256 valid species names is the second most speciose tribe of the subfamily. It has exclusively west Palaearctic distribution and is known from the whole Mediterranean region, going from Portugal up to Turkey, northern Africa, Romania and now also from Serbia. Three Balkan genera, (the distribution Figs 89, 90) can be separated by the following key:</p><p>1 All tarsi with three tarsomeres............................................................  Allotyphlus Coiffait</p><p>- All tarsi with two tarsomeres............................................................................ 2</p><p>2 Antennal club composed of two antennomeres; antennomere 5 about as large as 4 and 6; aedeagus symmetric; parameres strongly enlarged, largely covering distal part of median lobe.................................  Cyrtotyphlus Dodero</p><p>- Antennal club composed of five antennomeres; antennomere 5 larger than 4 and 6; aedeagus asymmetric, parameres of different shape...........................................................................  Romanotyphlus Struyve</p></div>	https://treatment.plazi.org/id/8E7F87986461FFF6FF047802B1BC3FA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFFAFF047F71B597392D.text	8E7F87986461FFFAFF047F71B597392D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus (Moreotyphlus) stolensis Hlavac, Janak & Banar 2025	<div><p>Allotyphlus (Moreotyphlus) stolensis Hlaváč, Janák &amp; Baňař,  sp. nov.</p><p>Figs 1–18, 83–84</p><p>Material studied.   Holotype, ♂: SERBIA with one label “SERBIA or. 26.v.2024 / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.11492&amp;materialsCitation.latitude=44.16446" title="Search Plazi for locations around (long 22.11492/lat 44.16446)">Mt. Stol</a>, soil washing, 660m / 44.16446N, 22.11492E / J. Janák lgt., hornbeam forest” [white, printed] (JJRC)  .  Paratypes: same data as holotype, 4 ♂♂, 4 ♀♀: (3 ♂♂, 3 ♀♀ in JJRC, 1 ♂, 1 ♀ in MMBC) .</p><p>Diagnosis. Maxillary palpi (Fig. 5) with palpomere 2 dilated, about twice as wide as 3; palpomere 4 small, about three times shorter and narrower than 3; gular sutures confluent anteriorly at about midlength of head capsule, after meeting point again separated to two divergent sutures (Fig. 6); antennae with long scape about as long as terminal antennomere, about 1.6 times as long as wide; pedicel about as long as wide; antennomeres 3–10 transverse; terminal antennomere subtriangular, pointed to apex, slightly longer than wide (Fig.7); visible tergites 1–5 and visible sternite 2–4 with shallow basal impression (Fig. 3); visible sternite 5 (VII) with deep transversal sulcus (Fig. 3, arrow), visible sternite 6 (VIII) of male symmetric, with median posterior excision which is prolonged with wide, deep impression (Figs 11–12); tarsal formula 3-3-3 (Figs 15–16); aedeagus with median lobe prolonged by long slender ventral lamina; parameres short, ventrally left paramere longer than right one, with 3 distal setae (Fig. 17).</p><p>Description. Body (Figs 1, 2) about 1.4 mm long (with abdomen slightly extended posteriad), completely depigmented.</p><p>Head capsule (Fig. 4) slightly longer than wide; wider than pronotum, posterior part deeply inserted into pronotum; basal capsule separated from neck region by occipital constriction, part of head capsule and neck region retracted into pronotum; gular sutures (Fig. 6) confluent anteriorly at about midlength of head capsule, after meeting point again separated to two divergent arms; maxillary palpi (Fig. 5) with palpomere 2 dilated, about twice as wide as 3; palpomere 4 small, about three times shorter and narrower than 3.</p><p>Antennae (Fig. 7) long, scape about as long as terminal antennomere, about 1.6 times as long as wide; pedicel as long as wide, as wide as scape, twice as long as antennomere 3; antennomeres 3–10 transverse; 3, 4 and 6 subequal in length; antennomere 5 larger than 4 and 6; 5 and 7 subequal in length; 8 as long as 7; 8 and 9 twice as wide as long; 9 1.25 times as long as 8; 10 1.2 times as long as 9; terminal antennomere subtriangular, pointed to apex, slightly longer than wide.</p><p>Pronotum (Fig. 8) about 1.1 times as long as wide, widest in anterior quarter, after that convergent posteriad; anterior angles widely rounded; posterior angles sharper; about 1.2 times as long as elytra. Prosternum (Fig. 9) fused with proepisternum, procoxal cavities rounded on anterior margin.</p><p>Elytra (Fig. 8) about as long as wide.</p><p>Abdomen (Figs 1–2) long, considerably longer than length of head capsule, pronotum and elytra combined. Visible tergites 1–5 (IV–VIII) and visible sternites 2–4 (IV–VI) with shallow basal impression, visible sternite 5 (VII) with deep transverse sulcus (Fig. 3, arrow). Visible tergite 6 (IX) of male expanded in posterior half and with rounded apical margin (Fig. 10). Visible sternite 6 (VIII) of male (Figs 11, 12) symmetric, setose in posterior half, about as long as wide, with median posterior excision which is prolonged with wide, deep impression, its surface shagreened and lateral sides bordered by line of short setae. Terminal abdominal segments of female (Fig. 13) with two paired longitudinal sclerites and half-circled shaped structure in middle. Male sternite IX asymmetric, with posterior margin rounded, lateral parts of tergite IX asymmetric, strongly chitinised, tergite X with short pointed projections laterally (Fig. 14).</p><p>Legs short, all tibia distally dilated, tarsal formula 3-3-3 (Figs 15–16).</p><p>Aedeagus (Figs 17, 18) about 0.26 mm long, median lobe prolonged by long slender distal process, basal part about 1.4 times as long as distal process, apex of distal process variable, rounded and slightly curved at apex in lateral view, with sharp, short preapical tooth; sclerite of median lobe well-defined; parameres short, in ventral view left paramere longer than right one, with 3 distal setae (Fig. 17d).</p><p>Etymology. Locotypic, named after Stol planina, the type locality of the species.</p><p>Natural History. All specimens were collected by the soil-washing in a belt of hornbeam forest with hazel, maple and oak on a north-facing slope above a stream, together with  Banatotyphlus siclus . The volume of the sample had about 100 liters (Figs 83–84).</p><p>Remarks.  Allotyphlus (Moreotyphlus) stolensis has similar aedeagus as  A. candicus and  A. foedatus (both from Greece, Crete) with long ventral lamina. It can be readily separated from late two species by the distal part of the ventral lamina slightly curved with rounded apex. The apex of the median lobe of  A. candicus and  A. foedatus is sharp, not curved and it is much shorter in the relation of the length of basal part of median lobe.</p><p>Distribution. Serbia.</p></div>	https://treatment.plazi.org/id/8E7F87986461FFFAFF047F71B597392D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646DFFFAFF047F1FB6033AFE.text	8E7F8798646DFFFAFF047F1FB6033AFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus achileus Coiffait 1959	<div><p>achileus Coiffait, 1959: 402</p><p>( Allotyphlus, TL: Ayos Deka [Ayioi Dhéka], près de l’Achileion [Akhílion] à quelques kilomètres au sud de Corfou (Corfou, Iles Ioniennes)).</p><p>Types: MNHN HT ♂, 2PT (MNHN) .</p><p>Other localities: PT: Ayos Theodori, près de l’Achileion [Akhílion] à quelques kilomètres au sud de Corfou (Corfou, Iles Ioniennes).</p><p>Distribution: Greece (Kérkira).</p></div>	https://treatment.plazi.org/id/8E7F8798646DFFFAFF047F1FB6033AFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646DFFFAFF047FC6B6033B96.text	8E7F8798646DFFFAFF047FC6B6033B96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus corcyranus Coiffait 1959	<div><p>corcyranus Coiffait, 1959: 401</p><p>( Allotyphlus, TL: Dassia, au pied de Mont Pandokrator à quelques kilomètres au nord de Corfou (Corfou, Iles Ioniennes)).</p><p>Types: MNHN HT ♂, 10PT (MNHN) .</p><p>Other localities: PT: Gouvia and Ipsos, au pied de Mont Pandokrator à quelques kilomètres au nord de Corfou (Corfou, Iles Ioniennes)).</p><p>Distribution: Greece (Kérkira).</p></div>	https://treatment.plazi.org/id/8E7F8798646DFFFAFF047FC6B6033B96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646DFFFBFF047EAFB6033CE4.text	8E7F8798646DFFFBFF047EAFB6033CE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus corcyricus Scheerpetlz 1958	<div><p>corcyricus Scheerpetlz, 1958: 390</p><p>( Tropidotyphlus, TL: im Val de Ropa auf der Insel Korfu)</p><p>Types: 7ST (FMNH, NMW) .</p><p>Reference: Pace, 1983a: 452 (supplement description).</p><p>Distribution: Greece (Kérkira).</p></div>	https://treatment.plazi.org/id/8E7F8798646DFFFBFF047EAFB6033CE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047B77B6373F22.text	8E7F8798646CFFFBFF047B77B6373F22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus ambracius Scheerpeltz 1958	<div><p>ambracius Scheerpeltz, 1958: 392</p><p>( Tropidotyphlus, TL: Umbebung von Arta auf einem trockenen Hügelrücken, 50 m).</p><p>Types: 13 ST (NMW) .</p><p>References: Pace, 1983a: 455 (supplement description).</p><p>Distribution: Greece (Epirus).</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFFBFF047B77B6373F22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047AD0B62A38AD.text	8E7F8798646CFFFBFF047AD0B62A38AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus candicus Assing 2018	<div><p>candicus Assing, 2018: 9</p><p>( Allotyphlus, TL: Greece: Crete, SW Sitia, Kimouriotis, 35°10‘40‘‘N, 26°03‘00‘‘E, 110 m).</p><p>Types: NMW (Assing coll.) HT ♂, 11 PT (NMW (Assing coll.)) .</p><p>Other localities: Greece: Crete: W Kritsa, 1050 m; S Kritsa, SW Kroustas, 960 m; SW Sitia, NW Magrigialos, 70 m.</p><p>Distribution: Greece (Crete).</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFFBFF047AD0B62A38AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047DB9B61C3938.text	8E7F8798646CFFFBFF047DB9B61C3938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus cephallenicus Pace 1983	<div><p>cephallenicus Pace, 1983a: 456</p><p>( Allotyphlus, TL: Cephallinia, M. de Rudi).</p><p>Types: NMW HT ♂, 3 PT (NMW, Pace coll.))</p><p>Distribution: Greece (Kefaloniá).</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFFBFF047DB9B61C3938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF0479C1B6033DB8.text	8E7F8798646CFFFBFF0479C1B6033DB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus dexters Coiffait 1959	<div><p>dexters Coiffait, 1959: 402</p><p>( Allotyphlus, TL: Doukades [Dhoukádes], nord-ouest de l’île de Corfou (Iles Ioniennes)).</p><p>Types: 6ST (MNHN) .</p><p>Other localities: PT: Paleokastritsa, nord-ouest de l’île de Corfou (Iles Ioniennes)).</p><p>Distribution: Greece (Kérkira).</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFFBFF0479C1B6033DB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047C0DB62A3A18.text	8E7F8798646CFFFBFF047C0DB62A3A18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus foedatus Assing 2019	<div><p>foedatus Assing, 2019: 261</p><p>( Allotyphlus, TL: Greece: Crete, SW Chania, Mavros stream valley, 35°24‘10“N, 23°53‘53“E, 290 m).</p><p>Types: NMW (Assing coll.) HT ♂, 136PT (NMW (Assing coll.), ZMHB) .</p><p>Other localities: Greece, Crete: SW Chania, W Lakki, 390 m; S Chania, N Thersio, 610 m; SW Chania, WSW Chliaro, 210 m; SW Chania, W Phrases, 640 m; SW Chania, S Nteres, 280 m; S Kissamos, NE Sirikari, 500 m; S Kissamos, Archontiko, 580 m; S Kissamos, Strovles, 470 m; S Kissamos, Elos, 580 m; E Kandanos, Anisaraki, 670 m; S Asigonia, E Kallikratis, 770 m.</p><p>Distribution: Greece (Crete).</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFFBFF047C0DB62A3A18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047F2EB6433A87.text	8E7F8798646CFFFBFF047F2EB6433A87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus karamani Struyve 2024	<div><p>karamani Struyve, 2024: 275</p><p>( Allotyphlus, TL: Greece, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.485&amp;materialsCitation.latitude=36.774" title="Search Plazi for locations around (long 22.485/lat 36.774)">Pelopponese</a>, Mani, Gytheo Platanos, 36.774°N, 22.485°E).</p><p>Types: MMBC HT ♂, 7PT (coll. Pavićević, coll. Struyve)</p><p>Distribution: Greece (Peloponnisos).</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFFBFF047F2EB6433A87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047FDFB6373B12.text	8E7F8798646CFFFBFF047FDFB6373B12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus loebli Coiffait 1973	<div><p>loebli Coiffait, 1973: 220</p><p>( Allotyphlus, TL: Grèce, Philiates).</p><p>Types: MHNG HT ♂, 2PT (MHNG, MNHN) .</p><p>Distribution: Greece (Epirus).</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFFBFF047FDFB6373B12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047E33B61C3B82.text	8E7F8798646CFFFBFF047E33B61C3B82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus moczarskii Pace 1983	<div><p>moczarskii Pace, 1983a: 457</p><p>( Allotyphlus, TL: Cephalonia, Argostoli).</p><p>Types: NMW HT ♂, 7 PT (NMW, MCNV) .</p><p>Other localities: Cephallonia, Megalo Vuno; Cephallonia, Krano; Cephallonia, M. Nero.</p><p>Distribution: Greece (Kefaloniá).</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFFBFF047E33B61C3B82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF04788DB5703E6F.text	8E7F8798646CFFFBFF04788DB5703E6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus sinester Coiffait 1959	<div><p>sinester Coiffait, 1959: 402</p><p>( Allotyphlus, TL: Paleokastritsa and Lakonès, nord-ouest de l’île de Corfou (Iles Ioniennes)).</p><p>Types: MNHN HT ♂, 5PT (MNHN)</p><p>Distribution: Greece (Kérkira).</p><p>sbg. Moreotyphlus Coiffait, 1959: 310, 312. Type species:  Allotyphlus spartensis Coiffait, 1959 . Original designation.</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFFBFF04788DB5703E6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047A03B60A3FF6.text	8E7F8798646CFFFBFF047A03B60A3FF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotyphlus aspericeps Scheerpeltz 1931	<div><p>aspericeps Scheerpeltz, 1931: 396</p><p>( Leptotyphlus, TL: bei Kaligoni [Kalligónion] auf Levkas).</p><p>Types: NMW HT ♂, 9 PT (NMW) .</p><p>Other localities: Perivolakia.</p><p>References: Pace, 1983a: 454 (supplement description).</p><p>Distribution: Greece (Lefkáda).</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFFBFF047A03B60A3FF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFF8FF047EA4B6433CE4.text	8E7F8798646CFFF8FF047EA4B6433CE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus naupliensis Coiffait 1959	<div><p>naupliensis Coiffait, 1959: 404</p><p>( Allotyphlus, TL: Yannouleika [Yiannoulaíika] près de Nauplion [Návplion] (Péloponnèse)).</p><p>Types: MNHN HT ♂, 1PT (MNHN) .</p><p>Distribution: Greece (Peloponnisos).</p></div>	https://treatment.plazi.org/id/8E7F8798646CFFF8FF047EA4B6433CE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF0479C1B6433D70.text	8E7F8798646FFFF8FF0479C1B6433D70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus patrensis Coiffait 1959	<div><p>patrensis Coiffait, 1959: 404</p><p>( Allotyphlus, TL: Kamarès [Kamáres] près de Patras (Péloponnèse)).</p><p>Types: MNHN HT ♂.</p><p>Distribution: Greece (Peloponnisos).</p></div>	https://treatment.plazi.org/id/8E7F8798646FFFF8FF0479C1B6433D70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047855B6433DC4.text	8E7F8798646FFFF8FF047855B6433DC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus spartensis Coiffait 1959	<div><p>spartensis Coiffait, 1959: 404</p><p>( Allotyphlus, TL: Tripi [Trípi] près de Sparte, au pied du versant Est du Taygète [Taygetos] (Péloponnèse)).</p><p>Types: MNHN HT ♂, 9PT (MNHN) .</p><p>Other localities: PT: Piyai [Piyaí] sur versant Ouest du Taygète [Taygetos] (Péloponnèse).</p><p>Distribution: Greece (Peloponnisos).</p></div>	https://treatment.plazi.org/id/8E7F8798646FFFF8FF047855B6433DC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF0478E1B58D3E0C.text	8E7F8798646FFFF8FF0478E1B58D3E0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allotyphlus stolensis Hlaváč & Janák & Baňař 2025	<div><p>stolensis sp. nov.</p><p>Distribution: Serbia.</p></div>	https://treatment.plazi.org/id/8E7F8798646FFFF8FF0478E1B58D3E0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047A01B74838C7.text	8E7F8798646FFFF8FF047A01B74838C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyrtoptyphlus Dodero 1900	<div><p>Cyrtoptyphlus Dodero, 1900</p><p>Cyrtotyphlus Dodero, 1900a: 401 .</p><p>Type species:  Cyrtotyphlus convexus Dodero, 1900 . Designation by monotypy.</p><p>Cyrtotyphlus Dodero: Coiffait, 1957: 61 (in key); Coiffait, 1959: 276 (redescription, key to species); Pace, 1976: 45 (diagnosis, distribution); Pace, 1977b: 196 (key to species of Apennine Mountains); Pace, 1996: 75 (redescription, key to Italian species); Struyve, 2022: 10 (in key).</p><p>Diagnosis. Gular sutures confluent in anterior third of head, finished in confluent point; antennae with 10 or 11 antennomeres; antennal club composed of two antennomeres; antennomere 5 about as large as 4 and 6; tarsal formula 2-2-2; aedeagus symmetric with strongly enlarged parameres, largely covering distal part of median lobe, with some distal setae.</p><p>Remarks. Smaller genus, with 7 species, 3 species are in France and Italy, one Italian species is also recorded from Slovenia, two species are in Greece, one in Eastern Macedonia and Thrace and one on the island Samothráki. One species is known from Montenegro and one from Romania.</p></div>	https://treatment.plazi.org/id/8E7F8798646FFFF8FF047A01B74838C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047CABB6733A6F.text	8E7F8798646FFFF8FF047CABB6733A6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyrtotyphlus carpathicus Struyve 2022	<div><p>carpathicus Struyve, 2022: 8</p><p>( Cyrtotyphlus, TL: Romania, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.393&amp;materialsCitation.latitude=45.171" title="Search Plazi for locations around (long 24.393/lat 45.171)">Sânbotin</a>, 45.171°N, 24.393°E). Types: HNHM HT ♂; 2PT (coll. Struyve).</p><p>Distribution: Romania.</p><p>Note. The species was originally described as  C. carpathica .  Cyrtotyphlus is a masculine, thus the specific name must be changed to  C. carpathicus .</p></div>	https://treatment.plazi.org/id/8E7F8798646FFFF8FF047CABB6733A6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047F77B5CE3AFB.text	8E7F8798646FFFF8FF047F77B5CE3AFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyrtotyphlus kaiseri Pace 1983	<div><p>kaiseri Pace, 1983a: 451</p><p>( Cyrtotyphlus, TL: Begovina bei Castelnuovo [Herzeg Novi], Süd-Dalmatien).</p><p>Types: NMW HT ♂ .</p><p>Distribution: Montenegro.</p></div>	https://treatment.plazi.org/id/8E7F8798646FFFF8FF047F77B5CE3AFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047FCBB74B3BF7.text	8E7F8798646FFFF8FF047FCBB74B3BF7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyrtotyphlus winkleri Breit 1913	<div><p>winkleri Breit, 1913: 235</p><p>( Cyrtotyphlus, TL: Nanos in Krain).</p><p>Types: unknown.</p><p>Other localities: Italy: Veneto (Recoaro: Fongara; Monti Berici: Zovencedo, Arcugnano, Lumignano; Arquà Petrarca: Monte Fasolo; Monte Venda, Marostica, Soligo), Friuli-Venezia Giulia (Taipana, Strgna, Cormons, Premariacco,  Dolegna del Collio); Slovenia: Planina; Logateč.</p><p>References: Coiffait, 1962: 159 (upgrade from subspecies to species level).</p><p>Distribution: Italy (Veneto, Friuli-Venezia Giulia), Slovenia.</p></div>	https://treatment.plazi.org/id/8E7F8798646FFFF8FF047FCBB74B3BF7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047D9EB05F39EF.text	8E7F8798646EFFF9FF047D9EB05F39EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotyphlini Fauvel 1874	<div><p>Leptotyphlini Fauvel, 1874</p><p>Diagnosis. Maxillary palpomere 2 strongly dilated, much more developed than palpomere 3; tarsi with 2 or 3 tarsomeres; prosternal coxal cavities rounded on anterior margin, lacking excision; lacking suture separating proepisternum from prosternum; all visible tergites with well-defined paratergites; all visible abdominal sternites lacking deep basal transverse sulcus.</p><p>Remarks. The richest tribe of the subfamily. It holds 10 genera and 264 valid species names. The large majority of species is in the Mediterranean region. Out of this region, there are three species in the Democratic Republic of Congo, one in Tanzania, one in Indonesia (West Papua) and one,  Leptotyphlus kovaci Šustek, 2000, in Slovakia. Only one genus,  Kenotyphlus (distribution on Fig. 91) is present in the Balkan Peninsula.</p></div>	https://treatment.plazi.org/id/8E7F8798646EFFF9FF047D9EB05F39EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047855B7093D9D.text	8E7F8798646EFFF9FF047855B7093D9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Egeotyphlus thracicus Coiffait 1957	<div><p>thracicus Coiffait, 1957: 62</p><p>( Cyrtotyphlus, TL: Thrace grecque: Sapé).</p><p>Types: unknown.</p><p>Distribution: Greece (Eastern Macedonia and Thrace).</p></div>	https://treatment.plazi.org/id/8E7F8798646EFFF9FF047855B7093D9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047F2EB0283BB5.text	8E7F8798646EFFF9FF047F2EB0283BB5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kenotyphlus Coiffait 1957	<div><p>Kenotyphlus Coiffait, 1957</p><p>Kenotyphlus Coiffait, 1957: 80 .</p><p>Type species:  Kenotyphlus turcicus Coiffait, 1957 . Original description.</p><p>Kenotyphlus Coiffait: Coiffait, 1957: 61 (in key), 80 (diagnosis); Coiffait, 1972: 602 (redescription, key to species of Turkey).</p><p>Diagnosis. Gular sutures structure unknown; labrum dentate, with two teeth; premolar tooth well-defined; antennae with 11 antennomeres, antennomere 5 much larger than 4 but about as large as 6, antennal club composed from 5 antennomeres; terminal antennomeres lacking sensorial appendages (phaneres); tarsal formula 2-2-2; genital armature of female indistinct.</p><p>Remarks.  Kenotyphlus has 6 species, three in Greece and three in Turkey.</p></div>	https://treatment.plazi.org/id/8E7F8798646EFFF9FF047F2EB0283BB5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047989B66F3D71.text	8E7F8798646EFFF9FF047989B66F3D71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metrotyphlus samothracicus Assing 2019	<div><p>samothracicus Assing, 2019: 892</p><p>( Cyrtotyphlus, TL: Greece: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.690277&amp;materialsCitation.latitude=40.46139" title="Search Plazi for locations around (long 25.690277/lat 40.46139)">Samothráki</a>, ESE Ano Meria, 40°27‘41‘‘N, 25°41‘25‘‘E, 30 m).</p><p>Types: NMW (Assing coll.) HT ♂, 5 PT (NMW (Assing coll.)) .</p><p>Other localities: Greece: Samothráki, Therma.</p><p>Distribution: Greece (Samothráki).</p></div>	https://treatment.plazi.org/id/8E7F8798646EFFF9FF047989B66F3D71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047AD2B6E0381F.text	8E7F8798646EFFF9FF047AD2B6E0381F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Romanotyphlus metaliferensis Struyve 2022	<div><p>metaliferensis Struyve, 2022: 11</p><p>( Romanotyphlus, TL: Romania, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.369&amp;materialsCitation.latitude=46.005" title="Search Plazi for locations around (long 23.369/lat 46.005)">Blandiana</a>, 46.005°N, 23.369°E).</p><p>Types: HNHM HT ♂; 20 PT (HNHM, coll. Struyve)) .</p><p>Distribution: Romania (Apuseni Mountains).</p></div>	https://treatment.plazi.org/id/8E7F8798646EFFF9FF047AD2B6E0381F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF0478E1B7903F3F.text	8E7F8798646EFFF9FF0478E1B7903F3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Romanotyphlus Struyve 2022	<div><p>Romanotyphlus Struyve, 2022</p><p>Romanotyphlus Struyve, 2022: 9 .</p><p>Type species:  Romanotyphlus metaliferensis Struyve, 2022 . Original designation.</p><p>Diagnosis. Gular sutures confluent in midlength of head, from confluent point again separated in two, divergent sutures; labrum edentate; antennae with 11 antennomeres, antennomeres 5 larger than 4 and 6, antennal club with 3 antennomeres; tarsal formula 2-2-2; aedeagus asymmetric, parameres short with one strong, distal seta.</p><p>Remarks. Monospecific genus, recently described from Romania.</p></div>	https://treatment.plazi.org/id/8E7F8798646EFFF9FF0478E1B7903F3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF047A00B1BD3BCA.text	8E7F87986469FFFEFF047A00B1BD3BCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metrotyphlini Coiffait 1963	<div><p>Metrotyphlini Coiffait, 1963</p><p>Diagnosis. Maxillary palpomeres 2 and 3 strongly and about equally dilated; mandibles lacking premolar tooth; tarsi with 2 or 3 tarsomeres; prosternal coxal cavities rounded on anterior margin, lacking excision; lacking suture separating proepisternum from prosternum; all visible tergites with well-defined paratergites; all visible abdominal sternites lacking deep basal transverse sulcus.</p><p>Remarks. The tribe  Metrotyphlini has actually, together with newly described genus, nine genera and 41 valid species names. Six genera are known in the Palaearctic region and two, monospecific genera, occur in South America,  Apotyphlus Coiffait &amp; Saiz, 1965 in Chile and  Venezillotyphlus Decu, 1990 in Venezuela. Un undescribed taxon is reported from Mexico (Navarrete-Heredia et al. 2002). The following key separates  Metrotyphlini of the Palaearctic region.</p><p>1 All legs with three tarsomeres........................................................................... 2</p><p>- All legs with two tarsomeres............................................................................ 7</p><p>2 Antenommere 5 larger than antennomeres 4 and 6........................................................... 3</p><p>- Antenommere 5 not larger than antennomeres 4 and 6........................................................ 5</p><p>3 Antennal club composed of 3 antennomeres................................................  Gynotyphlus Coiffait</p><p>- Antennal club composed of 5 antennomeres................................................................ 4</p><p>4 Labrum edentate, in middle slightly concave; last 4 antennomeres with large sensorial appendages (phaneres) inserted on upper part and curved towards lower part of antennomeres; aedeagus robust, asymmetric, parameres fused to median lobe, with four strong, distal setae....................................................................  Egeotyphlus Coiffait</p><p>- Labrum dentate; last 4 antennomeres with or without sensorial appendages (phaneres) but those if present always inserted on lower part and oriented towards upper part of antennomeres; aedeagus slender, symmetric, lacking parameres................................................................................................  Epirotyphlus Pace</p><p>5 Gular sutures close but not confluent, clearly separated.......................................  Cretotyphlus Assing</p><p>- Gular sutures confluent in about midlength of head or more anteriorly........................................... 6</p><p>6 Gular sutures confluent in about midlenght of head, from confluent point fused in one long straight suture; labrum tridentate; mandible lacking molar tooth.....................................................  Rhopalotyphlus Scheerpeltz</p><p>- Gular sutures confluent in about anterior third of head, from confluent point separated again in two divergent sutures; labrum with one, small median tooth; mandible with well-defined molar tooth...........................  Banatotyphlus Decu</p><p>7 Aedeagus with median lobe falciform; parameres very short, at most reaching half of length of median lobe; genital armature of females distinct.............................................................  Metrotyphlus (s.str.) Coiffait</p><p>- Aedeagus with median lobe stout with complicated distal part bearing some apophysis; parameres longer, almost reaching distal part of the median lobe; genital armature of females indistinct..............  Metrotyphlus (Agnotyphlus) Coiffait</p></div>	https://treatment.plazi.org/id/8E7F87986469FFFEFF047A00B1BD3BCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF04781CB62A3DB9.text	8E7F87986469FFFEFF04781CB62A3DB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kenotyphlus creticus Assing 2018	<div><p>creticus Assing, 2018: 12</p><p>( Kenotyphlus, TL: Greece: Crete, SW Malia, Gonies env., 35°14’43’’N, 25°25’34’’E, 220 m).</p><p>Types: NMW (Assing coll.) HT ♂, 6 PT (NMW (Assing coll.))</p><p>Distribution: Greece (Crete).</p></div>	https://treatment.plazi.org/id/8E7F87986469FFFEFF04781CB62A3DB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF04788CB6353DC5.text	8E7F87986469FFFEFF04788CB6353DC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kenotyphlus rhodiensis Coiffait 1973	<div><p>rhodiensis Coiffait, 1973: 222</p><p>( Kenotyphlus, TL: Ile de Rhodes, Petaloudes).</p><p>Types: MHNG HT ♂, 11PT (MHNG, MNHN)</p><p>Distribution: Greece (Ródos).</p></div>	https://treatment.plazi.org/id/8E7F87986469FFFEFF04788CB6353DC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF0478E0B62A3EFD.text	8E7F87986469FFFEFF0478E0B62A3EFD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kenotyphlus virgatus Assing 2019	<div><p>virgatus Assing, 2019: 262</p><p>( Kenotyphlus, TL: Greece: Crete, S Kissamos, NE Sirikari, 35°24’44”N, 23°38’36”E, 500 m).</p><p>Types: NMW (Assing coll.) HT ♂, 33 PT (NMW (Assing coll.), ZMHB) .</p><p>Other localities: Greece: Crete: S Kissamos, NE Elos, 460 m; S Kissamos, Aikirgianis, 290 m; SW Chania, Mavros, 380 m;   Asigonia, 380 m.</p><p>Distribution: Greece (Crete).</p></div>	https://treatment.plazi.org/id/8E7F87986469FFFEFF0478E0B62A3EFD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047C08B5EA3949.text	8E7F87986468FFFFFF047C08B5EA3949.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Banatotyphlus albensis Struyve 2022	<div><p>albensis Struyve, 2022: 16</p><p>( Banatotyphlus, TL: Romania, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.369&amp;materialsCitation.latitude=46.005" title="Search Plazi for locations around (long 23.369/lat 46.005)">Blandiana</a>, 46.005°N, 23.369°E). Types: HNHM HT ♂; 2 PT (coll. Struyve).</p><p>Distribution: Romania.</p></div>	https://treatment.plazi.org/id/8E7F87986468FFFFFF047C08B5EA3949	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047988B6E73E0D.text	8E7F87986468FFFFFF047988B6E73E0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Banatotyphlus Decu 1982	<div><p>Banatotyphlus Decu, 1982</p><p>Banatotyphlus Decu, 1982: 63 .</p><p>Type species:  Banatotyphlus racovitzai Decu, 1982 . Designated by monotypy.</p><p>Diagnosis. Gular sutures confluent in anterior third of head length, from confluent point again separated in two, divergent sutures; labrum with small, median tooth; antennae with 11 antennomeres, antennomeres 5 about as large as 4 and 6, antennal club with 5 antennomeres; tarsal formula 3-3-3; ventral part of tarsomere 2 and 3 with spatulate setae; aedeagus asymmetric, parameres completely reduced, invisible.</p><p>Remarks. The genus is separated from other  Metrotyphlini by palpomere 2 and 3 subequal in width; antennae short, antennomeres 3–10 transverse, 3–6 subequal in size, antennomeres 9–11 with sensorial appendages (phaneres); tarsi with 3 tarsomeres. The genus was erected for one species,  Banatotyphlus racovitzai Decu 1982 and another three species were described later (Struyve 2022).</p></div>	https://treatment.plazi.org/id/8E7F87986468FFFFFF047988B6E73E0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047C9CB5EA39D5.text	8E7F87986468FFFFFF047C9CB5EA39D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Banatotyphlus orsovae Struyve 2022	<div><p>orsovae Struyve, 2022: 15</p><p>( Banatotyphlus, TL: Romania, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.355&amp;materialsCitation.latitude=44.738" title="Search Plazi for locations around (long 22.355/lat 44.738)">OrŞova</a>, 44.738°N, 22.355°E). Types: HNHM HT ♂; 5 PT (coll. Struyve).</p><p>Distribution: Romania.</p></div>	https://treatment.plazi.org/id/8E7F87986468FFFFFF047C9CB5EA39D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047CF0B5EA3AA9.text	8E7F87986468FFFFFF047CF0B5EA3AA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Banatotyphlus racovitzai Decu 1982	<div><p>racovitzai Decu, 1982: 64</p><p>( Banatotyphlus, TL: vallée d’OgaŞu Gǎurii (Monts Locvei)).</p><p>Types: HT ♂, deposition unknown.</p><p>Other localities: Romania, Şopotu Nou, 44.809°N, 21.842°E .</p><p>References: Struyve 2022: 15.</p><p>Distribution: Romania.</p></div>	https://treatment.plazi.org/id/8E7F87986468FFFFFF047CF0B5EA3AA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047FBCB6003B7D.text	8E7F87986468FFFFFF047FBCB6003B7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Banatotyphlus siclus Struyve 2022	<div><p>siclus Struyve, 2022: 15</p><p>( Banatotyphlus, TL: Eibenthal, 44.546°N, 22.205°E).</p><p>Types: HNHM HT ♂; 15 PT (HNHM, coll. Struyve) .</p><p>Other localities: Cozia (sic!) [= Cozla], 44.631°N, 22.007°E and 44.628°N, 22.039°E;</p><p>New records: Serbia or., 14.x.2022 and 26.v.2024, Mt. Stol, 660m, 44.16446N, 22.11492E.</p><p>Distribution: Romania, Serbia.</p></div>	https://treatment.plazi.org/id/8E7F87986468FFFFFF047FBCB6003B7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047B90B4D03869.text	8E7F87986468FFFFFF047B90B4D03869.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Banatotyphlus siclus Struyve 2022	<div><p>Banatotyphlus siclus Struyve, 2022</p><p>Figs 19–24, 83–84</p><p>Material examined.   SERBIA: 1♂, 1♀: with one label “SERBIA or. 14.x.2022 / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.11492&amp;materialsCitation.latitude=44.16446" title="Search Plazi for locations around (long 22.11492/lat 44.16446)">Mt. Stol</a>, soil washing, 660m / 44.16446N, 22.11492E / J. Janák lgt., hornbeam forest” [white, printed] (JJRC)  .  16♂♂, 6♀♀: same locality label but collected on 26.v.2024 (JJRC, MMBC) .</p><p>Supplement description. Visible tergite 6 (IX) of male elongate with rounded posterior margin (Fig. 19). Visible sternite 6 (VIII) of male with deep emargination in posterior third (Fig. 20), male tergites IX and X as in Fig. 21, male sternite IX apically rounded (Fig. 22). Aedeagus (Figs 23–24) dorsaly elongate, slender, four times as long as wide, lateraly with apophysis in shape of “S”. First record for Serbia.</p><p>Natural History. The same as for  Allotyphlus stolensis sp. nov. (Figs 83–84).</p><p>Distribution. Romania, Serbia (Western Carpathians, on both side of the border between Romania and Serbia).</p></div>	https://treatment.plazi.org/id/8E7F87986468FFFFFF047B90B4D03869	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047988B4C93E24.text	8E7F8798646BFFFCFF047988B4C93E24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cretotyphlus Assing 2019	<div><p>Cretotyphlus Assing, 2019</p><p>Cretotyphlus Assing, 2019: 263 .</p><p>Type species:  Cretotyphlus hamatus Assing, 2019 . Original designation.</p><p>Diagnosis. Mandibles apically bifid; gular sutures close, but not confluent anteriorly; labrum tridentate, maxillary palpi with palpomeres 2 and 3 enlarged and somewhat flattened, 3 larger than 2; antenna with 11 antennomeres, moderately incrassate, apical antennomeres lacking sensorial appendages, antennal club composed of three last antennomeres; tarsi with 3 tarsommeres; aedeagus with long, slender, and distally acute ventral process, lacking parameres; female genital segment with more or less distinct pair of sclerites.</p><p>Remarks. The genus was erected for three new species from Crete and later another species was discovered in Sámos.</p></div>	https://treatment.plazi.org/id/8E7F8798646BFFFCFF047988B4C93E24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047BCEB62A3F68.text	8E7F8798646BFFFCFF047BCEB62A3F68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cretotyphlus chanianus Assing 2019	<div><p>chanianus Assing, 2019: 264</p><p>( Cretotyphlus, TL: Greece: Crete, SW Chania, Mavros stream valley, 35°24‘10“N, 23°53‘53“E, 290 m).</p><p>Types: NMW (Assing coll.) HT ♂, 2 PT (NMW (Assing coll.)) .</p><p>Distribution: Greece (Crete).</p></div>	https://treatment.plazi.org/id/8E7F8798646BFFFCFF047BCEB62A3F68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047A7EB62A3FD8.text	8E7F8798646BFFFCFF047A7EB62A3FD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cretotyphlus hamatus Assing 2019	<div><p>hamatus Assing, 2019: 263</p><p>( Cretotyphlus, TL: Greece: Crete, NE Lassithi Plateau, 35°12‘45“N, 25°31‘53“E, 850 m).</p><p>Types: NMW (Assing coll.) HT ♂, 3 PT (NMW (Assing coll.)) .</p><p>Distribution: Greece (Crete).</p></div>	https://treatment.plazi.org/id/8E7F8798646BFFFCFF047A7EB62A3FD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047AEEB62A3848.text	8E7F8798646BFFFCFF047AEEB62A3848.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cretotyphlus idanus Assing 2019	<div><p>idanus Assing, 2019: 264</p><p>( Cretotyphlus, TL: Greece: Crete, Psiloritis NE Livadia, 35°18‘56“N, 24°49‘07“E, 440 m).</p><p>Types: NMW (Assing coll.) HT ♂, 2 PT (NMW (Assing coll.)) .</p><p>Distribution: Greece (Crete).</p></div>	https://treatment.plazi.org/id/8E7F8798646BFFFCFF047AEEB62A3848	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047D9EB6383938.text	8E7F8798646BFFFCFF047D9EB6383938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cretotyphlus kerkisicus Assing 2022	<div><p>kerkisicus Assing, 2022: 233</p><p>( Cretotyphlus, TL: Greece: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.657778&amp;materialsCitation.latitude=37.7425" title="Search Plazi for locations around (long 26.657778/lat 37.7425)">Samos</a>, Oros Kerkis, NE-slope, 37°44‘33“N, 26°39‘28“E, 740 m).</p><p>Types: NMW (Assing coll.) HT ♂ .</p><p>Distribution: Greece (Sámos).</p></div>	https://treatment.plazi.org/id/8E7F8798646BFFFCFF047D9EB6383938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047C46B13D3B1D.text	8E7F8798646BFFFCFF047C46B13D3B1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Egeotyphlus Coiffait 1957	<div><p>Egeotyphlus Coiffait, 1957</p><p>Egeotyphlus Coiffait, 1957: 80 .</p><p>Type species:  Egeotyphlus thracicus Coiffait, 1957 . Original designation.</p><p>Egeotyphlus Coiffait: Coiffait, 1957: 61 (in key), 80 (diagnosis); Coiffait, 1959: 388 (redescription); Coiffait, 1972: 614 (redescription); Pace, 1983a: 460 (diagnosis in key); Pace, 1984a: 17 ( E. thracicus in key); Pace, 1995: 29 (diagnosis in key).</p><p>Diagnosis. Gular sutures confluent in anterior third of length of head capsule, from confluent point prolonged ahead as one straight suture; labrum edentate, its anterior margin in middle concave; antennae short, antennae with 11 antennomeres, all antennomeres transverse, 5 clearly larger than 4 and 6, antennomeres 8–11 with sensorial appendages (phaneres), inserted on upper part and curved towards lower part of antennomeres; antennal club composed from 5 antennomeres; all tarsi with 3 tarsomeres.</p><p>Remarks. The diagnosis of  Egeotyphlus was updated by Coiffait (1972).After transferring  Egeotyphlus zecevici to the genus  Epirotyphlus, the genus  Egeotyphlus became monospecific (see remarks for  Epirotyphlus).</p></div>	https://treatment.plazi.org/id/8E7F8798646BFFFCFF047C46B13D3B1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFFDFF04781CB7093D55.text	8E7F8798646AFFFDFF04781CB7093D55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Egeotyphlus thracicus Coiffait 1957	<div><p>thracicus Coiffait, 1957: 81</p><p>( Egeotyphlus, TL: Thrace grecque: Makri [Mákri]).</p><p>Types: depository unknown.</p><p>Distribution: Greece (Eastern Macedonia and Thrace).</p></div>	https://treatment.plazi.org/id/8E7F8798646AFFFDFF04781CB7093D55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFFDFF0478A8B5F3395E.text	8E7F8798646AFFFDFF0478A8B5F3395E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epirotyphlus Pace 1995	<div><p>Epirotyphlus Pace, 1995,  new status</p><p>Epirotyphlus Pace, 1995: 27 .</p><p>Type species:  Egeotyphlus leucadiensis Pace, 1995 . Original designation. Described as a subgenus of the genus  Egeotyphlus . New status.</p><p>Diagnosis. Gular sutures confluent in about midlength of head capsule, from this confluent point they continue as two, separate, short divergent sutures (Figs 31, 61); labrum dentate (Figs 30, 59); antennae with 11 antennomeres, antennomere 5 larger than 4 and 6; antennal club composed from 5 antennomeres (Figs. 32, 62), last 4 antennomeres with (Fig. 63) or without sensorial appendages (phaneres) but those if present always inserted on lower part and oriented towards upper part of antennomeres.</p><p>Remarks.  Epirotyphlus was originally described as a subgenus of  Egeotyphlus for a new species,  Egeotyphlus (Epirotyphlus) leucadiensis from the Greek island Lefkáda.  Epirotyphlus was separated from the nominal subgenus by 1) lacking large sensorial appendages (phaneres) on the last four antennomeres inserted on upper part and curved towards lower part, 2) having dentate labrum and 3) aedeagus lacking parameres. Pace (1995), when describing  Egeotyphlus (Epirotyphlus) leucadiensis recognized that this species is more similar to his previously described species  Egeotyphlus zecevici from Serbia (Pace 1984b) than to the type species of the genus,  Egeotyphlus thracicus . These both species share all characters mentioned above and more, they have very similar structure of the aedeagus. Aedeagus is slender, symmetric, parameres are reduced, lacking distal setae. The discovery of two new species, which are described here below and assigned to  Epirotyphlus, allowed us to study the ventral part of head and it seems that the structure of gular sutures can be an important character to separate  Epirotyphlus from  Egeotyphlus . Both new species, have gular sutures confluent in about midlength of head capsule and from this confluent point they continue as two, separate, short divergent sutures, opposite to  Egeotyphlus, where they continue as a simple, straight suture. Unfortunately, Pace (1984b, 1995) did not provide any information about the structure of gular sutures for either  Egeotyphlus (Epirotyphlus) leucadiensis or  Egeotyphlus zecevici . Thus, the importance of the structure of gular sutures for the definition of  Epirotyphlus will need further study, especially when the type species,  Epirotyphlus leucadiensis will be available. Regardless this, we are of the opinion that  Epirotyphlus deserves to be upgraded to the genus level.</p></div>	https://treatment.plazi.org/id/8E7F8798646AFFFDFF0478A8B5F3395E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFE6FF047CDFB5D83EC1.text	8E7F8798646AFFE6FF047CDFB5D83EC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epirotyphlus dusicae Hlavac, Janak & Banar 2025	<div><p>Epirotyphlus dusicae Hlaváč, Janák &amp; Baňař,  sp. nov.</p><p>Figs 25–54, 85</p><p>Material studied.   Holotype, ♂: MONTENEGRO with one label “MONTENEGRO SW013, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.97889&amp;materialsCitation.latitude=42.19889" title="Search Plazi for locations around (long 18.97889/lat 42.19889)">Đurovići</a> / 42°11’56’’N, 18°58’44’’E / 60 m, 23.III.2024, soil / washing near oak tree, P. Baňař lgt” [white, printed] (MMBC)  .  Paratypes: 7♂, 7♀, same data as holotype: (3♂, 4♀ in MMBC, 4♂, 3♀ in JJRC) .   4♂, 5♀: “MONTENEGRO SW009, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.97889&amp;materialsCitation.latitude=42.19889" title="Search Plazi for locations around (long 18.97889/lat 42.19889)">Đurovići</a>, 2.v.2023, 42°11‘56‘‘N, 18°58‘44‘‘E, 60 m, soil washing near oak tree, P. Baňař &amp; P.Hlaváč lgt.” [white, printed] (BMNH, MMBC, JJRC)  .   1♂: MONTENEGRO SW002, Rumija, road from Bar → <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.159721&amp;materialsCitation.latitude=42.116943" title="Search Plazi for locations around (long 19.159721/lat 42.116943)">Mikulići</a>, 30.iv.2023, 42°07‘01‘‘N, 19°09‘35‘‘E, 675 m, soil washing near oak tree, P. Baňař lgt.” [white, printed] (MMBC).   5♀: MONTENEGRO SW010, Rumija, road from Bar → Mikulići, 21.iii.2024, 42°07‘01‘‘N, 19°09‘35‘‘E, 674 m, soil washing near  Carpinus tree, P. Baňař lgt.” [white, printed] (MMBC)  .   2♂, 2♀, “MONTENEGRO SW014, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.979445&amp;materialsCitation.latitude=42.198612" title="Search Plazi for locations around (long 18.979445/lat 42.198612)">Đurovići</a> / 42°11‘55‘‘N, 18°58‘46‘‘E / 58 m, 23.III.2024, soil / washing close stream, P. Baňař lgt” [white, printed] (MMBC)  .   1♂, 2♀: “MONTENEGRO SW006, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.980278&amp;materialsCitation.latitude=42.19917" title="Search Plazi for locations around (long 18.980278/lat 42.19917)">Đurovići</a>, 2.v.2023, 42°11‘57‘‘N, 18°58‘49‘‘E, 70 m, soil washing near olive tree, P. Baňař lgt.” [white, printed] (MMBC)  .   2♀: MONTENEGRO SW007, near  Blizikuće, near petrol station, 29.iv.2023, soil washing near  Juniperus tree, P. Baňař &amp; P. Hlaváč lgt. [white, printed]” (MMBC)  .</p><p>Diagnosis. Head longer than wide (Fig. 28), longer than pronotum; maxillary palpi with palpomeres 2 and 3 dilated, 3 larger than 2 (Fig. 29, 45); ligula indistinct (Fig. 44); mandibles falciform (Fig. 45); labrum dentate (Fig. 30); gular suture confluent anteriorly in about midlength of head capsule (Fig. 31). Antennae with 11 antennomeres, antennal club composed of 5 antennomeres, with sensorial appendages which are inserted on lower part and oriented towards upper part of antennomere, antennomeres 3–10 strongly transverse (Figs 32, 49). Elytra tetragonal, dilated posteriad (Fig. 33), shorter than pronotum. Procoxal cavities (Figs 34, 50) rounded on anterior margin, lacking excision, prosternum and proepisternum fused.Abdominal sternites lacking deep, transversal sulcus, visible sternite 2–5 with shallow basal impression (Figs 27, 46). Tarsal formula 3-3-3, tarsomere 1 minuscule, 2 slightly longer, 3 longest, clearly longer than 1 and 2 combined (Figs 38–40, 52). Aedeagus elongate, slender and nearly symmetrical (Fig. 41, 42); lacking parameres, lateral protrusions of median lobe long (Fig. 42d, arrows). Terminal abdominal segments of female (Fig. 37) with one paired longitudinal sclerite.</p><p>Description. Body (Figs 25, 26, 46, 48) depigmented, most part of surface shagreened (Figs 43, 47, 54), body length about 0.95 mm.</p><p>Head capsule (Fig. 28, 43) wider than long, about 1.2 times as wide as long, shorter than pronotum; maxillary palpi (Fig. 29, 45) with palpomeres 2 and 3 dilated, 3 larger than 2; ligula indistinct (Fig. 29, 44); mandibles falciform (Fig. 45); labrum dentate (Fig. 30); gular suture confluent anteriorly behind midlength of head capsule, after meeting point again separated to two divergent arms (Fig. 31).</p><p>Antennae (Fig. 32, 49) with 11 antennomeres, scape bulbous, larger than pedicel, pedicel dilated distad, distally truncate, antennomeres 3–10 strongly transverse, antennomere 5 larger than 4 and 6, antennal club composed of 5 antennomeres, with sensorial appendages which are inserted on lower part and oriented towards upper part of antennomere.</p><p>Pronotum (Fig. 33, 43) wider than long, convergent posteriad.</p><p>Elytra (Fig. 33, 47, el) tetragonal, dilated posteriad, shorter than pronotum, external posterior angles rounded, internal ones right-angled.</p><p>Venter, procoxal cavities (Fig. 34, 50) rounded on anterior margin, anterior margin with inconspicuous excision (50, arrow), lacking suture separating proepisternum from prosternum, lateral portions of prosternum shagreened, medium portion smooth.</p><p>Abdominal sternites (Fig 27, 46, 48) lacking deep, transversal sulcus, visible sternite 2–6 with shallow basal impression (Fig. 27).</p><p>Legs (Figs 38–40, 52, 53) short, all tibiae dilated distad, tarsal formula 3-3-3, tarsomere about half of length of 3, tarsomere 2 shortest, about 3 times shorter than 1.</p><p>Aedeagus (Figs 41, 42) about 0.17 mm long, median lobe with long lateral protrusions (Fig. 42d, arrows).</p><p>Terminal abdominal segments of female (Fig. 37) with one paired longitudinal sclerite.</p><p>Etymology. Named after our friend Dušica Šoljaga, owner of a beautiful house which is a base of our expeditions to the coastal part of Montenegro.</p><p>Natural History. The majority of specimens (holotype and 27 paratypes) of newly described species was collected by the soil washing of a sample taken near and around a solitary oak (Fig. 85) on the border of terrace garden together with  Ubychia ganglbaueri (Reitter, 1912) and an undesribed species of  Cephennium and  Tychobythinus in May 2023 and March 2024. Several specimens were collected in Rumija Mts. in about 675 m a.s.l. and two females also in coastal Mediterranean forest under junipers tree near Blizikuće village.</p><p>Distribution. Montenegro.</p></div>	https://treatment.plazi.org/id/8E7F8798646AFFE6FF047CDFB5D83EC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986471FFE5FF047A5DB5973BE7.text	8E7F87986471FFE5FF047A5DB5973BE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epirotyphlus janae Hlavac, Janak & Banar 2025	<div><p>Epirotyphlus janae Hlaváč, Janák &amp; Baňař,  sp. nov.</p><p>Figs 55–76, 86–87</p><p>Material studied.  Holotype, ♂: SERBIA with one label “SERBIA or. 22.v.2024 / Majdan Kučajna, 230m / 44.44561 N, 21.63905 E / soil washing, J. Janák lgt. / hornbeam forest“ [white, printed] (JJRC) .  Paratypes: 2♂♂, 3♀♀ same data as for holotype (1♂, 3♀♀ in JJRC, 1♂ in MMBC) .</p><p>Diagnosis. Gular sutures confluent anteriorly slightly behind midlength of head capsule (Fig. 61); antennae long, antennomeres 3–10 transverse; antennomeres 5 larger than 4 and 6; antennal club composed from five antennomeres (Figs 62–63), incrassate, last four antennomeres setose, with sensorial appendages which are inserted on lower part and oriented towards upper part of antennomere (Fig. 63); procoxal cavities rounded on anterior margin, with small excision on lateral part (Fig. 65, arrow); tergites 1–5 (IV–VIII) and visible sternites 2–4 (IV–VI) (Fig. 57) lacking basal impression (Fig. 57); tarsal formula 3-3-3 (Figs 71–73); aedeagus elongate, about 3 times as long as wide; distal, ventral part of median lobe with deep excision, lateral arms truncate; lacking parameres, lateral protrusions of median lobe short (Figs 74, 75). Terminal abdominal segments of female (Fig. 76) with one paired longitudinal sclerite and large asymmetric structure in middle.</p><p>Description. Body (Figs 55, 56) about 0.9 mm long, completely depigmented.</p><p>Head capsule (Fig. 58) about 1.2 wider than long; about as wide and as long as pronotum; head capsule separated from neck region by occipital constriction, neck region retracted into pronotum; gular sutures (Fig. 61) confluent anteriorly slightly behind midlength of head capsule; maxillary palpi (Fig. 60) with palpomere 2 dilated, about as wide as 3 and about half of length of 3; palpomere 4 very small.</p><p>Antennae (Figs 62, 63) long, scape about 1.45 times as long as pedicel, about 1.2 times as long as wide; pedicel as long as wide, 3.5 times as long as antennomere 3; antennomeres 3–10 transverse; antennomeres 5 larger than 4 and 6; antennal club composed from five antennomeres, incrassate, 10 slightly wider than long, elliptical, terminal antennomere subspherical. Last four antennomeres (Fig. 63) setose, with sensorial appendages which are inserted on lower part and oriented towards upper part of antennomere.</p><p>Pronotum (Fig. 64) about 1.1 times as wide as long, widest in anterior third, after that convergent posteriad; anterior angles widely rounded; posterior angles sharper; about 1.1 times as long as elytra.</p><p>Venter, prosternum (Fig. 65) fused with proepisternum, procoxal cavities rounded on anterior margin, with small excision (Fig. 65, arrow) on lateral part.</p><p>Elytra (Fig. 64) about as long as wide, widest on posterior margin.</p><p>Abdomen long, considerably longer than length of head capsule, pronotum and elytra combined. Visible tergites 1–5 (IV–VIII) and visible sternites 2–4 (IV–VI) (Fig. 57) lacking basal impression. Visible sternite 6 (IX) of male (Figs 67, 68) symmetric, visible sternite 7 (IX) in male (Fig. 70) with small setae; sparsely setose in posterior half, visible tergite 7 (X) with median posterior excision which is prolonged with wide, deep impression (Fig. 69). Visible tergite 5 (VIII) (Fig. 66) 1.25 times as long as wide.</p><p>Legs (Figs 71–73) short, all tibia distally dilated, tarsal formula 3-3-3; first tarsomere longer than 2, 3 longer than 1 and 2 combined.</p><p>Aedeagus (Figs 74, 75) about 0.17 mm long, elongate, about 3 times as long as wide; distal, ventral part of median lobe with deep excision, lateral arms truncate; basal part less than half of length of median lobe; with elongate, long, internal apophysis; lacking parameres, lateral protrusions of median lobe short. Terminal abdominal segments of female (Fig. 76) with one paired longitudinal sclerite and large asymmetric structure in middle.</p><p>Etymology. Patronymic, named after Jana Janáková, the wife of the second author to thank her for being a companion on trips to the Balkan countries.</p><p>Remarks.  E. janae has the same type of the aedeagus as  E. zecevici but it differs from latter mainly by apically narrowed lateral projections and truncate lateral arms of median lobe (Pace 1984, Figs 2–4).</p><p>Natural History. All specimens were collected by soil-washing on a south-facing slope above the road at the edge of a deciduous forest (hornbeam, hazel, maple, oak). Volume of sample was about 100 liters (Figs 86–87).</p><p>Distribution. Serbia.</p></div>	https://treatment.plazi.org/id/8E7F87986471FFE5FF047A5DB5973BE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFEBFF047F61B58D3AD0.text	8E7F8798647CFFEBFF047F61B58D3AD0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Egeotyphlus zecevici Pace 1984	<div><p>zecevici Pace, 1984b: 17</p><p>( Egeotyphlus, TL: Serbia, Stol planina, 800 m), new combination.</p><p>Types: 4 ST (coll. Nonveiller, coll. Pace).</p><p>Distribution: Serbia.</p></div>	https://treatment.plazi.org/id/8E7F8798647CFFEBFF047F61B58D3AD0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFEBFF047C9EB5CE39F0.text	8E7F8798647CFFEBFF047C9EB5CE39F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epirotyphlus dusicae Hlaváč & Janák & Baňař 2025	<div><p>dusicae sp. nov.</p><p>Distribution: Montenegro.</p></div>	https://treatment.plazi.org/id/8E7F8798647CFFEBFF047C9EB5CE39F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFEBFF047F29B58D3A44.text	8E7F8798647CFFEBFF047F29B58D3A44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epirotyphlus janae Hlaváč & Janák & Baňař 2025	<div><p>janae sp. nov.</p><p>Distribution: Serbia.</p></div>	https://treatment.plazi.org/id/8E7F8798647CFFEBFF047F29B58D3A44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFEBFF047CD6B60A3A1C.text	8E7F8798647CFFEBFF047CD6B60A3A1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epirotyphlus leucadiensis Pace 1995	<div><p>leucadiensis Pace, 1995: 27</p><p>( Egeotyphlus, TL: Isola di Leukas,  Apolpena (Grecia)), new combination.</p><p>Types: NMW (Assing coll.) HT ♂.</p><p>Distribution: Greece (Lefkáda).</p></div>	https://treatment.plazi.org/id/8E7F8798647CFFEBFF047CD6B60A3A1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFE9FF047E38B7853E59.text	8E7F8798647CFFE9FF047E38B7853E59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gynotyphlus Coiffait 1955	<div><p>Gynotyphlus Coiffait, 1955</p><p>Gynotyphlus Coiffait, 1955: 72 .</p><p>Type species:  Leptotyphlus perpusillus Dodero, 1900 . Original designation.</p><p>Oligotyphlus Scheerpeltz, 1940: 28 nomen nudum, no species designation.</p><p>syn.  Oligotyphlus Scheerpeltz, 1956: 460 . Type species:  Leptotyphlus perpusillus Dodero, 1900 . Validation of  Oligotyphlus Scheerpeltz 1940 by inclusion of species and designation. Synonymy in Coiffait, 1959: 390.</p><p>syn.  Austriacotyphlus Scheerpeltz, 1959: 4 . Type species:  Austriacotyphlus piffli Scheerpeltz, 1959 . Original designation. Synonymy in Pace, 1983b: 43.</p><p>Gynotyphlus Coiffait: Coiffait, 1957: 61 (in key); Coiffait, 1959: 390 (redescription, key to subspecies of  G. perpusillus); Coiffait, 1964: 696 (distribution); Coiffait, 1972: 617 (redescription, key to subspecies of  G. perpusillus); Coiffait, 1984: 117 (distribution); Pace, 1996: 302 (redescription, key to subspecies of  G. perpusillus of Italy).</p><p>Austriacotyphlus Scheerpeltz: Coiffait, 1972: 616 (redescription).</p><p>Diagnosis. Gular sutures confluent in anterior third of length of head capsule, from confluent point not prolonged ahead; labrum dentate; antennae short, with 11 antennomeres, antennomeres 5–10 strongly transverse, 5 wider than 4 and about as wide as 6, antennal club composed from 5 antennomeres, terminal antennomeres lacking sensorial appendages (phaneres); all tarsi with 3 tarsomeres. Genital armature of female indistinct.</p><p>Remarks. The genus has three species, all are present in the Balkan Peninsula. It is believed that the natural area of the distribution is southern France, Italy, the Balkans and Turkey and that the parthenogenetic species  Gynotyphlus perpusillus was introduced in many countries in Europe.</p></div>	https://treatment.plazi.org/id/8E7F8798647CFFE9FF047E38B7853E59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFE9FF047D91B02139F8.text	8E7F8798647EFFE9FF047D91B02139F8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genotyphlus perpusillus Dodero 1900	<div><p>perpusillus Dodero, 1900b: 573</p><p>( Leptotyphlus; TL: Genova).</p><p>Remarks: Eleven subpecies of  Gynotyphlus perpusillus were described (Löbl &amp; Löbl 2015: 915), two from the territory of the Balkans,  Gynotyphlus perpusillus dalmaticus Coiffait, 1955: 72 (TL: Dalmatie: Susurac [Sućurac] from southern Dalmatia, Croatia) and  Gynotyphlus perpusillus macedonicus Coiffait, 1957: 81 (TL: Macédoine Yugoslave [North Macedonia]: Katlanovo). The validity of all ten subspecies is doubtfull and it will need the revison (Struyve 2022: 7).</p><p>Reference: Struyve 2022: 7 (distribution in Romania, note on the validity of subspecies).</p><p>Distribution: Portugal, France, Corsica, Italy, Sardinia, Sicily, Switzerland, Austria, Greece, Ionian Islands, Croatia, North Macedonia, Turkey, Morocco, Romania (Struyve 2022: 7).</p></div>	https://treatment.plazi.org/id/8E7F8798647EFFE9FF047D91B02139F8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFE9FF047A39B6373FE0.text	8E7F8798647EFFE9FF047A39B6373FE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gynotyphlus beierianus Scheerpeltz 1958	<div><p>beierianus Scheerpeltz, 1958: 396</p><p>( Oligotyphlus, TL: südlichen Epiros, Umgebung von Nisista, Xerovuni, 700–800 m).</p><p>Types: NHM HT ♀, 6PT ♀ (NMW).</p><p>Reference: Violi &amp; Fancello, 2022: 1 (the necessity to verify the validity of the species).</p><p>Distribution: Greece (Epirus).</p></div>	https://treatment.plazi.org/id/8E7F8798647EFFE9FF047A39B6373FE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFE9FF047AC5B60338B4.text	8E7F8798647EFFE9FF047AC5B60338B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gynotyphlus corcyrensis Scheerpeltz 1958	<div><p>corcyrensis Scheerpeltz, 1958: 394</p><p>( Oligotyphlus, TL: Gasturi aus Korfu).</p><p>Types: 6ST (NMW) .</p><p>Other localities: Val de Ropa auf Korfu.</p><p>Reference: Violi &amp; Fancello, 2022: 1 (the necessity to verify the validity of the species).</p><p>Distribution: Greece (Kérkira).</p></div>	https://treatment.plazi.org/id/8E7F8798647EFFE9FF047AC5B60338B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFEEFF047F05B5053913.text	8E7F8798647EFFEEFF047F05B5053913.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metrotyphlus Coiffait 1959	<div><p>Metrotyphlus Coiffait, 1959</p><p>Metrotyphlus as a subgenus of  Megatyphlus Coiffait, 1959: 328 .</p><p>Type species:  Megatyphlus hervei Coiffait, 1957 . Original designation.</p><p>Metrotyphlus Coiffait: Coiffait, 1972: 609 (redescription, key to subgenera and species); Pace, 1996: 286 (redescription, key to subgenera and species of Italy).</p><p>sbg.  Agnotyphlus Coiffait, 1972: 613 . Type species:  Metrotyphlus istriensis Coiffait, 1972 . Original designation.</p><p>sbg.  Agnatotyphlus: Struyve, 2022: 13 (lapsus calami).</p><p>Diagnosis. Gular sutures confluent in anterior third of length of head capsule, from confluent point prolonged ahead by two short, divergent sutures; labrum dentate; antennae with 11 antennomeres, 5 slightly larger than 4 and 6, antennal club composed from 3 antennomeres; terminal antennomeres lacking sensorial appendages (phaneres); all tarsi with 2 tarsomeres. Genital armature of female distinct or indistinct.</p><p>Remarks.  Metrotyphlus, together with  Apotyphlus and  Venezillotyphlus have tarsal formula 2-2-2. The genus has 2 subgenera differentiated by the structure of aedeagus and female genital armature. All balkanic species belong to the subgenus  Agnotyphlus Coiffait, 1972 . The nominal subgenus includes four species in France, Italy and Switzerland.</p></div>	https://treatment.plazi.org/id/8E7F8798647EFFEEFF047F05B5053913	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986479FFEEFF047CFEB6743A57.text	8E7F87986479FFEEFF047CFEB6743A57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metrotyphlus aristotelis Violi & Fancello 2022	<div><p>aristotelis Violi &amp; Fancello, 2022: 3</p><p>( Metrotyphlus sbg. Agnotyphlus, TL: Grecia —Penisola Calcidica, Galatista: 3,5 Km ad Est di Galatista, 40°27’41.6’’ 23°19’20.7’’, 520 m).</p><p>Types: MCSN HT ♂.</p><p>Distribution: Greece (Khalkidhiki).</p></div>	https://treatment.plazi.org/id/8E7F87986479FFEEFF047CFEB6743A57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986479FFEEFF047F6EB5EA3B2B.text	8E7F87986479FFEEFF047F6EB5EA3B2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metrotyphlus banati Struyve 2022	<div><p>banati Struyve, 2022: 13</p><p>( Metrotyphlus, subg. Agnatotyphlus [sic], TL: Romania, Eibenthal, 44. 546°N, 22.205°E).</p><p>Types: HNHM HT ♂; 13 PT (HNHM, coll. Struyve) .</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.039&amp;materialsCitation.latitude=44.628" title="Search Plazi for locations around (long 22.039/lat 44.628)">Other</a> records: Romania, Cozia (sic!) [= Cozla], 44.628°N, 22.039°E  .</p><p>Distribution: Romania.</p></div>	https://treatment.plazi.org/id/8E7F87986479FFEEFF047F6EB5EA3B2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986479FFEEFF047E3AB62F3BFF.text	8E7F87986479FFEEFF047E3AB62F3BFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metrotyphlus goritianus Pace 1977	<div><p>goritianus Pace, 1977a: 8</p><p>( Metrotyphlus, sbg. Agnotyphlus, TL: monti del Carso di Monfalcone, presso il centro abitato di Devetachi, a circa 215 metri d’altitudine).</p><p>Types: MSNV HT ♂; 13 PT (MSNV, MCST, coll. Pace) .</p><p>Other records: Slovenia, Lokvica, 215m.</p><p>Distribution: Italia, Slovenia.</p></div>	https://treatment.plazi.org/id/8E7F87986479FFEEFF047E3AB62F3BFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986478FFEFFF047989B62A3CC8.text	8E7F87986478FFEFFF047989B62A3CC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metrotyphlus istriensis Coiffait 1972	<div><p>istriensis Coiffait, 1972: 613</p><p>( Metrotyphlus, sbg. Agnotyphlus, TL: Istrie, Monte Maggiore [Učka]).</p><p>Types: SOFM HT ♂, 1PT (SOFM) .</p><p>Distribution: Croatia (Istria).</p></div>	https://treatment.plazi.org/id/8E7F87986478FFEFFF047989B62A3CC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986478FFEFFF04781DB66F3DB8.text	8E7F87986478FFEFFF04781DB66F3DB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metrotyphlus samothracicus Assing 2019	<div><p>samothracicus Assing, 2019: 897</p><p>( Metrotyphlus, TL: Greece: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.601389&amp;materialsCitation.latitude=40.494446" title="Search Plazi for locations around (long 25.601389/lat 40.494446)">Samothráki</a>, Therma, 40°29’40’’N, 25°36’05’’E, 110 m).</p><p>Types: NMW (Assing coll.) HT ♂ .</p><p>Distribution: Greece (Samothráki).</p></div>	https://treatment.plazi.org/id/8E7F87986478FFEFFF04781DB66F3DB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986478FFEFFF047F21B08F3BF9.text	8E7F87986478FFEFFF047F21B08F3BF9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhopalotyphlus Scheerpeltz 1956	<div><p>Rhopalotyphlus Scheerpeltz, 1956</p><p>Rhopalotyphlus Scheerpeltz, 1956: 462 .</p><p>Type species:  Leptotyphlus beieri Scheerpeltz, 1931 . Original designation.</p><p>Validation of  Rhopalotyphlus Scheerpeltz 1940 by inclusion of species and designation of type species.</p><p>Rhopalotyphlus Scheerpeltz: Scheerpeltz, 1940: 29 (nomen nudum); Coiffait, 1972: 625 (redescription); Pace, 1983a: 460 (diagnosis in key).</p><p>Diagnosis. Gular sutures confluent in half of length of head capsule, from confluent point prolonged ahead by long, single suture; labrum dentate, with three prominent teeth; antennae with 11 antennomeres, antennal club composed from 3 antennomeres; terminal antennomeres lacking sensorial appendages (phaneres); tarsal formula 3-3-3; genital armature of female distinct.</p><p>Remarks. Monospecific genus known only from the type material, the male is unknown.</p></div>	https://treatment.plazi.org/id/8E7F87986478FFEFFF047F21B08F3BF9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647BFFECFF04781CB60A3DB9.text	8E7F8798647BFFECFF04781CB60A3DB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotyphlus beieri Scheerpeltz 1931	<div><p>beieri Scheerpeltz, 1931: 394</p><p>( Leptotyphlus, TL: bei Kaligoni auf Levkas).</p><p>Types: 3ST (NMW) .</p><p>References: Pace, 1983a: 459 (supplement description).</p><p>Distribution: Greece (Lefkáda).</p></div>	https://treatment.plazi.org/id/8E7F8798647BFFECFF04781CB60A3DB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647AFFEDFF047DAAB1BD3BA4.text	8E7F8798647AFFEDFF047DAAB1BD3BA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neotyphlini Coiffait 1963	<div><p>Neotyphlini Coiffait, 1963</p><p>Diagnosis. Maxillary palpi with palpomeres 2 and 3 strongly, equally dilated; ligula on anterior margin with two widely well-separated projections; mandibles lacking premolar tooth; coxal cavities with lateral excision on anterior margin which is prolonged by suture separating prosternum from proepisternum; all visible abdominal sternite lacking deep transversal sulcus.</p><p>Remarks. The most widespread tribe of the family with 20 genera and 51 valid species names which is considered to be the most primitive leptotyphlines (Smetana 1986). The tribe is richest in taxa in the New World with 16 genera and 42 species, where Chile with 5 genera and 23 species and California (USA) with 7 genera and 12 species are the richest known regions. One genus with two species is known from South Africa and three genera and seven species from Europe.  Kladotyphlus and  Megatyphlus are present in the Balkan Peninsula.  Apheliotyphlus is monospecific genus known in Italy (Friuli-Venezia Giulia) and because of the vicinity of this region to Slovenia, the presence of the genus in Slovenia can be plausible. All three European genera can be separated by the following key (Pace 1999, translated and modified).</p><p>1 All tarsi with 2 tarsommeres; labrum on anterior margin more or less rectilinear, aedeagus with parameres.............. 2</p><p>- All tarsi with 3 tarsommeres; labrum with anterior margin with deep median excision forming long, lateral, divergent branches; aedeagus lacking parameres..............................................................  Kladotyphlus Pace</p><p>2 Head capsule lacking sulcus in shape of V; aedeagus short, parameres strongly curved, with two distal setae, Italy (FriuliVenezia Giulia)......................................................................  Apheliotyphlus Pace</p><p>- Head capsule with sulcus in shape of V; aedeagus very long, parameres rectilinear, with one distal seta  Megatyphlus Coiffait</p></div>	https://treatment.plazi.org/id/8E7F8798647AFFEDFF047DAAB1BD3BA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047B79B5EA3EF8.text	8E7F87986445FFD2FF047B79B5EA3EF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kladotyphlus ariesensis Struyve 2022	<div><p>ariesensis ariesensis Struyve, 2022: 13</p><p>( Kladotyphlus, TL: Romania,  BrăzeŞti, 46°409N, 23.334°E).</p><p>Types: HNHM HT ♂; 3PT (coll. Struyve)).</p><p>Distribution: Romania.</p></div>	https://treatment.plazi.org/id/8E7F87986445FFD2FF047B79B5EA3EF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047BCDB5EA3F04.text	8E7F87986445FFD2FF047BCDB5EA3F04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kladotyphlus ariesensis Struyve 2022	<div><p>ariesensis orientalis Struyve, 2022: 13</p><p>( Kladotyphlus, TL: Romania,  Iara, 46°527N, 23.577°E)</p><p>Types: HNHM HT ♂.</p><p>Distribution: Romania.</p></div>	https://treatment.plazi.org/id/8E7F87986445FFD2FF047BCDB5EA3F04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047A22B5E73FF4.text	8E7F87986445FFD2FF047A22B5E73FF4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kladotyphlus kahleni Pace 1999	<div><p>kahleni Pace, 1999: 79</p><p>( Kladotyphlus, TL: Slovenia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.016666&amp;materialsCitation.latitude=45.85" title="Search Plazi for locations around (long 15.016666/lat 45.85)">Suha Krajina</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.016666&amp;materialsCitation.latitude=45.85" title="Search Plazi for locations around (long 15.016666/lat 45.85)">Trebnje</a>, Rdeći Kal, vicino della grotta [close to the cave] Koprivnica, 340 m [15°01‘E, 45°51‘N]).</p><p>Types: TLMF HT ♂; 6 PT (TLMF)) .</p><p>Distribution: Slovenia.</p></div>	https://treatment.plazi.org/id/8E7F87986445FFD2FF047A22B5E73FF4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047988B7223D98.text	8E7F87986445FFD2FF047988B7223D98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kladotyphlus Pace 1999	<div><p>Kladotyphlus Pace, 1999</p><p>Kladotyphlus Pace, 1999: 78 .</p><p>Type species:  Kladotyphlus kahleni Pace, 1999 . Original description.</p><p>Diagnosis. Gular sutures confluent, confluent point in about midlength of head capsule, from here prolonged ahead as one straight suture; ligula with two separated projectins; labrum with deep median incision and two wide lateral branches; antennae with 11 antennomeres, antennal club not clearly delimited; tarsal formula 3-3-3.</p><p>Remarks. The genus has only two Balkan species.</p></div>	https://treatment.plazi.org/id/8E7F87986445FFD2FF047988B7223D98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047FFBB5E73B82.text	8E7F87986445FFD2FF047FFBB5E73B82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotyphlus carniolicus Bernhauer 1908	<div><p>carniolicus carniolicus Bernhauer, 1908: 331</p><p>( Leptotyphlus, TL: Savetal in der Wochein [Bohinj]).</p><p>Types: more ST, depository unknown but most probably in FMNH .</p><p>Remarks: Two another subspecies,  M. carniolicus goritianus Pace, 1979 is known from Italy and  M. carniolicus strupianus Pace, 1979 from Austria.</p><p>References: Pace, 1979: 469 (diagnosis), 473 (in key).</p><p>Distribution: Slovenia.</p></div>	https://treatment.plazi.org/id/8E7F87986445FFD2FF047FFBB5E73B82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047D0AB51E3A16.text	8E7F87986445FFD2FF047D0AB51E3A16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megatyphlus Coiffait 1955	<div><p>Megatyphlus Coiffait, 1955</p><p>Megatyphlus Coiffait, 1955: 68 .</p><p>Type species:  Leptotyphlus curtii Breit, 1911 . Original designation.</p><p>syn.  Glyptotyphlus Scheerpeltz, 1956: 462 . Type species:  Leptotyphlus carniolicus Bernhauer, 1908 . Original designation. Validation of  Glyptotyphlus Scheerpeltz 1940 by inclusion of species and designation of type species.</p><p>Glyptotyphlus Scheerpeltz, 1940: 29 nomen nudum, no species designation.</p><p>Glyptotyphlus Scheerpeltz, in litt.: Hölzel, 1944: 67 nomen nudum, no description.</p><p>Megatyphlus Coiffait: Coiffait, 1957: 71 (in key); Coiffait, 1959: 327 (redescription); Coiffait, 1972: 622 (redescription, key to species); Pace, 1979 (revision, key to species); Pace, 1996: 309 (redescription, key to species of Italy).</p><p>Diagnosis. Gular sutures confluent, confluent point in about midlength of head capsule, from here prolonged ahead as one straight suture; ligula reduced, with two small lateral lobes; labrum with anterior margin rectilinear or deeply concave, bearing minuscule teeth; antennae with 11 antennomeres, antennomere 5 about as large as 6, antennal club composed of three antennomeres; tarsal formula 2-2-2.</p><p>Remarks.  Megatyphlus has 3 species and its members with the body length 1.5–1.9 mm are the largest leptotyphlines.</p></div>	https://treatment.plazi.org/id/8E7F87986445FFD2FF047D0AB51E3A16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD3FF047EA3B62A3CC9.text	8E7F87986445FFD3FF047EA3B62A3CC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotyphlus curtii Breit 1911	<div><p>curtii Breit, 1911: 169</p><p>( Leptotyphlus, TL: Monte Maggiore bei Abazzia [Učka near Opatija])</p><p>Types: more ST, depository unknown.</p><p>Synonymy:  Glyptotyphlus jonicus Scheerpeltz, in litt., unavailable name, in Pace, 1983a: 460.</p><p>References: Pace, 1979: 473 (in key).</p><p>Distribution: Croatia (Istria).</p></div>	https://treatment.plazi.org/id/8E7F87986445FFD3FF047EA3B62A3CC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986444FFD3FF04781DB6583D55.text	8E7F87986444FFD3FF04781DB6583D55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megatyphlus uhligi Thelot 1981	<div><p>uhligi Thélot, 1981: 47</p><p>( Megatyphlus, TL: Yugoslavie: Hercegovina, Jablanica).</p><p>Types: ZMHB HT ♂ .</p><p>Distribution: Bosnia and Hercegovina.</p></div>	https://treatment.plazi.org/id/8E7F87986444FFD3FF04781DB6583D55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hlaváč, Peter;Janák, Jiří;Baňař, Petr	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
