taxonID	type	description	language	source
F1763122DBFE5093903B67A818528ED9.taxon	diagnosis	Diagnosis. These insects are well characterized by the synapomorphic presence in the forewing of large median cells and strong angles in the courses of R and M, all absent in the Lophioneuridae (sensu Nel et al. 2012, 2014) and Moundthripidae Nel, Azar, & Nel, 2007. These three families have a CuA re-emerging first from R + M + CuA but CuA is not orthogonal to R + M in the Moundthripidae versus orthogonal in Iotacyphidae and Lophioneuridae (see below).	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
22E6595FCC9E5AF98C6C59F6598F1A19.taxon	diagnosis	Diagnosis. Forewing with CuA re-emerging orthogonally from R + M + CuA.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
AF2C7FF3A7105D6C85FAD56905D9004A.taxon	type_taxon	(Carboniferous Westphalothripidesidae P. Nel, Azar, Prokop, Roques, Hodebert, & A. Nel, 2012 to crown-Thysanoptera) (= “ Thripida Fallén, 1814 ” sensu P. Nel et al. 2014)	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
AF2C7FF3A7105D6C85FAD56905D9004A.taxon	diagnosis	Diagnosis. Left mandibular stylet, right absent; piercing lacinial stylets; pretarsus with eversible arolium; pretarsal claws reduced in adults; wings with fringe of long setae and CuA simple.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
13725FD7043C51B98F4BAF5807C83B3B.taxon	type_taxon	Type species. Triassocypha prima sp. nov.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
13725FD7043C51B98F4BAF5807C83B3B.taxon	etymology	Etymology. The name is an artificial combination of Triassic, referring to period from which the fossils originate, and the Ancient Greek adjective κῡφός (kūphós, meaning, “ bent forward ” or “ curved ”; feminine κῡφή). The gender of the name is feminine.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
13725FD7043C51B98F4BAF5807C83B3B.taxon	diagnosis	Diagnosis. Forewing ScP re-emerging from R as a short, straight veinlet perpendicular to C; a closed cell between MA and MP; two longitudinal cells between R and M / MA; CuP + PCu well separated from A.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
8033400C167853E18DD29D3F80C6CB52.taxon	description	Figures 2 A, B, D, 3 A	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
8033400C167853E18DD29D3F80C6CB52.taxon	etymology	Etymology. The specific epithet is the Latin adjective prīmus, meaning, “ first ” or “ early ”, and refers to this as the first described species of the genus.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
8033400C167853E18DD29D3F80C6CB52.taxon	diagnosis	Diagnosis. Forewing characters only: ma-mp longer than r-ma; distal-most cell between R and MA three times as long as wide.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
8033400C167853E18DD29D3F80C6CB52.taxon	description	Description. Body 1.3 mm long, presence of long macrosetae on the thorax and some wing veins. Head: head apparently orthognathous to prognathous, with a relatively long buccal cone; compound eyes rounded with diameter 0.1 mm; antenna elongate, with flagellomeres elongate, distal most flagellomeres shorter than other flagellomeres, scape large, pedicel apparently small. Thorax: thorax 0.5 mm long; all legs preserved and of similar shapes, apparently only two tarsomeres, basal one light brown and distal one dark brown. Abdomen: abdomen elongate and broad, 0.7 mm long, 0.55 mm wide; cerci absent. Forewing venation: forewing 1.6 mm long, 0.6 mm wide, pear-shaped, rounded apically, narrow basally; a common stem ScP + R + M + CuA from which CuA separates 0.2 mm from wing base, perpendicular to ScP + R + M and short before cua-cup terminates into vein; cua-cup 0.07 mm long, straight, aligned with distal part of CuA; CuA simple and curved distally, ending at wing margin 0.5 mm from wing base; CuP + PCu straight, well separated from A; A short, weakly curved; ScP re-emerging from R 0.8 mm from wing base, perpendicular to C and R, straight, 0.12 mm long; M separating from R 0.1 mm distal of base of CuA; stem of M curved, 0.25 mm long, separating into MA and MP and delimiting a broad elongate closed cell, 0.5 mm long, 0.2 mm wide; both MA and MP with a strong angle distally and ending on posterior margin at right angle; a long ma-mp perpendicular to MA and MP; two elongate, broad cells between R and MA, separated by r-ma perpendicular to R and MA, basal-most cell 0.35 mm long, 0.07 mm wide; distal-most cell 0.6 mm long, 0.2 mm wide; R forming an angle distal of ScP and straight distally; separating into RA and RP 0.45 mm distal of ScP; RA straight, short, rather oblique, nearly orthogonal with RP; RP forming a strong angle and reaching wing apex; straight, elongate rp-ma, 0.1 mm long.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
05146F1C0D605D0C87B67D81E8B3237C.taxon	description	Figures 2 C, E, 3 B	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
05146F1C0D605D0C87B67D81E8B3237C.taxon	etymology	Etymology. The specific epithet is the Latin numeral secundus, meaning “ second ”, and refers to this as the second described species of the genus.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
05146F1C0D605D0C87B67D81E8B3237C.taxon	diagnosis	Diagnosis. Forewing characters only: ma-mp as long as r-ma; distal-most cell between R and MA two times as long as wide.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
05146F1C0D605D0C87B67D81E8B3237C.taxon	description	Description. Body 1.4 mm long, presence of long macrosetae on the head, thorax, and some wing veins. Head: head apparently orthognathous to prognathous, with a relatively long buccal cone; compound eyes rounded with diameter 0.1 mm; [antenna not preserved]. Thorax: 0.37 mm long, 0.4 mm high; legs partly preserved. Abdomen: broad, 0.9 mm long, 0.7 mm wide; cerci absent. Forewing venation: forewing 1.7 mm long, 0.8 mm wide, pear-shaped, rounded apically, narrow basally; a common stem ScP + R + M + CuA; CuA, cua-cup, and CuP not visible; ScP re-emerging from R 0.8 mm from wing base, perpendicular to C and R, straight, 0.15 mm long; M separating from R 0.6 mm distal of base of CuA; stem of M curved, 0.5 mm long, separating into MA and MP and delimiting a broad elongate closed cell, 0.4 mm long, 0.19 mm wide; both MA and MP with a strong angle distally and ending on posterior margin at right angle; a long ma-mp perpendicular to MA and MP; two elongate, broad cells between R and MA, separated by r-ma perpendicular to R and MA, basal-most cell 0.4 mm long, 0.15 mm wide; distal-most cell 0.46 mm long, 0.23 mm wide; R forming an angle distal of ScP and straight distally, separating into RA and RP 0.5 mm distal of ScP; RA straight, short, rather oblique, nearly orthogonal with RP; RP forming a strong angle, reaching wing apex; straight, elongate rp-ma, 0.12 mm long.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
CF6F0A2E69C4546DBB9958ECC8B0F5CB.taxon	type_taxon	Type genus. Triassocypha gen. nov.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
CF6F0A2E69C4546DBB9958ECC8B0F5CB.taxon	diagnosis	Diagnosis. Two closed cells between R and M and a closed cell between the branches of M.	en	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
