taxonID	type	description	language	source
8E7F87986466FFF6FF047AE9B1BC3CFB.taxon	diagnosis	Diagnosis. Very small beetles, body length about 0.8 – 1.9 mm; depigmented; eyes completely atrophied. Head capsule large, neck region short; gular sutures always well-defined, confluent or not in middle or anterior third of head length; mandibles strong, symmetric with one or more teeth; maxillary palpi composed from four palpomeres, palpomere 2 and 3 of different shape and size, tribe characteristic. Antennae with 10 or 11 antennomeres, scape and pedicel always large; antennal club composed from two to five antennomeres. Pronotum about as long as head or slightly shorter. Elytra reduced, slightly wider than long; lacking hind wings. Abdomen longer than head, pronotum and elytra combined; visible sternites simple, slightly impressed or with deep transverse sulcus at posterior region. Leptotyphlinae are actually classified into six tribes which can be separated by following key.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFF6FF047AFDB6183A2C.taxon	type_taxon	Type species: Leptotyphlus aspericeps Scheerpeltz, 1931. Original designation.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFF6FF047AFDB6183A2C.taxon	diagnosis	Diagnosis. Gular sutures confluent in anterior part of head length, from confluent point continuing as single, straight, short suture or again separated to two divergent sutures; antennae with 11 antennomeres, antennomeres 5 larger than 4 and 6, antennal club with 3 antennomeres; labrum dentate with anterior margin rectilinear; tarsal formula 3 - 3 - 3; aedeagus asymmetric, parameres of different shape, with distal setae.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFF6FF047AFDB6183A2C.taxon	discussion	Remarks. Large genus with 25 valid species names assigned in two subgenera. Five species are known from Italy (Trentino-Alto Adige, Veneto, Puglia), three species from Turkey, all from the province Burdur and 17 species from Greece, 10 from mainland (Epirus, Peloponnese, Teygetos, Eastern Macedonia and Thrace) and 7 from islands (Crete, Kefaloniá, Kérkira, Lefkáda).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFF6FF047802B1BC3FA0.taxon	diagnosis	Diagnosis. Mandibles lacking premolar tooth; antennae with 10 or 11 antennomeres; all visible tergites with well-defined paratergites; deep, basal transversal sulcus at least on visible abdominal sternite 5, except for Romanotyphlus and Paratyphlus where it is present also on visible sternites 2 – 4, later genus known from northern Africa, Balearic Islands, Iberian Peninsula and Sardinia); maxillary palpi with palpomere 2 strongly dilated, distinctly larger than palpomere 3.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFF6FF047802B1BC3FA0.taxon	discussion	Remarks. Entomoculiini with seven genera and 256 valid species names is the second most speciose tribe of the subfamily. It has exclusively west Palaearctic distribution and is known from the whole Mediterranean region, going from Portugal up to Turkey, northern Africa, Romania and now also from Serbia. Three Balkan genera, (the distribution Figs 89, 90) can be separated by the following key:	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFFAFF047F71B597392D.taxon	description	Figs 1 – 18, 83 – 84	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFFAFF047F71B597392D.taxon	materials_examined	Material studied. Holotype, ♂: SERBIA with one label “ SERBIA or. 26. v. 2024 / Mt. Stol, soil washing, 660 m / 44.16446 N, 22.11492 E / J. Janák lgt., hornbeam forest ” [white, printed] (JJRC). Paratypes: same data as holotype, 4 ♂♂, 4 ♀♀: (3 ♂♂, 3 ♀♀ in JJRC, 1 ♂, 1 ♀ in MMBC).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFFAFF047F71B597392D.taxon	diagnosis	Diagnosis. Maxillary palpi (Fig. 5) with palpomere 2 dilated, about twice as wide as 3; palpomere 4 small, about three times shorter and narrower than 3; gular sutures confluent anteriorly at about midlength of head capsule, after meeting point again separated to two divergent sutures (Fig. 6); antennae with long scape about as long as terminal antennomere, about 1.6 times as long as wide; pedicel about as long as wide; antennomeres 3 – 10 transverse; terminal antennomere subtriangular, pointed to apex, slightly longer than wide (Fig. 7); visible tergites 1 – 5 and visible sternite 2 – 4 with shallow basal impression (Fig. 3); visible sternite 5 (VII) with deep transversal sulcus (Fig. 3, arrow), visible sternite 6 (VIII) of male symmetric, with median posterior excision which is prolonged with wide, deep impression (Figs 11 – 12); tarsal formula 3 - 3 - 3 (Figs 15 – 16); aedeagus with median lobe prolonged by long slender ventral lamina; parameres short, ventrally left paramere longer than right one, with 3 distal setae (Fig. 17).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFFAFF047F71B597392D.taxon	description	Description. Body (Figs 1, 2) about 1.4 mm long (with abdomen slightly extended posteriad), completely depigmented. Head capsule (Fig. 4) slightly longer than wide; wider than pronotum, posterior part deeply inserted into pronotum; basal capsule separated from neck region by occipital constriction, part of head capsule and neck region retracted into pronotum; gular sutures (Fig. 6) confluent anteriorly at about midlength of head capsule, after meeting point again separated to two divergent arms; maxillary palpi (Fig. 5) with palpomere 2 dilated, about twice as wide as 3; palpomere 4 small, about three times shorter and narrower than 3. Antennae (Fig. 7) long, scape about as long as terminal antennomere, about 1.6 times as long as wide; pedicel as long as wide, as wide as scape, twice as long as antennomere 3; antennomeres 3 – 10 transverse; 3, 4 and 6 subequal in length; antennomere 5 larger than 4 and 6; 5 and 7 subequal in length; 8 as long as 7; 8 and 9 twice as wide as long; 9 1.25 times as long as 8; 10 1.2 times as long as 9; terminal antennomere subtriangular, pointed to apex, slightly longer than wide. Pronotum (Fig. 8) about 1.1 times as long as wide, widest in anterior quarter, after that convergent posteriad; anterior angles widely rounded; posterior angles sharper; about 1.2 times as long as elytra. Prosternum (Fig. 9) fused with proepisternum, procoxal cavities rounded on anterior margin. Elytra (Fig. 8) about as long as wide. Abdomen (Figs 1 – 2) long, considerably longer than length of head capsule, pronotum and elytra combined. Visible tergites 1 – 5 (IV – VIII) and visible sternites 2 – 4 (IV – VI) with shallow basal impression, visible sternite 5 (VII) with deep transverse sulcus (Fig. 3, arrow). Visible tergite 6 (IX) of male expanded in posterior half and with rounded apical margin (Fig. 10). Visible sternite 6 (VIII) of male (Figs 11, 12) symmetric, setose in posterior half, about as long as wide, with median posterior excision which is prolonged with wide, deep impression, its surface shagreened and lateral sides bordered by line of short setae. Terminal abdominal segments of female (Fig. 13) with two paired longitudinal sclerites and half-circled shaped structure in middle. Male sternite IX asymmetric, with posterior margin rounded, lateral parts of tergite IX asymmetric, strongly chitinised, tergite X with short pointed projections laterally (Fig. 14). Legs short, all tibia distally dilated, tarsal formula 3 - 3 - 3 (Figs 15 – 16). Aedeagus (Figs 17, 18) about 0.26 mm long, median lobe prolonged by long slender distal process, basal part about 1.4 times as long as distal process, apex of distal process variable, rounded and slightly curved at apex in lateral view, with sharp, short preapical tooth; sclerite of median lobe well-defined; parameres short, in ventral view left paramere longer than right one, with 3 distal setae (Fig. 17 d).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFFAFF047F71B597392D.taxon	etymology	Etymology. Locotypic, named after Stol planina, the type locality of the species.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFFAFF047F71B597392D.taxon	biology_ecology	Natural History. All specimens were collected by the soil-washing in a belt of hornbeam forest with hazel, maple and oak on a north-facing slope above a stream, together with Banatotyphlus siclus. The volume of the sample had about 100 liters (Figs 83 – 84).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFFAFF047F71B597392D.taxon	discussion	Remarks. Allotyphlus (Moreotyphlus) stolensis has similar aedeagus as A. candicus and A. foedatus (both from Greece, Crete) with long ventral lamina. It can be readily separated from late two species by the distal part of the ventral lamina slightly curved with rounded apex. The apex of the median lobe of A. candicus and A. foedatus is sharp, not curved and it is much shorter in the relation of the length of basal part of median lobe.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986461FFFAFF047F71B597392D.taxon	distribution	Distribution. Serbia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646DFFFAFF047F1FB6033AFE.taxon	materials_examined	Types: MNHN HT ♂, 2 PT (MNHN).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646DFFFAFF047F1FB6033AFE.taxon	distribution	Other localities: PT: Ayos Theodori, près de l’Achileion [Akhílion] à quelques kilomètres au sud de Corfou (Corfou, Iles Ioniennes). Distribution: Greece (Kérkira).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646DFFFAFF047FC6B6033B96.taxon	materials_examined	Types: MNHN HT ♂, 10 PT (MNHN).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646DFFFAFF047FC6B6033B96.taxon	distribution	Other localities: PT: Gouvia and Ipsos, au pied de Mont Pandokrator à quelques kilomètres au nord de Corfou (Corfou, Iles Ioniennes )). Distribution: Greece (Kérkira).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646DFFFBFF047EAFB6033CE4.taxon	materials_examined	Types: 7 ST (FMNH, NMW).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646DFFFBFF047EAFB6033CE4.taxon	distribution	Reference: Pace, 1983 a: 452 (supplement description). Distribution: Greece (Kérkira).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047B77B6373F22.taxon	materials_examined	Types: 13 ST (NMW). References: Pace, 1983 a: 455 (supplement description).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047B77B6373F22.taxon	distribution	Distribution: Greece (Epirus).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047AD0B62A38AD.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂, 11 PT (NMW (Assing coll. )).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047AD0B62A38AD.taxon	distribution	Other localities: Greece: Crete: W Kritsa, 1050 m; S Kritsa, SW Kroustas, 960 m; SW Sitia, NW Magrigialos, 70 m. Distribution: Greece (Crete).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047DB9B61C3938.taxon	materials_examined	Types: NMW HT ♂, 3 PT (NMW, Pace coll. ))	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047DB9B61C3938.taxon	distribution	Distribution: Greece (Kefaloniá).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF0479C1B6033DB8.taxon	materials_examined	Types: 6 ST (MNHN).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF0479C1B6033DB8.taxon	distribution	Other localities: PT: Paleokastritsa, nord-ouest de l’île de Corfou (Iles Ioniennes )). Distribution: Greece (Kérkira).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047C0DB62A3A18.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂, 136 PT (NMW (Assing coll.), ZMHB).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047C0DB62A3A18.taxon	distribution	Other localities: Greece, Crete: SW Chania, W Lakki, 390 m; S Chania, N Thersio, 610 m; SW Chania, WSW Chliaro, 210 m; SW Chania, W Phrases, 640 m; SW Chania, S Nteres, 280 m; S Kissamos, NE Sirikari, 500 m; S Kissamos, Archontiko, 580 m; S Kissamos, Strovles, 470 m; S Kissamos, Elos, 580 m; E Kandanos, Anisaraki, 670 m; S Asigonia, E Kallikratis, 770 m. Distribution: Greece (Crete).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047F2EB6433A87.taxon	materials_examined	Types: MMBC HT ♂, 7 PT (coll. Pavićević, coll. Struyve)	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047F2EB6433A87.taxon	distribution	Distribution: Greece (Peloponnisos).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047FDFB6373B12.taxon	materials_examined	Types: MHNG HT ♂, 2 PT (MHNG, MNHN).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047FDFB6373B12.taxon	distribution	Distribution: Greece (Epirus).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047E33B61C3B82.taxon	materials_examined	Types: NMW HT ♂, 7 PT (NMW, MCNV).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047E33B61C3B82.taxon	distribution	Other localities: Cephallonia, Megalo Vuno; Cephallonia, Krano; Cephallonia, M. Nero. Distribution: Greece (Kefaloniá).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF04788DB5703E6F.taxon	materials_examined	Types: MNHN HT ♂, 5 PT (MNHN)	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF04788DB5703E6F.taxon	distribution	Distribution: Greece (Kérkira). sbg. Moreotyphlus Coiffait, 1959: 310, 312. Type species: Allotyphlus spartensis Coiffait, 1959. Original designation.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047A03B60A3FF6.taxon	materials_examined	Types: NMW HT ♂, 9 PT (NMW).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFFBFF047A03B60A3FF6.taxon	distribution	Other localities: Perivolakia. References: Pace, 1983 a: 454 (supplement description). Distribution: Greece (Lefkáda).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFF8FF047EA4B6433CE4.taxon	materials_examined	Types: MNHN HT ♂, 1 PT (MNHN).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646CFFF8FF047EA4B6433CE4.taxon	distribution	Distribution: Greece (Peloponnisos).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF0479C1B6433D70.taxon	materials_examined	Types: MNHN HT ♂.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF0479C1B6433D70.taxon	distribution	Distribution: Greece (Peloponnisos).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047855B6433DC4.taxon	materials_examined	Types: MNHN HT ♂, 9 PT (MNHN).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047855B6433DC4.taxon	distribution	Other localities: PT: Piyai [Piyaí] sur versant Ouest du Taygète [Taygetos] (Péloponnèse). Distribution: Greece (Peloponnisos).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF0478E1B58D3E0C.taxon	distribution	Distribution: Serbia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047A01B74838C7.taxon	type_taxon	Type species: Cyrtotyphlus convexus Dodero, 1900. Designation by monotypy.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047A01B74838C7.taxon	diagnosis	Diagnosis. Gular sutures confluent in anterior third of head, finished in confluent point; antennae with 10 or 11 antennomeres; antennal club composed of two antennomeres; antennomere 5 about as large as 4 and 6; tarsal formula 2 - 2 - 2; aedeagus symmetric with strongly enlarged parameres, largely covering distal part of median lobe, with some distal setae.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047A01B74838C7.taxon	discussion	Remarks. Smaller genus, with 7 species, 3 species are in France and Italy, one Italian species is also recorded from Slovenia, two species are in Greece, one in Eastern Macedonia and Thrace and one on the island Samothráki. One species is known from Montenegro and one from Romania.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047CABB6733A6F.taxon	distribution	Distribution: Romania.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047CABB6733A6F.taxon	discussion	Note. The species was originally described as C. carpathica. Cyrtotyphlus is a masculine, thus the specific name must be changed to C. carpathicus.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047F77B5CE3AFB.taxon	materials_examined	Types: NMW HT ♂.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047F77B5CE3AFB.taxon	distribution	Distribution: Montenegro.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047FCBB74B3BF7.taxon	materials_examined	Types: unknown.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646FFFF8FF047FCBB74B3BF7.taxon	distribution	Other localities: Italy: Veneto (Recoaro: Fongara; Monti Berici: Zovencedo, Arcugnano, Lumignano; Arquà Petrarca: Monte Fasolo; Monte Venda, Marostica, Soligo), Friuli-Venezia Giulia (Taipana, Strgna, Cormons, Premariacco, Dolegna del Collio); Slovenia: Planina; Logateč. References: Coiffait, 1962: 159 (upgrade from subspecies to species level). Distribution: Italy (Veneto, Friuli-Venezia Giulia), Slovenia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047D9EB05F39EF.taxon	diagnosis	Diagnosis. Maxillary palpomere 2 strongly dilated, much more developed than palpomere 3; tarsi with 2 or 3 tarsomeres; prosternal coxal cavities rounded on anterior margin, lacking excision; lacking suture separating proepisternum from prosternum; all visible tergites with well-defined paratergites; all visible abdominal sternites lacking deep basal transverse sulcus.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047D9EB05F39EF.taxon	discussion	Remarks. The richest tribe of the subfamily. It holds 10 genera and 264 valid species names. The large majority of species is in the Mediterranean region. Out of this region, there are three species in the Democratic Republic of Congo, one in Tanzania, one in Indonesia (West Papua) and one, Leptotyphlus kovaci Šustek, 2000, in Slovakia. Only one genus, Kenotyphlus (distribution on Fig. 91) is present in the Balkan Peninsula.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047855B7093D9D.taxon	materials_examined	Types: unknown.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047855B7093D9D.taxon	distribution	Distribution: Greece (Eastern Macedonia and Thrace).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047F2EB0283BB5.taxon	type_taxon	Type species: Kenotyphlus turcicus Coiffait, 1957. Original description.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047F2EB0283BB5.taxon	diagnosis	Diagnosis. Gular sutures structure unknown; labrum dentate, with two teeth; premolar tooth well-defined; antennae with 11 antennomeres, antennomere 5 much larger than 4 but about as large as 6, antennal club composed from 5 antennomeres; terminal antennomeres lacking sensorial appendages (phaneres); tarsal formula 2 - 2 - 2; genital armature of female indistinct.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047F2EB0283BB5.taxon	discussion	Remarks. Kenotyphlus has 6 species, three in Greece and three in Turkey.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047989B66F3D71.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂, 5 PT (NMW (Assing coll. )).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047989B66F3D71.taxon	distribution	Other localities: Greece: Samothráki, Therma. Distribution: Greece (Samothráki).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047AD2B6E0381F.taxon	materials_examined	Types: HNHM HT ♂; 20 PT (HNHM, coll. Struyve )).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF047AD2B6E0381F.taxon	distribution	Distribution: Romania (Apuseni Mountains).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF0478E1B7903F3F.taxon	type_taxon	Type species: Romanotyphlus metaliferensis Struyve, 2022. Original designation.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF0478E1B7903F3F.taxon	diagnosis	Diagnosis. Gular sutures confluent in midlength of head, from confluent point again separated in two, divergent sutures; labrum edentate; antennae with 11 antennomeres, antennomeres 5 larger than 4 and 6, antennal club with 3 antennomeres; tarsal formula 2 - 2 - 2; aedeagus asymmetric, parameres short with one strong, distal seta.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646EFFF9FF0478E1B7903F3F.taxon	discussion	Remarks. Monospecific genus, recently described from Romania.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF047A00B1BD3BCA.taxon	diagnosis	Diagnosis. Maxillary palpomeres 2 and 3 strongly and about equally dilated; mandibles lacking premolar tooth; tarsi with 2 or 3 tarsomeres; prosternal coxal cavities rounded on anterior margin, lacking excision; lacking suture separating proepisternum from prosternum; all visible tergites with well-defined paratergites; all visible abdominal sternites lacking deep basal transverse sulcus.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF047A00B1BD3BCA.taxon	discussion	Remarks. The tribe Metrotyphlini has actually, together with newly described genus, nine genera and 41 valid species names. Six genera are known in the Palaearctic region and two, monospecific genera, occur in South America, Apotyphlus Coiffait & Saiz, 1965 in Chile and Venezillotyphlus Decu, 1990 in Venezuela. Un undescribed taxon is reported from Mexico (Navarrete-Heredia et al. 2002). The following key separates Metrotyphlini of the Palaearctic region.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF04781CB62A3DB9.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂, 6 PT (NMW (Assing coll. ))	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF04781CB62A3DB9.taxon	distribution	Distribution: Greece (Crete).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF04788CB6353DC5.taxon	materials_examined	Types: MHNG HT ♂, 11 PT (MHNG, MNHN)	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF04788CB6353DC5.taxon	distribution	Distribution: Greece (Ródos).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF0478E0B62A3EFD.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂, 33 PT (NMW (Assing coll.), ZMHB).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986469FFFEFF0478E0B62A3EFD.taxon	distribution	Other localities: Greece: Crete: S Kissamos, NE Elos, 460 m; S Kissamos, Aikirgianis, 290 m; SW Chania, Mavros, 380 m; Asigonia, 380 m. Distribution: Greece (Crete).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047C08B5EA3949.taxon	distribution	Distribution: Romania.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047988B6E73E0D.taxon	type_taxon	Type species: Banatotyphlus racovitzai Decu, 1982. Designated by monotypy.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047988B6E73E0D.taxon	diagnosis	Diagnosis. Gular sutures confluent in anterior third of head length, from confluent point again separated in two, divergent sutures; labrum with small, median tooth; antennae with 11 antennomeres, antennomeres 5 about as large as 4 and 6, antennal club with 5 antennomeres; tarsal formula 3 - 3 - 3; ventral part of tarsomere 2 and 3 with spatulate setae; aedeagus asymmetric, parameres completely reduced, invisible.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047988B6E73E0D.taxon	discussion	Remarks. The genus is separated from other Metrotyphlini by palpomere 2 and 3 subequal in width; antennae short, antennomeres 3 – 10 transverse, 3 – 6 subequal in size, antennomeres 9 – 11 with sensorial appendages (phaneres); tarsi with 3 tarsomeres. The genus was erected for one species, Banatotyphlus racovitzai Decu 1982 and another three species were described later (Struyve 2022).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047C9CB5EA39D5.taxon	distribution	Distribution: Romania.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047CF0B5EA3AA9.taxon	materials_examined	Types: HT ♂, deposition unknown.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047CF0B5EA3AA9.taxon	distribution	Other localities: Romania, Şopotu Nou, 44.809 ° N, 21.842 ° E. References: Struyve 2022: 15. Distribution: Romania.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047FBCB6003B7D.taxon	materials_examined	Types: HNHM HT ♂; 15 PT (HNHM, coll. Struyve). Other localities: Cozia (sic!) [= Cozla], 44.631 ° N, 22.007 ° E and 44.628 ° N, 22.039 ° E; New records: Serbia or., 14. x. 2022 and 26. v. 2024, Mt. Stol, 660 m, 44.16446 N, 22.11492 E.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047FBCB6003B7D.taxon	distribution	Distribution: Romania, Serbia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047B90B4D03869.taxon	description	Figs 19 – 24, 83 – 84	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047B90B4D03869.taxon	materials_examined	Material examined. SERBIA: 1 ♂, 1 ♀: with one label “ SERBIA or. 14. x. 2022 / Mt. Stol, soil washing, 660 m / 44.16446 N, 22.11492 E / J. Janák lgt., hornbeam forest ” [white, printed] (JJRC). 16 ♂♂, 6 ♀♀: same locality label but collected on 26. v. 2024 (JJRC, MMBC).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047B90B4D03869.taxon	description	Supplement description. Visible tergite 6 (IX) of male elongate with rounded posterior margin (Fig. 19). Visible sternite 6 (VIII) of male with deep emargination in posterior third (Fig. 20), male tergites IX and X as in Fig. 21, male sternite IX apically rounded (Fig. 22). Aedeagus (Figs 23 – 24) dorsaly elongate, slender, four times as long as wide, lateraly with apophysis in shape of “ S ”. First record for Serbia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047B90B4D03869.taxon	biology_ecology	Natural History. The same as for Allotyphlus stolensis sp. nov. (Figs 83 – 84).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986468FFFFFF047B90B4D03869.taxon	distribution	Distribution. Romania, Serbia (Western Carpathians, on both side of the border between Romania and Serbia).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047988B4C93E24.taxon	type_taxon	Type species: Cretotyphlus hamatus Assing, 2019. Original designation.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047988B4C93E24.taxon	diagnosis	Diagnosis. Mandibles apically bifid; gular sutures close, but not confluent anteriorly; labrum tridentate, maxillary palpi with palpomeres 2 and 3 enlarged and somewhat flattened, 3 larger than 2; antenna with 11 antennomeres, moderately incrassate, apical antennomeres lacking sensorial appendages, antennal club composed of three last antennomeres; tarsi with 3 tarsommeres; aedeagus with long, slender, and distally acute ventral process, lacking parameres; female genital segment with more or less distinct pair of sclerites.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047988B4C93E24.taxon	discussion	Remarks. The genus was erected for three new species from Crete and later another species was discovered in Sámos.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047BCEB62A3F68.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂, 2 PT (NMW (Assing coll. )).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047BCEB62A3F68.taxon	distribution	Distribution: Greece (Crete).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047A7EB62A3FD8.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂, 3 PT (NMW (Assing coll. )).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047A7EB62A3FD8.taxon	distribution	Distribution: Greece (Crete).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047AEEB62A3848.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂, 2 PT (NMW (Assing coll. )).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047AEEB62A3848.taxon	distribution	Distribution: Greece (Crete).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047D9EB6383938.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047D9EB6383938.taxon	distribution	Distribution: Greece (Sámos).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047C46B13D3B1D.taxon	type_taxon	Type species: Egeotyphlus thracicus Coiffait, 1957. Original designation.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047C46B13D3B1D.taxon	diagnosis	Diagnosis. Gular sutures confluent in anterior third of length of head capsule, from confluent point prolonged ahead as one straight suture; labrum edentate, its anterior margin in middle concave; antennae short, antennae with 11 antennomeres, all antennomeres transverse, 5 clearly larger than 4 and 6, antennomeres 8 – 11 with sensorial appendages (phaneres), inserted on upper part and curved towards lower part of antennomeres; antennal club composed from 5 antennomeres; all tarsi with 3 tarsomeres.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646BFFFCFF047C46B13D3B1D.taxon	discussion	Remarks. The diagnosis of Egeotyphlus was updated by Coiffait (1972). After transferring Egeotyphlus zecevici to the genus Epirotyphlus, the genus Egeotyphlus became monospecific (see remarks for Epirotyphlus).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFFDFF04781CB7093D55.taxon	materials_examined	Types: depository unknown.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFFDFF04781CB7093D55.taxon	distribution	Distribution: Greece (Eastern Macedonia and Thrace).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFFDFF0478A8B5F3395E.taxon	type_taxon	Type species: Egeotyphlus leucadiensis Pace, 1995. Original designation. Described as a subgenus of the genus Egeotyphlus. New status.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFFDFF0478A8B5F3395E.taxon	diagnosis	Diagnosis. Gular sutures confluent in about midlength of head capsule, from this confluent point they continue as two, separate, short divergent sutures (Figs 31, 61); labrum dentate (Figs 30, 59); antennae with 11 antennomeres, antennomere 5 larger than 4 and 6; antennal club composed from 5 antennomeres (Figs. 32, 62), last 4 antennomeres with (Fig. 63) or without sensorial appendages (phaneres) but those if present always inserted on lower part and oriented towards upper part of antennomeres.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFFDFF0478A8B5F3395E.taxon	discussion	Remarks. Epirotyphlus was originally described as a subgenus of Egeotyphlus for a new species, Egeotyphlus (Epirotyphlus) leucadiensis from the Greek island Lefkáda. Epirotyphlus was separated from the nominal subgenus by 1) lacking large sensorial appendages (phaneres) on the last four antennomeres inserted on upper part and curved towards lower part, 2) having dentate labrum and 3) aedeagus lacking parameres. Pace (1995), when describing Egeotyphlus (Epirotyphlus) leucadiensis recognized that this species is more similar to his previously described species Egeotyphlus zecevici from Serbia (Pace 1984 b) than to the type species of the genus, Egeotyphlus thracicus. These both species share all characters mentioned above and more, they have very similar structure of the aedeagus. Aedeagus is slender, symmetric, parameres are reduced, lacking distal setae. The discovery of two new species, which are described here below and assigned to Epirotyphlus, allowed us to study the ventral part of head and it seems that the structure of gular sutures can be an important character to separate Epirotyphlus from Egeotyphlus. Both new species, have gular sutures confluent in about midlength of head capsule and from this confluent point they continue as two, separate, short divergent sutures, opposite to Egeotyphlus, where they continue as a simple, straight suture. Unfortunately, Pace (1984 b, 1995) did not provide any information about the structure of gular sutures for either Egeotyphlus (Epirotyphlus) leucadiensis or Egeotyphlus zecevici. Thus, the importance of the structure of gular sutures for the definition of Epirotyphlus will need further study, especially when the type species, Epirotyphlus leucadiensis will be available. Regardless this, we are of the opinion that Epirotyphlus deserves to be upgraded to the genus level.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFE6FF047CDFB5D83EC1.taxon	description	Figs 25 – 54, 85	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFE6FF047CDFB5D83EC1.taxon	materials_examined	Material studied. Holotype, ♂: MONTENEGRO with one label “ MONTENEGRO SW 013, Đurovići / 42 ° 11 ’ 56 ’’ N, 18 ° 58 ’ 44 ’’ E / 60 m, 23. III. 2024, soil / washing near oak tree, P. Baňař lgt ” [white, printed] (MMBC). Paratypes: 7 ♂, 7 ♀, same data as holotype: (3 ♂, 4 ♀ in MMBC, 4 ♂, 3 ♀ in JJRC). 4 ♂, 5 ♀: “ MONTENEGRO SW 009, Đurovići, 2. v. 2023, 42 ° 11 ‘ 56 ‘‘ N, 18 ° 58 ‘ 44 ‘‘ E, 60 m, soil washing near oak tree, P. Baňař & P. Hlaváč lgt. ” [white, printed] (BMNH, MMBC, JJRC). 1 ♂: MONTENEGRO SW 002, Rumija, road from Bar → Mikulići, 30. iv. 2023, 42 ° 07 ‘ 01 ‘‘ N, 19 ° 09 ‘ 35 ‘‘ E, 675 m, soil washing near oak tree, P. Baňař lgt. ” [white, printed] (MMBC). 5 ♀: MONTENEGRO SW 010, Rumija, road from Bar → Mikulići, 21. iii. 2024, 42 ° 07 ‘ 01 ‘‘ N, 19 ° 09 ‘ 35 ‘‘ E, 674 m, soil washing near Carpinus tree, P. Baňař lgt. ” [white, printed] (MMBC). 2 ♂, 2 ♀, “ MONTENEGRO SW 014, Đurovići / 42 ° 11 ‘ 55 ‘‘ N, 18 ° 58 ‘ 46 ‘‘ E / 58 m, 23. III. 2024, soil / washing close stream, P. Baňař lgt ” [white, printed] (MMBC). 1 ♂, 2 ♀: “ MONTENEGRO SW 006, Đurovići, 2. v. 2023, 42 ° 11 ‘ 57 ‘‘ N, 18 ° 58 ‘ 49 ‘‘ E, 70 m, soil washing near olive tree, P. Baňař lgt. ” [white, printed] (MMBC). 2 ♀: MONTENEGRO SW 007, near Blizikuće, near petrol station, 29. iv. 2023, soil washing near Juniperus tree, P. Baňař & P. Hlaváč lgt. [white, printed] ” (MMBC).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFE6FF047CDFB5D83EC1.taxon	diagnosis	Diagnosis. Head longer than wide (Fig. 28), longer than pronotum; maxillary palpi with palpomeres 2 and 3 dilated, 3 larger than 2 (Fig. 29, 45); ligula indistinct (Fig. 44); mandibles falciform (Fig. 45); labrum dentate (Fig. 30); gular suture confluent anteriorly in about midlength of head capsule (Fig. 31). Antennae with 11 antennomeres, antennal club composed of 5 antennomeres, with sensorial appendages which are inserted on lower part and oriented towards upper part of antennomere, antennomeres 3 – 10 strongly transverse (Figs 32, 49). Elytra tetragonal, dilated posteriad (Fig. 33), shorter than pronotum. Procoxal cavities (Figs 34, 50) rounded on anterior margin, lacking excision, prosternum and proepisternum fused. Abdominal sternites lacking deep, transversal sulcus, visible sternite 2 – 5 with shallow basal impression (Figs 27, 46). Tarsal formula 3 - 3 - 3, tarsomere 1 minuscule, 2 slightly longer, 3 longest, clearly longer than 1 and 2 combined (Figs 38 – 40, 52). Aedeagus elongate, slender and nearly symmetrical (Fig. 41, 42); lacking parameres, lateral protrusions of median lobe long (Fig. 42 d, arrows). Terminal abdominal segments of female (Fig. 37) with one paired longitudinal sclerite.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFE6FF047CDFB5D83EC1.taxon	description	Description. Body (Figs 25, 26, 46, 48) depigmented, most part of surface shagreened (Figs 43, 47, 54), body length about 0.95 mm. Head capsule (Fig. 28, 43) wider than long, about 1.2 times as wide as long, shorter than pronotum; maxillary palpi (Fig. 29, 45) with palpomeres 2 and 3 dilated, 3 larger than 2; ligula indistinct (Fig. 29, 44); mandibles falciform (Fig. 45); labrum dentate (Fig. 30); gular suture confluent anteriorly behind midlength of head capsule, after meeting point again separated to two divergent arms (Fig. 31). Antennae (Fig. 32, 49) with 11 antennomeres, scape bulbous, larger than pedicel, pedicel dilated distad, distally truncate, antennomeres 3 – 10 strongly transverse, antennomere 5 larger than 4 and 6, antennal club composed of 5 antennomeres, with sensorial appendages which are inserted on lower part and oriented towards upper part of antennomere. Pronotum (Fig. 33, 43) wider than long, convergent posteriad. Elytra (Fig. 33, 47, el) tetragonal, dilated posteriad, shorter than pronotum, external posterior angles rounded, internal ones right-angled. Venter, procoxal cavities (Fig. 34, 50) rounded on anterior margin, anterior margin with inconspicuous excision (50, arrow), lacking suture separating proepisternum from prosternum, lateral portions of prosternum shagreened, medium portion smooth. Abdominal sternites (Fig 27, 46, 48) lacking deep, transversal sulcus, visible sternite 2 – 6 with shallow basal impression (Fig. 27). Legs (Figs 38 – 40, 52, 53) short, all tibiae dilated distad, tarsal formula 3 - 3 - 3, tarsomere about half of length of 3, tarsomere 2 shortest, about 3 times shorter than 1. Aedeagus (Figs 41, 42) about 0.17 mm long, median lobe with long lateral protrusions (Fig. 42 d, arrows). Terminal abdominal segments of female (Fig. 37) with one paired longitudinal sclerite.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFE6FF047CDFB5D83EC1.taxon	etymology	Etymology. Named after our friend Dušica Šoljaga, owner of a beautiful house which is a base of our expeditions to the coastal part of Montenegro.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFE6FF047CDFB5D83EC1.taxon	biology_ecology	Natural History. The majority of specimens (holotype and 27 paratypes) of newly described species was collected by the soil washing of a sample taken near and around a solitary oak (Fig. 85) on the border of terrace garden together with Ubychia ganglbaueri (Reitter, 1912) and an undesribed species of Cephennium and Tychobythinus in May 2023 and March 2024. Several specimens were collected in Rumija Mts. in about 675 m a. s. l. and two females also in coastal Mediterranean forest under junipers tree near Blizikuće village.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798646AFFE6FF047CDFB5D83EC1.taxon	distribution	Distribution. Montenegro.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986471FFE5FF047A5DB5973BE7.taxon	description	Figs 55 – 76, 86 – 87	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986471FFE5FF047A5DB5973BE7.taxon	materials_examined	Material studied. Holotype, ♂: SERBIA with one label “ SERBIA or. 22. v. 2024 / Majdan Kučajna, 230 m / 44.44561 N, 21.63905 E / soil washing, J. Janák lgt. / hornbeam forest “ [white, printed] (JJRC). Paratypes: 2 ♂♂, 3 ♀♀ same data as for holotype (1 ♂, 3 ♀♀ in JJRC, 1 ♂ in MMBC).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986471FFE5FF047A5DB5973BE7.taxon	diagnosis	Diagnosis. Gular sutures confluent anteriorly slightly behind midlength of head capsule (Fig. 61); antennae long, antennomeres 3 – 10 transverse; antennomeres 5 larger than 4 and 6; antennal club composed from five antennomeres (Figs 62 – 63), incrassate, last four antennomeres setose, with sensorial appendages which are inserted on lower part and oriented towards upper part of antennomere (Fig. 63); procoxal cavities rounded on anterior margin, with small excision on lateral part (Fig. 65, arrow); tergites 1 – 5 (IV – VIII) and visible sternites 2 – 4 (IV – VI) (Fig. 57) lacking basal impression (Fig. 57); tarsal formula 3 - 3 - 3 (Figs 71 – 73); aedeagus elongate, about 3 times as long as wide; distal, ventral part of median lobe with deep excision, lateral arms truncate; lacking parameres, lateral protrusions of median lobe short (Figs 74, 75). Terminal abdominal segments of female (Fig. 76) with one paired longitudinal sclerite and large asymmetric structure in middle.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986471FFE5FF047A5DB5973BE7.taxon	description	Description. Body (Figs 55, 56) about 0.9 mm long, completely depigmented. Head capsule (Fig. 58) about 1.2 wider than long; about as wide and as long as pronotum; head capsule separated from neck region by occipital constriction, neck region retracted into pronotum; gular sutures (Fig. 61) confluent anteriorly slightly behind midlength of head capsule; maxillary palpi (Fig. 60) with palpomere 2 dilated, about as wide as 3 and about half of length of 3; palpomere 4 very small. Antennae (Figs 62, 63) long, scape about 1.45 times as long as pedicel, about 1.2 times as long as wide; pedicel as long as wide, 3.5 times as long as antennomere 3; antennomeres 3 – 10 transverse; antennomeres 5 larger than 4 and 6; antennal club composed from five antennomeres, incrassate, 10 slightly wider than long, elliptical, terminal antennomere subspherical. Last four antennomeres (Fig. 63) setose, with sensorial appendages which are inserted on lower part and oriented towards upper part of antennomere. Pronotum (Fig. 64) about 1.1 times as wide as long, widest in anterior third, after that convergent posteriad; anterior angles widely rounded; posterior angles sharper; about 1.1 times as long as elytra. Venter, prosternum (Fig. 65) fused with proepisternum, procoxal cavities rounded on anterior margin, with small excision (Fig. 65, arrow) on lateral part. Elytra (Fig. 64) about as long as wide, widest on posterior margin. Abdomen long, considerably longer than length of head capsule, pronotum and elytra combined. Visible tergites 1 – 5 (IV – VIII) and visible sternites 2 – 4 (IV – VI) (Fig. 57) lacking basal impression. Visible sternite 6 (IX) of male (Figs 67, 68) symmetric, visible sternite 7 (IX) in male (Fig. 70) with small setae; sparsely setose in posterior half, visible tergite 7 (X) with median posterior excision which is prolonged with wide, deep impression (Fig. 69). Visible tergite 5 (VIII) (Fig. 66) 1.25 times as long as wide. Legs (Figs 71 – 73) short, all tibia distally dilated, tarsal formula 3 - 3 - 3; first tarsomere longer than 2, 3 longer than 1 and 2 combined. Aedeagus (Figs 74, 75) about 0.17 mm long, elongate, about 3 times as long as wide; distal, ventral part of median lobe with deep excision, lateral arms truncate; basal part less than half of length of median lobe; with elongate, long, internal apophysis; lacking parameres, lateral protrusions of median lobe short. Terminal abdominal segments of female (Fig. 76) with one paired longitudinal sclerite and large asymmetric structure in middle.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986471FFE5FF047A5DB5973BE7.taxon	etymology	Etymology. Patronymic, named after Jana Janáková, the wife of the second author to thank her for being a companion on trips to the Balkan countries.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986471FFE5FF047A5DB5973BE7.taxon	discussion	Remarks. E. janae has the same type of the aedeagus as E. zecevici but it differs from latter mainly by apically narrowed lateral projections and truncate lateral arms of median lobe (Pace 1984, Figs 2 – 4).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986471FFE5FF047A5DB5973BE7.taxon	biology_ecology	Natural History. All specimens were collected by soil-washing on a south-facing slope above the road at the edge of a deciduous forest (hornbeam, hazel, maple, oak). Volume of sample was about 100 liters (Figs 86 – 87).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986471FFE5FF047A5DB5973BE7.taxon	distribution	Distribution. Serbia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFEBFF047F61B58D3AD0.taxon	materials_examined	Types: 4 ST (coll. Nonveiller, coll. Pace).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFEBFF047F61B58D3AD0.taxon	distribution	Distribution: Serbia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFEBFF047C9EB5CE39F0.taxon	distribution	Distribution: Montenegro.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFEBFF047F29B58D3A44.taxon	distribution	Distribution: Serbia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFEBFF047CD6B60A3A1C.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFEBFF047CD6B60A3A1C.taxon	distribution	Distribution: Greece (Lefkáda).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFE9FF047E38B7853E59.taxon	type_taxon	Type species: Leptotyphlus perpusillus Dodero, 1900. Original designation.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFE9FF047E38B7853E59.taxon	diagnosis	Diagnosis. Gular sutures confluent in anterior third of length of head capsule, from confluent point not prolonged ahead; labrum dentate; antennae short, with 11 antennomeres, antennomeres 5 – 10 strongly transverse, 5 wider than 4 and about as wide as 6, antennal club composed from 5 antennomeres, terminal antennomeres lacking sensorial appendages (phaneres); all tarsi with 3 tarsomeres. Genital armature of female indistinct.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647CFFE9FF047E38B7853E59.taxon	discussion	Remarks. The genus has three species, all are present in the Balkan Peninsula. It is believed that the natural area of the distribution is southern France, Italy, the Balkans and Turkey and that the parthenogenetic species Gynotyphlus perpusillus was introduced in many countries in Europe.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFE9FF047D91B02139F8.taxon	discussion	Remarks: Eleven subpecies of Gynotyphlus perpusillus were described (Löbl & Löbl 2015: 915), two from the territory of the Balkans, Gynotyphlus perpusillus dalmaticus Coiffait, 1955: 72 (TL: Dalmatie: Susurac [Sućurac] from southern Dalmatia, Croatia) and Gynotyphlus perpusillus macedonicus Coiffait, 1957: 81 (TL: Macédoine Yugoslave [North Macedonia]: Katlanovo). The validity of all ten subspecies is doubtfull and it will need the revison (Struyve 2022: 7).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFE9FF047D91B02139F8.taxon	distribution	Reference: Struyve 2022: 7 (distribution in Romania, note on the validity of subspecies). Distribution: Portugal, France, Corsica, Italy, Sardinia, Sicily, Switzerland, Austria, Greece, Ionian Islands, Croatia, North Macedonia, Turkey, Morocco, Romania (Struyve 2022: 7).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFE9FF047A39B6373FE0.taxon	materials_examined	Types: NHM HT ♀, 6 PT ♀ (NMW).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFE9FF047A39B6373FE0.taxon	distribution	Reference: Violi & Fancello, 2022: 1 (the necessity to verify the validity of the species). Distribution: Greece (Epirus).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFE9FF047AC5B60338B4.taxon	materials_examined	Types: 6 ST (NMW).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFE9FF047AC5B60338B4.taxon	distribution	Other localities: Val de Ropa auf Korfu. Reference: Violi & Fancello, 2022: 1 (the necessity to verify the validity of the species). Distribution: Greece (Kérkira).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFEEFF047F05B5053913.taxon	type_taxon	Type species: Megatyphlus hervei Coiffait, 1957. Original designation.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFEEFF047F05B5053913.taxon	diagnosis	Diagnosis. Gular sutures confluent in anterior third of length of head capsule, from confluent point prolonged ahead by two short, divergent sutures; labrum dentate; antennae with 11 antennomeres, 5 slightly larger than 4 and 6, antennal club composed from 3 antennomeres; terminal antennomeres lacking sensorial appendages (phaneres); all tarsi with 2 tarsomeres. Genital armature of female distinct or indistinct.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647EFFEEFF047F05B5053913.taxon	discussion	Remarks. Metrotyphlus, together with Apotyphlus and Venezillotyphlus have tarsal formula 2 - 2 - 2. The genus has 2 subgenera differentiated by the structure of aedeagus and female genital armature. All balkanic species belong to the subgenus Agnotyphlus Coiffait, 1972. The nominal subgenus includes four species in France, Italy and Switzerland.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986479FFEEFF047CFEB6743A57.taxon	materials_examined	Types: MCSN HT ♂.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986479FFEEFF047CFEB6743A57.taxon	distribution	Distribution: Greece (Khalkidhiki).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986479FFEEFF047F6EB5EA3B2B.taxon	materials_examined	Types: HNHM HT ♂; 13 PT (HNHM, coll. Struyve). Other records: Romania, Cozia (sic!) [= Cozla], 44.628 ° N, 22.039 ° E.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986479FFEEFF047F6EB5EA3B2B.taxon	distribution	Distribution: Romania.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986479FFEEFF047E3AB62F3BFF.taxon	materials_examined	Types: MSNV HT ♂; 13 PT (MSNV, MCST, coll. Pace). Other records: Slovenia, Lokvica, 215 m.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986479FFEEFF047E3AB62F3BFF.taxon	distribution	Distribution: Italia, Slovenia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986478FFEFFF047989B62A3CC8.taxon	materials_examined	Types: SOFM HT ♂, 1 PT (SOFM).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986478FFEFFF047989B62A3CC8.taxon	distribution	Distribution: Croatia (Istria).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986478FFEFFF04781DB66F3DB8.taxon	materials_examined	Types: NMW (Assing coll.) HT ♂.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986478FFEFFF04781DB66F3DB8.taxon	distribution	Distribution: Greece (Samothráki).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986478FFEFFF047F21B08F3BF9.taxon	type_taxon	Type species: Leptotyphlus beieri Scheerpeltz, 1931. Original designation.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986478FFEFFF047F21B08F3BF9.taxon	diagnosis	Diagnosis. Gular sutures confluent in half of length of head capsule, from confluent point prolonged ahead by long, single suture; labrum dentate, with three prominent teeth; antennae with 11 antennomeres, antennal club composed from 3 antennomeres; terminal antennomeres lacking sensorial appendages (phaneres); tarsal formula 3 - 3 - 3; genital armature of female distinct.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986478FFEFFF047F21B08F3BF9.taxon	discussion	Remarks. Monospecific genus known only from the type material, the male is unknown.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647BFFECFF04781CB60A3DB9.taxon	materials_examined	Types: 3 ST (NMW).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647BFFECFF04781CB60A3DB9.taxon	distribution	References: Pace, 1983 a: 459 (supplement description). Distribution: Greece (Lefkáda).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647AFFEDFF047DAAB1BD3BA4.taxon	diagnosis	Diagnosis. Maxillary palpi with palpomeres 2 and 3 strongly, equally dilated; ligula on anterior margin with two widely well-separated projections; mandibles lacking premolar tooth; coxal cavities with lateral excision on anterior margin which is prolonged by suture separating prosternum from proepisternum; all visible abdominal sternite lacking deep transversal sulcus.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F8798647AFFEDFF047DAAB1BD3BA4.taxon	discussion	Remarks. The most widespread tribe of the family with 20 genera and 51 valid species names which is considered to be the most primitive leptotyphlines (Smetana 1986). The tribe is richest in taxa in the New World with 16 genera and 42 species, where Chile with 5 genera and 23 species and California (USA) with 7 genera and 12 species are the richest known regions. One genus with two species is known from South Africa and three genera and seven species from Europe. Kladotyphlus and Megatyphlus are present in the Balkan Peninsula. Apheliotyphlus is monospecific genus known in Italy (Friuli-Venezia Giulia) and because of the vicinity of this region to Slovenia, the presence of the genus in Slovenia can be plausible. All three European genera can be separated by the following key (Pace 1999, translated and modified).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047B79B5EA3EF8.taxon	materials_examined	Types: HNHM HT ♂; 3 PT (coll. Struyve )).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047B79B5EA3EF8.taxon	distribution	Distribution: Romania.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047BCDB5EA3F04.taxon	materials_examined	Types: HNHM HT ♂.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047BCDB5EA3F04.taxon	distribution	Distribution: Romania.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047A22B5E73FF4.taxon	materials_examined	Types: TLMF HT ♂; 6 PT (TLMF )).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047A22B5E73FF4.taxon	distribution	Distribution: Slovenia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047988B7223D98.taxon	type_taxon	Type species: Kladotyphlus kahleni Pace, 1999. Original description.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047988B7223D98.taxon	diagnosis	Diagnosis. Gular sutures confluent, confluent point in about midlength of head capsule, from here prolonged ahead as one straight suture; ligula with two separated projectins; labrum with deep median incision and two wide lateral branches; antennae with 11 antennomeres, antennal club not clearly delimited; tarsal formula 3 - 3 - 3.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047988B7223D98.taxon	discussion	Remarks. The genus has only two Balkan species.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047FFBB5E73B82.taxon	materials_examined	Types: more ST, depository unknown but most probably in FMNH.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047FFBB5E73B82.taxon	discussion	Remarks: Two another subspecies, M. carniolicus goritianus Pace, 1979 is known from Italy and M. carniolicus strupianus Pace, 1979 from Austria. References: Pace, 1979: 469 (diagnosis), 473 (in key).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047FFBB5E73B82.taxon	distribution	Distribution: Slovenia.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047D0AB51E3A16.taxon	type_taxon	Type species: Leptotyphlus curtii Breit, 1911. Original designation.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047D0AB51E3A16.taxon	diagnosis	Diagnosis. Gular sutures confluent, confluent point in about midlength of head capsule, from here prolonged ahead as one straight suture; ligula reduced, with two small lateral lobes; labrum with anterior margin rectilinear or deeply concave, bearing minuscule teeth; antennae with 11 antennomeres, antennomere 5 about as large as 6, antennal club composed of three antennomeres; tarsal formula 2 - 2 - 2.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD2FF047D0AB51E3A16.taxon	discussion	Remarks. Megatyphlus has 3 species and its members with the body length 1.5 – 1.9 mm are the largest leptotyphlines.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD3FF047EA3B62A3CC9.taxon	materials_examined	Types: more ST, depository unknown.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986445FFD3FF047EA3B62A3CC9.taxon	distribution	Distribution: Croatia (Istria).	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986444FFD3FF04781DB6583D55.taxon	materials_examined	Types: ZMHB HT ♂.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
8E7F87986444FFD3FF04781DB6583D55.taxon	distribution	Distribution: Bosnia and Hercegovina.	en	Hlaváč, Peter, Janák, Jiří, Baňař, Petr (2025): Endogean and cavernicolous Coleoptera of the Balkans. XXVIII. Synopsis of the subfamily Leptotyphlinae Fauvel, 1874 (Coleoptera, Staphylinidae) of the Balkan Peninsula, with the description of three new species from Serbia and Montenegro. Zootaxa 5594 (1): 61-97, DOI: 10.11646/zootaxa.5594.1.3, URL: https://doi.org/10.11646/zootaxa.5594.1.3
