identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A68334CB1AD35DAE8704526486E3E785.text	A68334CB1AD35DAE8704526486E3E785.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthomoplax Souma & Kamitani 2021	<div><p>Genus Acanthomoplax Souma &amp; Kamitani, 2021</p><p>Acanthomoplax Souma &amp; Kamitani, 2021: 4. Type species by original designation: Acanthomoplax tomokunii Souma &amp; Kamitani, 2021 .</p><p>Note.</p><p>For detailed diagnostic characters of the genus, see Souma and Kamitani (2021) and Souma (2022 a).</p><p>Remarks.</p><p>The monotypic genus Acanthomoplax, which is endemic to the Ogasawara Islands, Japan, comprises only A. tomokunii distributed in Chichijima and Hahajima groups (Souma and Kamitani 2021; Souma 2022 a).</p></div>	https://treatment.plazi.org/id/A68334CB1AD35DAE8704526486E3E785	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Souma, Jun	Souma, Jun (2025): An illustrated key to the lace bugs (Hemiptera, Heteroptera, Tingidae) from “ Oriental Galapagos ” (the Ogasawara Islands, Japan), with descriptions of three new species of the endemic genus Omoplax Horváth, 1912. ZooKeys 1250: 243-284, DOI: 10.3897/zookeys.1250.160064
728BA545518653418757B14F20C9E19A.text	728BA545518653418757B14F20C9E19A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthomoplax tomokunii Souma & Kamitani 2021	<div><p>Acanthomoplax tomokunii Souma &amp; Kamitani, 2021</p><p>Figs 1 A, 2 A, 3 A, 4 A, 5 A, 6 A, 7 A, 8 A, 9 A, 10 A, 11 A, 12 A, 13 A, 14 A, 15 A – E</p><p>Acanthomoplax tomokunii Souma &amp; Kamitani, 2021: 6. Holotype: ♀; type locality: Japan • Ogasawara Isls., Hahajima I., Mt. Chibusayama [= Ogasawara Islands, Hahajima Group, Hahajima Island, Mt. Chibusa]; ELKU.</p><p>References.</p><p>Souma (2022 a: 125) (distribution); Shimamoto and Ishikawa (2023: 93) (catalog); Souma (2023: 9) (monograph).</p><p>Material examined.</p><p>Non-types, Japan • 2 ♂♂ 1 ♀; Ogasawara Isls., Ototojima Is., Kurohama – Ichinotani; Machilus kobu; 5 Oct. 2024; J. Souma leg.; three third or fourth instar nymphs developed into adults by 15 Oct. 2024 by feeding on Machilus kobu in captivity; SIHU • 1 ♀; same collection data as for preceding; a single egg has developed into a second instar nymph until 17 Oct. 2024 by feeding on Machilus kobu in captivity, and a second instar nymph developed into an adult by 30 Oct. 2024 by feeding on Machilus thunbergii in captivity; SIHU.</p><p>Diagnosis.</p><p>Acanthomoplax tomokunii is recognized among other lace bug species based on diagnostic characters mentioned in previous studies (Souma and Kamitani 2021; Souma 2022 a) and can be distinguished from the seven other lace bug species occurring in the Ogasawara Islands based on a combination of the following characteristics: pair of frontal spines reaching beyond apex of clypeus (Figs 3 A, 4 A, 5 A, 6 A, 14 A); median spine reaching beyond bases of frontal spines; pair of occipital spines reaching beyond anterior margin of compound eye; hood medially with robust denticles throughout its length; median carina of pronotum with robust denticles throughout its length; paranotum narrowed posteriad; outer margin of paranotum with robust denticles throughout its length; Sc (subcosta) vein of hemelytron with robust denticles throughout its length; and R + M (fused radius and media) vein of hemelytron with robust denticles throughout its length (Figs 7 A, 8 A, 9 A, 10 A).</p><p>Remarks.</p><p>Segmental oligomery of the antenna was confirmed in Acanthomoplax tomokunii, and one examined specimen lacks the left antennal segment IV (Fig. 2 A), as reported in many lace bugs (Štusák and Stehlík 1978).</p><p>Distribution.</p><p>Japan: Ogasawara Islands: Chichijima Group (Anijima Island, Ototojima Island), Hahajima Group (Hahajima Island) (Fig. 18) (Souma and Kamitani 2021; Souma 2022 a). Acanthomoplax tomokunii, which is endemic to the Ogasawara Islands, was confirmed in Ototojima Island during the survey in 2024 but has not been collected in Anijima and Hahajima islands since 2014 and 1999, respectively, despite extensive field surveys by numerous investigators (cf. Souma and Kamitani 2021; Souma 2022 a).</p><p>Host plant.</p><p>Machilus kobu ( Lauraceae) (Fig. 17 A), which is also known as “ Kobugashi ” and is endemic to the Ogasawara Islands (Government of Japan 2010), was confirmed as a host plant for Acanthomoplax tomokunii through field and captive observations of adults and nymphs. In captivity, A. tomokunii fed on M. thunbergii Siebold &amp; Zucc., “ Tabunoki ” (Fig. 17 B), which is not distributed in the Ogasawara Islands (Tanaka and Matsui 2007–2025), and a single second instar nymph developed normally to an adult in at least 13 days, suggesting the possibility of rearing this species in captivity using closely related species of M. kobu, which are not found in the native distribution area of A. tomokunii . However, to the best of the author’s knowledge, A. tomokunii feeds only on M. kobu in the field and appears to be monophagous. Additionally, the single individual in the present study was reared on M. kobu to the second instar nymph. Therefore, it is unclear whether rearing from oviposition to emergence can be achieved successfully only by feeding on M. thunbergii .</p><p>Bionomics.</p><p>Acanthomoplax tomokunii inhabits an evergreen broad-leaved forest with a subtropical climate in the Ogasawara Islands (Souma and Kamitani 2021), and sucks sap on the abaxial side of the leaves of M. kobu, causing irregular yellowing on the adaxial side (Fig. 17 A). Adults were collected in March and April, and from June to August (Souma and Kamitani 2021; Souma 2022 a); nymphs were collected in October.</p></div>	https://treatment.plazi.org/id/728BA545518653418757B14F20C9E19A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Souma, Jun	Souma, Jun (2025): An illustrated key to the lace bugs (Hemiptera, Heteroptera, Tingidae) from “ Oriental Galapagos ” (the Ogasawara Islands, Japan), with descriptions of three new species of the endemic genus Omoplax Horváth, 1912. ZooKeys 1250: 243-284, DOI: 10.3897/zookeys.1250.160064
5B505D714ACA5F73B90C9255F47A87CA.text	5B505D714ACA5F73B90C9255F47A87CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Omoplax desecta (Horvath 1912)	<div><p>Omoplax desecta (Horváth, 1912)</p><p>Figs 1 B, 2 B, 3 B, 4 B, 5 B, 6 B, 7 B, 8 B, 9 B, 10 B, 11 B, 12 B, 13 B, 14 B, 15 F – H</p><p>Stephanitis (Omoplax) desecta Horváth, 1912: 337. Syntype (s): ♀; type locality: Japan • Ogasawara [= Ogasawara Islands]; Laboratory of Systematic Entomology, Faculty of Agriculture, Hokkaido University, Sapporo, Japan (Tomokuni 1994: 844; Souma 2022 a: 125) .</p><p>Omoplax desecta: Takeya (1962: 74) (new combination).</p><p>References.</p><p>Drake (1948: 55) (checklist: genus); Takeya (1951: 14) (checklist: eastern Asia); Drake and Maa (1953: 100) (checklist: genus); Drake (1956: 116) (distribution); Drake and Ruhoff (1960: 83) (catalog); Drake and Ruhoff (1965: 336) (catalog); Nakane (1970: 28) (checklist: Ogasawara Islands); Miyamoto and Yasunaga (1989: 167) (checklist: Japan); Nishimura and Arai (1989: 38) (distribution); Kato (1992: 79) (checklist: Ogasawara Islands); Yasunaga et al. (1993: 179) (monograph); Tomokuni (1994: 844) (type material); Yasunaga (1995: 20) (illustration); Péricart and Golub (1996: 51) (checklist: Palearctic); Miyano (1998: 8) (distribution); Guilbert (2001: 551) (distribution); Karube and Takakuwa (2004: 83) (illustration); Ohbayashi et al. (2004: 32) (checklist: Ogasawara Islands); Government of Japan (2010: 208) (checklist: Ogasawara Islands); Yamada and Tomokuni (2012: 198) (monograph); Takahashi et al. (2014: 163) (natural enemy); Yamada and Ishikawa (2016: 432) (checklist: Japan); Souma and Kamitani (2021: 8) (distribution: part); Souma (2022 a: 125) (distribution: part); Shimamoto and Ishikawa (2023: 93) (catalog: part); Souma (2023: 9) (monograph). Several studies conducted before the 1980 s recorded this species as Stephanitis (Omoplax) desecta (Drake 1948; Takeya 1951; Drake and Maa 1953; Drake 1956; Drake and Ruhoff 1960, 1965; Nakane 1970; Nishimura and Arai 1989).</p><p>Material examined.</p><p>Non-types, Japan • 1 ♂; Ogasawara Isls, Chichijima Is.; 6 May 1974; Y. Hori leg.; NSMT • 1 ♀; same locality data as for preceding; Rhaphiolepis indica var. tashiroi; 20 Mar. 2001; G. Tokihiro; NSMT • 1 ♀; Ogasawara Isls, Hahajima Is., Higashiko; 5 Jun. 1976; T. Nakane leg.; NSMT • 1 ♀; Ogasawara Isls, Hahajima Is., Mt. Chibusa; 8 Jun. 1976; T. Nakane leg.; NSMT • 1 ♀; same locality data as for preceding; 6 Jul. 1997; T. Kishimoto leg; NSMT • 2 ♀♀; same locality data as for preceding; 7 Jul. 1997; K. Matsumoto leg; NSMT • 1 ♀; same locality data as for preceding; 27 May 2022; T. Yoshida leg.; SIHU • 2 ♂♂ 5 ♀♀; same locality data as for preceding; Rhaphiolepis indica var. tashiroi; 2 Oct. 2024; J. Souma; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., Mt. Mikazuki; 12 Jun. 1976; T. Nakane leg.; NSMT • 23 ♂♂ 18 ♀♀ 1 fifth instar nymph; same locality data as for preceding; 18 Apr. 1997; K. Matsumoto leg.; NSMT • 1 ♀; same locality and date data as for preceding; T. Kishimoto leg.; NSMT • 1 ♂ 3 ♀♀; same locality data as for preceding; 23 Apr. 1997; K. Matsumoto leg.; NSMT • 2 ♂♂ 6 ♀♀; same locality data as for preceding; 17 May 2024; Y. Hisasue leg.; SIHU • 1 ♂ 1 ♀ 1 fifth instar nymph; same locality data as for preceding; Rhaphiolepis indica var. tashiroi; 21 Sep. 2024; J. Souma; SIHU • 2 ♀♀; same locality, host plant, and collector data as for preceding; 22 Sep. 2024; SIHU • 2 ♀♀; same locality data as for preceding; 25 Oct. 2024; Y. Hisasue; SIHU • 1 ♀; same locality and collector data as for preceding; 5 Nov. 2024; SIHU • 1 ♀; same locality and collector data as for preceding; 10 Jan. 2025; SIHU • 3 ♂♂ 4 ♀♀; same locality and collector data as for preceding; 15 May 2025; SIHU • 1 ♂ 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Shigure; 8 Dec. 1977; M. Tomokuni leg.; NSMT • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Takayama; 30 Jul. 1996; K. Matsumoto leg.; NSMT • 2 ♀♀; Ogasawara Isls, Hahajima Is., Kuwanoki-zawa; 4 Aug. 1996; K. Matsumoto leg.; NSMT • 1 ♂; Ogasawara Isls, Chichijima Is., Higashidaira; 15 Apr. 1997; K. Matsumoto leg.; NSMT • 14 ♂♂ 8 ♀♀ 3 fifth instar nymphs 1 fourth instar nymph; Ogasawara Isls, Chichijima Is., Buta Beach; 16 Apr. 1997; K. Matsumoto leg.; NSMT • 7 ♂♂ 4 ♀♀; same locality and date data as for preceding; T. Kishimoto leg.; NSMT • 1 ♀; Ogasawara Isls, Chichijima Is., Nagatani – Mt. Tsutsuji; 17 Apr. 1997; K. Matsumoto leg; NSMT • 18 ♂♂ 14 ♀♀; Ogasawara Isls, Hahajima Is., Minamizaki; 19 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♂; same locality data as for preceding; 4 Jul. 1997; T. Kishimoto leg; NSMT • 1 ♂; same locality data as for preceding; 15 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♀; same locality data as for preceding; Rhaphiolepis indica var. tashiroi; 27 Sep. 2024; J. Souma; SIHU • 1 ♂ 3 ♀♀; Ogasawara Isls, Hahajima Is., Nakanotaira – Minamizaki; 19 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♂; Ogasawara Isls, Hahajima Is., Mt. Mikazuki (non-existent place name: mislabeling?); 19 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♂ 1 ♀; Ogasawara Isls, Hahajima Is., Komoridani; 20 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♂ 1 ♀; Ogasawara Isls, Hahajima Is., Mt. Yakeyama; 20 Apr. 1997; K. Matsumoto leg; NSMT • 3 ♂♂; Ogasawara Isls, Hahajima Is., Oki-mura; 20 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♀; same locality and date data as for preceding; T. Kishimoto leg; NSMT • 1 ♀; Ogasawara Isls, Chichijima Is., Minamifukurozawa; 23 Apr. 1997; K. Matsumoto leg; NSMT • 3 ♀♀ 1 fifth instar nymph; same locality data as for preceding; 23 Mar. 2024; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Ichinotani; 26 Apr. 1997; K. Matsumoto leg; NSMT • 1 ♂; Ogasawara Isls, Ototojima Is., Kurohama; 27 Apr. 1997; K. Matsumoto leg; NSMT • 2 ♂♂; Ogasawara Isls, Hahajima Is., Nishiura; 4 Jul. 1997; K. Matsumoto leg; NSMT • 1 ♀; Ogasawara Isls, Hahajima Is., Mt. Kuwanoki; 4 Jul. 1997; T. Kishimoto leg; NSMT • 1 ♂ 1 ♀; Ogasawara Isls, Hahajima Is., Tamagawa Dam; 5 Jul. 1997; K. Matsumoto leg.; NSMT • 1 ♀; same locality data as for preceding; 14 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Tsutsuji; 10 Jul. 1997; T. Kishimoto leg.; NSMT • 1 ♂; Ogasawara Isls, Ototojima Is., Ainosawa; 1 Jul. 2021; T. Matsumoto &amp; S. Shimamoto leg.; SIHU • 6 ♂♂ 4 ♀♀; same locality data as for preceding; 3–4 Jul. 2024; N. Tsuji leg.; SIHU • 2 ♀♀; Ogasawara Isls, Anijima Is., Central Plateau; 1 Jul. 2021; T. Matsumoto &amp; S. Shimamoto leg.; SIHU • 1 ♂; same locality and collector data as for preceding; 2 Jul. 2021; SIHU • 1 ♀; Ogasawara Isls, Anijima Is., Mt. Togari; 29 Jul. 2021; T. Matsumoto &amp; S. Shimamoto leg.; SIHU • 1 ♂; Ogasawara Isls, Hahajima Is., Igumadani, Sekimon; alt. 370 m; 26 May 2022; T. Yoshida leg.; SIHU • 2 ♂♂; Ogasawara Isls, Chichijima Is., Susaki; 21 May 2023; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. One; 4 Feb. 2024; Y. Hisasue leg.; SIHU • 2 ♂♂ 1 ♀; Ogasawara Isls, Meijima Is.; 14 Jun. 2024; N. Tsuji leg.; SIHU • 1 ♂; Ogasawara Isls, Hahajima Is., Nishidai; 17 Jun. 2024; N. Tsuji leg.; SIHU • 1 ♂ 5 ♀♀; Ogasawara Isls, Hahajima Is., Shizukazawa; 20 Jun. 2024; Y. Hisasue leg.; SIHU • 7 ♂♂ 6 ♀♀; Ogasawara Isls, Chichijima Is., Mt. Ogami; alt. 10–90 m; 11 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♀; Ogasawara Isls, Ototojima Is., Mt. Sokuryogatake; 12 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♂ 1 ♀; Ogasawara Isls., Anijima Is., Mansaku – Kanaimisaki; 12 Jul. 2024; N. Tsuji leg.; SIHU • 1 ♀; Ogasawara Isls, Hahajima Is., Nijuccho Pass; alt. 160 m; 15 Jul. 2024; Y. Uehara leg.; SIHU • 5 ♂♂ 1 ♀; Ogasawara Isls, Mukohjima Is.; alt. 0–90 m; 16 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Tsuitate; 19 Jul. 2024; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Hatsuneura; 28 Jul. 2024; N. Tsuji leg.; SIHU • 1 ♂ 1 ♀; Ogasawara Isls, Chichijima Is., Higashi-machi; Calophyllum inophyllum; 21 Sep. 2024; J. Souma leg.; SIHU • 1 ♀; same locality, host plant, and collector data as for preceding; 8 Oct. 2024; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Yoake; Rhaphiolepis indica var. tashiroi; 22 Sep. 2024; J. Souma leg.; SIHU • 1 ♂ 1 ♀; Ogasawara Isls., Ototojima Is., Kurohama – Ichinotani; Rhaphiolepis indica var. tashiroi; 23 Sep. 2024; J. Souma leg.; SIHU • 3 ♂♂ 1 ♀; Ogasawara Isls., Anijima Is., Tamana Beach – Mt. Mikaeri; Rhaphiolepis indica var. tashiroi; 24 Sep. 2024; J. Souma leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Ogamiyama Park; Rhaphiolepis indica var. tashiroi; 25 Sep. 2024; J. Souma leg.; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., John Beach; Calophyllum inophyllum; 7 Oct. 2024; J. Souma leg.; SIHU • 2 ♀♀; Ogasawara Isls, Nishijima Is.; 17 Nov. 2024; N. Tsuji leg.; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., Oku-mura; 10 Dec. 2024; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Hahajima Is., Hyogidaira; Rhaphiolepis indica var. tashiroi; 9 Mar. 2025; J. Souma leg.; SIHU • 1 ♀; same locality, host plant, and collector data as for preceding; 11 Mar. 2025; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., Kominato Beach; 25 Apr. 2025; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Chichijima Is., Mt. Sakaiura; 27 Apr. 2025; Y. Hisasue leg.; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., Mulberry Bay; 29 Apr. 2025; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Hahajima Is., Nishiura; 4 May 2025; Y. Hisasue leg.; SIHU • 1 ♂ 4 ♀♀; Ogasawara Isls, Ototojima Is., Shikahama – Mt. Hirone; 27 May 2025; S. Shimamoto leg.; SIHU • 3 ♂♂ 5 ♀♀; Ogasawara Isls, Ototojima Is., Kohama; 2 Jun. 2025; Y. Hisasue leg.; SIHU . The four nymphs recorded above are in poor condition and are thus not described in the present study.</p><p>Diagnosis.</p><p>Omoplax desecta is recognized among the other Omoplax species based on a combination of the following characteristics: rostrum reaching posterior margin of metasternum (Fig. 11 B); pronotal disc pale brown (Figs 3 B, 4 B, 5 B, 6 B); hood more than 0.5 times as wide as maximum width of head across compound eyes, not reaching apex of clypeus (Fig. 14 B); paranotum without areolae in middle part, with areolae in remaining parts; anterior margin of hemelytron weakly curved downward in apical half (Figs 7 B, 8 B, 9 B, 10 B); subcostal and discoidal areas of hemelytron not united; costal area narrower than combined width of subcostal and discoidal areas; Sc (subcosta) vein of hemelytron distinct in apical part of dorsal view; R + M (fused radius and media) vein of hemelytron indistinct, not carinate; and ventral surface of body in various shades of brown (Figs 12 B, 13 B).</p><p>Remarks.</p><p>In general appearance, Omoplax desecta is very similar to O. majorcarinae, whose distribution range is consistent with that of O. desecta in Chichijima Group (Guilbert 2001; Souma and Kamitani 2021; Souma 2022 a); however, the former can be distinguished from the latter based on the following five main characters (Figs 3 B, G, 4 B, G, 5 B, G, 6 B, G, 7 B, G, 8 B, G, 9 B, G, 10 B, G, 11 B, G, 14 B, G): rostrum reaching posterior margin of metasternum (reaching middle part of mesosternum in O. majorcarinae); anterior margin of hemelytron weakly curved downward in apical half (strongly curved downward in apical half in O. majorcarinae); subcostal and discoidal areas of hemelytron not united (united in O. majorcarinae); Sc (subcosta) vein of hemelytron distinct in apical part of dorsal view (indistinct in apical part of dorsal view in O. majorcarinae); and R + M (fused radius and media) vein of hemelytron distinct, carinate (indistinct, not carinate in O. majorcarinae). Morphological differences between O. desecta and the five other Omoplax species are presented in the identification key below.</p><p>Distribution.</p><p>Japan: Ogasawara Islands: Chichijima Group (Anijima Island, Chichijima Island, Nishijima Island, Ototojima Island), Hahajima Group (Hahajima Island, Meijima Island, Mukohjima Island), Mukojima Group (Nakodojima Island) (Fig. 18) (Government of Japan 2010; Souma and Kamitani 2021; Souma 2022 a; Shimamoto and Ishikawa 2023). Omoplax desecta is endemic to the Ogasawara Islands.</p><p>Host plant.</p><p>Rhaphiolepis indica var. tashiroi ( Rosaceae) (Fig. 17 C) and Calophyllum inophyllum ( Clusiaceae) (Fig. 17 D), also known as “ Shimasharinbai ” and “ Terihaboku ”, respectively, were confirmed as host plants for Omoplax desecta by the field and captive observations of adults and nymphs, suggesting the possibility of polyphagy for this lace bug species. However, no feeding behavior of O. desecta was observed on Ardisia sp. ( Primulaceae), Bischofia javanica Blume ( Phyllanthaceae), Cinnamomum sp. ( Lauraceae), Ligustrum sp. ( Oleaceae), and Terminalia sp. ( Combretaceae), from which only a few adults were collected in previous studies (cf. Yasunaga et al. 1993; Guilbert 2001; Yamada and Tomokuni 2012). Therefore, these five tree species do not appear to be host plants for this lace bug species.</p><p>Bionomics.</p><p>Omoplax desecta inhabits an evergreen broad-leaved forest with a subtropical climate in the Ogasawara Islands (Souma and Kamitani 2021), and sucks sap on the abaxial side of the leaves of Rhaphiolepis indica var. tashiroi and Calophyllum inophyllum, causing irregular yellowing on the adaxial side (Fig. 17 C, D). Adults were collected in all seasons (Drake 1956; Nishimura and Arai 1989; Yasunaga et al. 1993; Tomokuni 1994; Miyano 1998; Guilbert 2001; Souma and Kamitani 2021; Souma 2022 a); nymphs were collected in March, April, and October. In Anijima Island, O. desecta has been confirmed in the stomach contents of the invasive green anole, Anolis carolinensis ( Squamata, Dactyloidae) (Takahashi et al. 2014).</p></div>	https://treatment.plazi.org/id/5B505D714ACA5F73B90C9255F47A87CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Souma, Jun	Souma, Jun (2025): An illustrated key to the lace bugs (Hemiptera, Heteroptera, Tingidae) from “ Oriental Galapagos ” (the Ogasawara Islands, Japan), with descriptions of three new species of the endemic genus Omoplax Horváth, 1912. ZooKeys 1250: 243-284, DOI: 10.3897/zookeys.1250.160064
B56AA5BF20905550A354519D2630BF7B.text	B56AA5BF20905550A354519D2630BF7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Omoplax hisasuei Souma 2025	<div><p>Omoplax hisasuei sp. nov.</p><p>Figs 1 C, 2 C, 3 C, 4 C, 5 C, 6 C, 7 C, 8 C, 9 C, 10 C, 11 C, 12 C, 13 C, 14 C, 15 I</p><p>Type material.</p><p>Holotype, Japan • ♂; Ogasawara Isls., Hahajima Is., Mt. Kuwanoki; Neolitsea sericea var. aurata; 29 Sep. 2024; J. Souma leg.; SIHU . Paratypes, Japan • 6 ♂♂ 5 ♀♀; same data as for holotype; SIHU • 1 ♂ 1 ♀; same locality and date data as for holotype; Y. Hisasue leg.; SIHU • 1 ♂; same locality, host plant, and collector data as for holotype; 8 Mar. 2025; SIHU • 2 ♂♂; same locality, host plant, and collector data as for holotype; 9 Mar. 2025; SIHU • 13 ♂♂ 14 ♀♀; same locality data as for holotype; 22 May 2025; S. Shimamoto leg.; SIHU .</p><p>Additional material examined.</p><p>Non-types, Japan • 1 fifth instar nymph; same locality and date data as for holotype; Y. Hisasue leg.; SIHU . The single nymph recorded above is in poor condition and is thus not described in the present study.</p><p>Diagnosis.</p><p>Omoplax hisasuei sp. nov. is recognized among the other Omoplax species based on a combination of the following characteristics: rostrum reaching middle part of mesosternum (Fig. 11 C); pronotal disc pale brown (Figs 3 C, 4 C, 5 C, 6 C); hood less than 0.5 times as wide as maximum width of head across compound eyes, not reaching apex of clypeus (Fig. 14 C); paranotum with areolae throughout its length; anterior margin of hemelytron strongly curved downward in apical half (Figs 7 C, 8 C, 9 C, 10 C); subcostal and discoidal areas of hemelytron united; costal area wider than fused subcostal and discoidal areas; Sc (subcosta) vein of hemelytron indistinct in apical part of dorsal view; R + M (fused radius and media) vein of hemelytron indistinct, not carinate; and ventral surface of body dark brown to black (Figs 12 C, 13 C).</p><p>Description.</p><p>Male. Head, antenna, calli, and legs in various shades of brown; pronotal disc, hood, median carina of pronotum, paranotum, posterior process, and hemelytron pale brown; markings on dorsum and ventral surface of body dark brown to black; compound eye dark red; areolae of pronotum and hemelytron transparent; pubescence on body yellowish (Figs 1 C, 3 C, 5 C, 7 C, 9 C, 11 C, 12 C).</p><p>Body ovate; pubescence on body distinctly shorter than radius of compound eye (Figs 1 C, 12 C). Head (Figs 3 C, 5 C) glabrous; pair of frontal spines separated from each other at apices, not reaching apex of clypeus, occasionally reduced; median spine not reaching bases of frontal spines, occasionally reduced; pair of occipital spines not reaching anterior margin of compound eyes, occasionally reduced; antenniferous tubercle obtuse, curved inward, longer than frontal spines; vertex and clypeus smooth. Compound eye round in dorsal view. Antenna densely covered with minute pubescence on segments I to III and long pubescence on segment IV; pubescence on segment IV longer than pubescence on other parts of body; segment I cylindrical, shorter than segment IV; segment II cylindrical, shortest among antennal segments; segment III linear, longest amongst antennal segments; segment IV fusiform. Bucculae closed at anterior ends, with 3 rows of areolae at highest part. Rostrum (Fig. 11 C) reaching middle part of mesosternum.</p><p>Pronotum (Figs 1 C, 3 C, 5 C) glabrous. Pronotal disc coarsely punctate. Hood shorter than median carina of pronotum, higher than median carina, with 3–4 rows of areolae at highest part, less than 0.5 times as wide as maximum width of head across compound eyes, not reaching apex of clypeus, without robust denticles throughout its length; dorsal margin arched; posterior margin extending to anterior part of pronotal disc. Collar not covering compound eye. Median carina straight, extending to apex of posterior process, with 1–2 rows of areolae at highest part, without robust denticles throughout its length; dorsal margin arched. Calli smooth. Paranotum subvertical, widened posteriad, with a single row of areolae in anterior half and 2 rows in posterior half; outer margin gently curved outward throughout its length, without robust denticles throughout its length. Posterior process triangular; apex rounded.</p><p>Hemelytron (Figs 7 C, 9 C) glabrous, extending beyond apex of abdomen; anterior margin strongly curved downward in apical half; apices separated from each other at rest; subcostal and discoidal areas united; costal area wider than fused subcostal and discoidal areas, with 4–5 rows of areolae at widest part; fused subcostal and discoidal areas with 7 rows of areolae at widest part; sutural area with 4–5 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; Sc (subcostal) and Hc (hypocosta) veins distinct throughout their respective length; R + M (fused radius and media) and Cu (cubitus) veins indistinct throughout their respective length, not carinate; Sc and R + M veins without robust denticles throughout their respective length; Sc vein indistinct in apical part of dorsal view.</p><p>Thoracic pleura smooth in anterior part, coarsely punctate in posterior part. Ostiolar peritreme oblong. Sternal laminae (Fig. 11 C) lower than bucculae; pro- and mesosternal laminae open at both anterior and posterior ends; metasternal laminae as high as mesosternal laminae, open at anterior ends, fused with each other at posterior ends. Legs (Fig. 1 C) smooth, covered with pubescence; femora thickest at middle. Abdomen ellipsoidal. Terminalia (Fig. 12 C) pentagonal in ventral view, covered with pubescence.</p><p>Measurements (n = 24). Body length with hemelytra 3.05–3.30 mm; maximum width of body across hemelytra 1.65–1.75 mm; length of antennal segments I to IV 0.20 mm, 0.10 mm, 1.20 mm, and 0.70 mm, respectively; pronotal length 1.10–1.20 mm; pronotal width across paranota 0.80–0.85 mm; hemelytral length 2.40–2.55 mm; maximum width of hemelytron 0.95–1.00 mm.</p><p>Female. General habitus very similar to that of male (Figs 2 C, 4 C, 6 C, 8 C, 10 C, 13 C) except for the following characters: body wider than in male; antennal segments III and IV shorter than in male; hemelytron usually wider than in male; and apical part of abdomen pentagonal in ventral view.</p><p>Measurements (n = 20). Body length with hemelytra 3.15–3.35 mm; maximum width of body across hemelytra 1.85–1.95 mm; length of antennal segments I to IV 0.20 mm, 0.10 mm, 1.10 mm, and 0.60 mm, respectively; pronotal length 1.15–1.20 mm; pronotal width across paranota 0.80–0.85 mm; hemelytral length 2.45–2.55 mm; maximum width of hemelytron 1.00– 1.05 mm.</p><p>Remarks.</p><p>Among all the species of Omoplax, O. hisasuei sp. nov. strongly resembles O. majorcarinae and O. mukojimensis in terms of its general habitus. However, based on a comparison of the type material of the new species together with the holotype or non-type material and the original descriptions (Guilbert 2001; Souma 2022 a) of O. majorcarinae and O. mukojimensis, three main characteristics were recognized to easily differentiate O. hisasuei sp. nov. from O. majorcarinae and O. mukojimensis (Figs 3 C, G, 4 C, G, H, 5 C, G, 6 C, G, H, 7 C, G, 8 C, G, H, 9 C, G, 10 C, G, H, 14 C, G, H): hood less than 0.5 times as wide as maximum width of head across compound eyes (more than 0.5 times as wide as maximum width of head across compound eyes in O. majorcarinae and O. mukojimensis); paranotum with areolae throughout its length (without areolae in middle part, with areolae in remaining parts in O. majorcarinae); and costal area wider than fused subcostal and discoidal areas (narrower than fused subcostal and discoidal areas in O. majorcarinae and O. mukojimensis). Morphological differences between O. hisasuei sp. nov. and the four other Omoplax species are presented in the identification key below.</p><p>Distribution.</p><p>Japan: Ogasawara Islands: Hahajima Group (Hahajima Island) (Fig. 19). Omoplax hisasuei sp. nov. is endemic to Hahajima Island.</p><p>Etymology.</p><p>This new species is named in honor of Yu Hisasue, a Japanese hymenopterist who collected some of the paratypes and has contributed to clarifying the species diversity of insects from the Ogasawara Islands.</p><p>Host plant.</p><p>Only Neolitsea sericea var. aurata ( Lauraceae) (Fig. 17 E), which is also known as “ Kinshokudamo ”, was confirmed as a host plant for Omoplax hisasuei sp. nov. through field and captive observations of adults and nymphs, suggesting the possibility of monophagy for this lace bug species.</p><p>Bionomics.</p><p>Omoplax hisasuei sp. nov. inhabits an evergreen broad-leaved forest with a subtropical climate in the Ogasawara Islands and sucks sap on the abaxial side of the leaves of Neolitsea sericea var. aurata, causing irregular yellowing on the adaxial side (Fig. 17 E). Adults were collected in March, May, and September; a single nymph was collected in September.</p></div>	https://treatment.plazi.org/id/B56AA5BF20905550A354519D2630BF7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Souma, Jun	Souma, Jun (2025): An illustrated key to the lace bugs (Hemiptera, Heteroptera, Tingidae) from “ Oriental Galapagos ” (the Ogasawara Islands, Japan), with descriptions of three new species of the endemic genus Omoplax Horváth, 1912. ZooKeys 1250: 243-284, DOI: 10.3897/zookeys.1250.160064
C031F63A342254E4842AA29F93B04299.text	C031F63A342254E4842AA29F93B04299.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Omoplax Horvath 1912	<div><p>Genus Omoplax Horváth, 1912</p><p>Omoplax Horváth, 1912: 336 (as subgenus of Stephanitis Stål, 1873; upgraded by Takeya 1962: 74). Type species by monotypy: Stephanitis (Omoplax) desecta Horváth, 1912 .</p><p>Note.</p><p>For detailed diagnostic characters of the genus, see Souma and Kamitani (2021) and Souma (2022 a).</p><p>Remarks.</p><p>The genus Omoplax, which is endemic to the Ogasawara Islands, Japan, previously comprised four species (Souma 2022 a); however, in the present study, the author describes three new species. The following seven Omoplax species are currently recognized: O. desecta, O. hisasuei sp. nov., O. inugusu sp. nov., O. karubei, O. kobugashi sp. nov., O. majorcarinae, and O. mukojimensis .</p></div>	https://treatment.plazi.org/id/C031F63A342254E4842AA29F93B04299	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Souma, Jun	Souma, Jun (2025): An illustrated key to the lace bugs (Hemiptera, Heteroptera, Tingidae) from “ Oriental Galapagos ” (the Ogasawara Islands, Japan), with descriptions of three new species of the endemic genus Omoplax Horváth, 1912. ZooKeys 1250: 243-284, DOI: 10.3897/zookeys.1250.160064
EB0E928C99185EA5A121E0509E4D7ECD.text	EB0E928C99185EA5A121E0509E4D7ECD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Omoplax inugusu Souma 2025	<div><p>Omoplax inugusu sp. nov.</p><p>Figs 1 D, 2 D, 3 D, 4 D, 5 D, 6 D, 7 D, 8 D, 9 D, 10 D, 11 D, 12 D, 13 D, 14 D, 16 A, B</p><p>Omoplax majorcarinae Guilbert, 2001: Souma and Kamitani (2021: 9) (distribution: part); Souma (2022 a: 126) (distribution: part); Shimamoto and Ishikawa (2023: 94) (catalog: part); Souma (2023: 9) (monograph). Misidentifications.</p><p>Type material.</p><p>Holotype, Japan • ♂; Ogasawara Isls., Hahajima Is., Tamagawa Dam; Machilus boninensis; 1 Oct. 2024; J. Souma leg.; SIHU . Paratypes, Japan • 2 ♂♂ 4 ♀♀; Ogasawara Isls., Hahajima Is., Kitamura; 4 Jun. 1976; T. Nakane leg.; 1 ♂ 3 ♀♀ referring to Souma (2022 a); NSMT • 1 ♂ 5 ♀♀; Ogasawara Isls., Hahajima Is., Mt. Chibusa; 21 Jun. 1994; Y. Kaneko leg.; referring to Souma and Kamitani (2021); TUA • 2 ♀♀; same locality data as for preceding; 7 Jul. 1997; K. Matsumoto leg.; referring to Souma (2022 a); NSMT • 1 ♂ 1 ♀; same locality and collector data as for preceding; 18 Jun. 2001; referring to Souma and Kamitani (2021); TUA • 1 ♂; same locality data as for preceding; 12 Jun. 2024; Y. Hisasue leg.; SIHU • 3 ♀♀; same locality data as for preceding; Machilus boninensis; 2 Oct. 2024; J. Souma leg.; SIHU • 1 ♀; Ogasawara Isls., Hahajima Is., “ 石門 ” [= Sekimon]; 27 Jun. 2009; Japan Forest Technology Association leg.; referring to Souma (2022 a); KPMNH • 2 ♂♂ 2 ♀♀; Ogasawara Isls., Hahajima Is., Mt. Sekimon; alt. 356 m; 22 Jun. 2022; S. Tomura leg.; SIHU • 2 ♀♀; same data as for holotype; SIHU • 1 ♂ 3 ♀♀; same locality data as for holotype; 27 Jun. 1996; K. Morimoto leg.; referring to Souma and Kamitani (2021); ELKU • 2 ♂♂ 4 ♀♀; same locality data as for holotype; 14 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♂; Ogasawara Isls., Hahajima Is., Mt. Kichibe; 16 Jun. 2024; N. Tsuji leg.; SIHU • 1 ♂ 1 ♀; Ogasawara Isls, Hahajima Is., Nishidai; 10 Jun. 2025; N. Tsuji leg.; SIHU .</p><p>Diagnosis.</p><p>Omoplax inugusu sp. nov. is recognized among the other Omoplax species based on a combination of the following characteristics: rostrum reaching middle part of mesosternum (Fig. 11 D); pronotal disc black (Figs 3 D, 4 D, 5 D, 6 D); hood more than 0.5 times as wide as maximum width of head across compound eyes, not reaching apex of clypeus (Fig. 14 D); paranotum with areolae throughout its length; anterior margin of hemelytron not curved downward in apical half (Figs 7 D, 8 D, 9 D, 10 D); subcostal and discoidal areas of hemelytron not united; costal area narrower than combined width of subcostal and discoidal areas; Sc (subcosta) vein of hemelytron visible in apical part of dorsal view; R + M (fused radius and media) vein of hemelytron indistinct, not carinate; and ventral surface of body dark brown to black (Figs 12 D, 13 D).</p><p>Description.</p><p>Male. Markings on dorsum, head, hood, median carina of pronotum, paranotum, calli, posterior process, hemelytron, and ventral surface of body dark brown to black; antenna and legs in various shades of brown; pronotal disc black; compound eye dark red; areolae of pronotum and hemelytron transparent; pubescence on body yellowish (Figs 1 D, 3 D, 5 D, 7 D, 9 D, 11 D, 12 D).</p><p>Body ovate; pubescence on body shorter than radius of compound eye (Figs 1 D, 12 D). Head (Figs 3 D, 5 D) glabrous; pair of frontal spines separated from each other at apices, not reaching apex of clypeus, occasionally reduced; median spine not reaching bases of frontal spines, occasionally reduced; pair of occipital spines not reaching anterior margin of compound eyes, occasionally reduced; antenniferous tubercle obtuse, curved inward, longer than frontal spines; vertex and clypeus smooth. Compound eye round in dorsal view. Antenna densely covered with minute pubescence on segments I to III and long pubescence on segment IV; pubescence on segment IV longer than pubescence on other parts of body; segment I cylindrical, shorter than segment IV; segment II cylindrical, shortest among antennal segments; segment III linear, longest amongst antennal segments; segment IV fusiform. Bucculae closed at anterior ends, with 3 rows of areolae at highest part. Rostrum (Fig. 11 D) reaching middle part of mesosternum.</p><p>Pronotum (Figs 1 D, 3 D, 5 D) glabrous. Pronotal disc coarsely punctate. Hood shorter than median carina of pronotum, higher than median carina, with 4–5 rows of areolae at highest part, more than 0.5 times as wide as maximum width of head across compound eyes, not reaching apex of clypeus, without robust denticles throughout its length; dorsal margin arched; posterior margin extending to anterior part of pronotal disc. Collar not covering compound eye. Median carina straight, extending to apex of posterior process, with 2 rows of areolae at highest part, without robust denticles throughout its length; dorsal margin arched. Calli smooth. Paranotum subvertical, widened posteriad, with a single row of areolae in anterior half and 2–3 rows in posterior half; outer margin gently curved outward throughout its length, without robust denticles throughout its length. Posterior process triangular; apex rounded.</p><p>Hemelytron (Figs 7 D, 9 D) glabrous, extending beyond apex of abdomen; anterior margin not curved downward in apical half; apices close to each other at rest; subcostal and discoidal areas not united; costal area narrower than combined width of subcostal and discoidal areas, with 4–5 rows of areolae at widest part; subcostal area with 3 rows of areolae at widest part; discoidal area with 4 rows of areolae at widest part; sutural area with 5 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; Sc (subcostal) and Hc (hypocosta) veins distinct throughout their respective length; R + M (fused radius and media) and Cu (cubitus) veins indistinct throughout their respective length, not carinate; Sc and R + M veins without robust denticles throughout their respective length; Sc vein distinct in apical part of dorsal view.</p><p>Thoracic pleura smooth in anterior part, coarsely punctate in posterior part. Ostiolar peritreme oblong. Sternal laminae (Fig. 11 D) lower than bucculae; pro- and mesosternal laminae open at both anterior and posterior ends; metasternal laminae as high as mesosternal laminae, open at anterior ends, fused with each other at posterior ends. Legs (Fig. 1 D) smooth, covered with pubescence; femora thickest at middle. Abdomen ellipsoidal. Terminalia (Fig. 12 D) pentagonal in ventral view, covered with pubescence.</p><p>Measurements (n = 14). Body length with hemelytra 2.75–3.10 mm; maximum width of body across hemelytra 1.30–1.55 mm; length of antennal segments I to IV 0.20 mm, 0.10 mm, 1.20 mm, and 0.60 mm, respectively; pronotal length 1.20–1.25 mm; pronotal width across paranota 0.75–0.80 mm; hemelytral length 2.05–2.30 mm; maximum width of hemelytron 0.80–0.90 mm.</p><p>Female. General habitus very similar to that of male (Figs 2 D, 4 D, 6 D, 8 D, 10 D, 13 D) except for the following characters: body usually longer and wider than in male; antennal segments III and IV shorter than in male; hemelytron usually longer and wider than in male; and apical part of abdomen pentagonal in ventral view.</p><p>Measurements (n = 31). Body length with hemelytra 2.90–3.15 mm; maximum width of body across hemelytra 1.55–1.65 mm; length of antennal segments I to IV 0.20 mm, 0.10 mm, 1.10 mm, and 0.50 mm, respectively; pronotal length 1.20–1.30 mm; pronotal width across paranota 0.80–0.85 mm; hemelytral length 2.25–2.45 mm; maximum width of hemelytron 0.85–0.95 mm.</p><p>Remarks.</p><p>In the previous studies (Souma and Kamitani 2021; Souma 2022 a), Omoplax inugusu sp. nov. was identified as O. majorcarinae, but the former differs from the original description and the illustrations of the latter (Guilbert 2001) based on the following characters: body length with hemelytra 2.75–3.15 mm (3.45 mm in type material) (Figs 1 D, 2 D); maximum width of body across hemelytra 1.30–1.65 mm (1.85 mm in type material); pronotal disc black (Figs 3 D, 4 D, 5 D, 6 D); paranotum with areolae throughout its length (Fig. 14 D); anterior margin of hemelytron not curved downward in apical half (Figs 7 D, 8 D, 9 D, 10 D); and Sc (subcosta) vein of hemelytron distinct in apical part of dorsal view. These features were considered intraspecific variations of O. majorcarinae in the previous studies (cf. Souma and Kamitani 2021; Souma 2022 a), but constitute interspecific variation in the present study based on the examination of dozens of specimens which show two morphological species, each respectively collected from different plant species.</p><p>Among all the Omoplax species, O. inugusu sp. nov. strongly resembles O. kobugashi sp. nov. in terms of its general habitus. However, based on a comparison of the type materials of O. inugusu sp. nov. and O. kobugashi sp. nov., three main characteristics were recognized to easily differentiate O. inugusu sp. nov. from O. kobugashi sp. nov. (Figs 3 D, F, 4 D, F, 5 D, F, 6 D, F, 7 D, F, 8 D, F, 9 D, F, 10 D, F, 11 D, F, 14 D, F): rostrum reaching middle part of mesosternum (reaching posterior margin of mesosternum in O. kobugashi sp. nov.); paranotum with areolae throughout its length (without areolae in middle part, with areolae in remaining parts in O. kobugashi sp. nov.); and anterior margin of hemelytron not curved downward in apical half (weakly curved downward in apical half in O. kobugashi sp. nov.). Morphological differences between O. inugusu sp. nov. and the five other Omoplax species are presented in the identification key below.</p><p>Distribution.</p><p>Japan: Ogasawara Islands: Hahajima Group (Hahajima Island) (Fig. 19) (Souma and Kamitani 2021; Souma 2022 a). Omoplax inugusu sp. nov. is endemic to Hahajima Island.</p><p>Etymology.</p><p>The specific epithet is the Japanese plant name “ Munin- inugusu ” [= Machilus boninensis], referring to the host plant of the new species; a noun in apposition.</p><p>Host plant.</p><p>Only Machilus boninensis ( Lauraceae) (Fig. 17 F), which is also known as “ Munin- inugusu ”, was confirmed as a host plant for Omoplax inugusu sp. nov. by the field and captive observations of adults, suggesting the possibility of monophagy for this lace bug species.</p><p>Bionomics.</p><p>Omoplax inugusu sp. nov. inhabits an evergreen broad-leaved forest with a subtropical climate in the Ogasawara Islands (Souma and Kamitani 2021), and sucks sap on the abaxial side of the leaves of Machilus boninensis, causing irregular yellowing on the adaxial side (Fig. 17 F). Adults were collected in June, July, and October (Souma and Kamitani 2021; Souma 2022 a); nymphs are unknown.</p></div>	https://treatment.plazi.org/id/EB0E928C99185EA5A121E0509E4D7ECD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Souma, Jun	Souma, Jun (2025): An illustrated key to the lace bugs (Hemiptera, Heteroptera, Tingidae) from “ Oriental Galapagos ” (the Ogasawara Islands, Japan), with descriptions of three new species of the endemic genus Omoplax Horváth, 1912. ZooKeys 1250: 243-284, DOI: 10.3897/zookeys.1250.160064
38E875D83E1B5BF2A11A862D26ED8A87.text	38E875D83E1B5BF2A11A862D26ED8A87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Omoplax karubei Souma 2022	<div><p>Omoplax karubei Souma, 2022</p><p>Figs 1 E, 2 E, 3 E, 4 E, 5 E, 6 E, 7 E, 8 E, 9 E, 10 E, 11 E, 12 E, 13 E, 14 E</p><p>Omoplax karubei Souma, 2022 a: 118. Holotype: ♂; type locality: Japan • “ 東京都小笠原村, 聟島 ” [= Ogasawara Islands, Mukojima Group, Mukojima Island]; KPMNH.</p><p>References.</p><p>Shimamoto and Ishikawa (2023: 94) (catalog); Souma (2023: 9) (monograph).</p><p>Material examined.</p><p>Non-types, Japan • 2 ♂♂ 2 ♀♀; Ogasawara Isls, Mukojima Is.; Rhaphiolepis indica var. tashiroi; 15–16 Jul. 2024; S. Yagi, M. Kimura &amp; J. - H. Park leg.; SIHU .</p><p>Diagnosis.</p><p>Omoplax karubei is recognized among the other Omoplax species based on a combination of the following characteristics: rostrum reaching posterior margin of metasternum (Fig. 11 E); pronotal disc pale brown (Figs 3 E, 4 E, 5 E, 6 E); hood more than 0.5 times as wide as maximum width of head across compound eyes, reaching beyond apex of clypeus (Fig. 14 E); paranotum with areolae throughout its length; anterior margin of hemelytron not curved downward in apical half (Figs 7 E, 8 E, 9 E, 10 E); subcostal and discoidal areas of hemelytron not united; costal area narrower than combined width of subcostal and discoidal areas; Sc (subcosta) vein of hemelytron distinct in apical part of dorsal view; R + M (fused radius and media) vein of hemelytron distinct, carinate; and ventral surface of body in various shades of brown (Figs 12 E, 13 E).</p><p>Remarks.</p><p>Omoplax karubei can be distinguished from O. mukojimensis, whose distribution range is consistent with that of O. karubei (Souma 2022 a), based on the following six main characters (Figs 3 E, 4 E, H, 5 E, 6 E, H, 7 E, 8 E, H, 9 E, 10 E, H, 11 E, H, 14 E, H): rostrum reaching posterior margin of metasternum (reaching middle part of mesosternum in O. mukojimensis); hood reaching beyond apex of clypeus (not reaching apex of clypeus in O. mukojimensis); anterior margin of hemelytron not curved downward in apical half (strongly curved downward in apical half in O. mukojimensis); subcostal and discoidal areas of hemelytron not united (united in O. mukojimensis); Sc (subcosta) vein of hemelytron distinct in apical part of dorsal view (indistinct in apical part of dorsal view in O. mukojimensis); and R + M (fused radius and media) vein of hemelytron distinct, carinate (indistinct, not carinate in O. mukojimensis). Morphological differences between O. karubei and the five other Omoplax species are presented in the identification key below.</p><p>Distribution.</p><p>Japan: Ogasawara Islands: Mukojima Group (Mukojima Island) (Fig. 19) (Souma 2022 a). Omoplax karubei is endemic to Mukojima Island.</p><p>Host plant.</p><p>Only Rhaphiolepis indica var. tashiroi ( Rosaceae) (Fig. 17 G), which is also known as “ Shimasharinbai ”, was confirmed as a host plant for Omoplax karubei by the field observation of adults, suggesting the possibility of monophagy for this lace bug species.</p><p>Bionomics.</p><p>Omoplax karubei inhabits an evergreen broad-leaved forest with a subtropical climate in the Ogasawara Islands (Souma 2022 a), and sucks sap on the abaxial side of the leaves of Rhaphiolepis indica var. tashiroi, causing irregular yellowing on the adaxial side (Fig. 17 G). Adults were collected in June and July; nymphs are unknown (Souma 2022 a).</p></div>	https://treatment.plazi.org/id/38E875D83E1B5BF2A11A862D26ED8A87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Souma, Jun	Souma, Jun (2025): An illustrated key to the lace bugs (Hemiptera, Heteroptera, Tingidae) from “ Oriental Galapagos ” (the Ogasawara Islands, Japan), with descriptions of three new species of the endemic genus Omoplax Horváth, 1912. ZooKeys 1250: 243-284, DOI: 10.3897/zookeys.1250.160064
718C9CE1C7BF5E0FA66F9390989F29BE.text	718C9CE1C7BF5E0FA66F9390989F29BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Omoplax kobugashi Souma 2025	<div><p>Omoplax kobugashi sp. nov.</p><p>Figs 1 F, 2 F, 3 F, 4 F, 5 F, 6 F, 7 F, 8 F, 9 F, 10 F, 11 F, 12 F, 13 F, 14 F, 16 C – F</p><p>Omoplax majorcarinae Guilbert, 2001: Souma and Kamitani (2021: 9) (distribution: part); Souma (2022 a: 126) (distribution: part); Shimamoto and Ishikawa (2023: 93) (catalog: part); Souma (2023: 9) (monograph). Misidentifications.</p><p>Type material.</p><p>Holotype, Japan • ♂; Ogasawara Isls., Chichijima Is., Mt. Mikazuki; Machilus kobu; 22 Sep. 2024; J. Souma leg.; SIHU . Paratypes, Japan • 1 ♂; Ogasawara Isls., Chichijima Is.; 4 May 1976; Y. Hori leg.; referring to Souma (2022 a); NSMT • 1 ♀; Ogasawara Isls., Chichijima Is., “ 大村 ” [= Omura]; 18 Jun. 1976; Y. Kurosawa leg.; referring to Souma (2022 a); NSMT • 1 ♂ 1 ♀; Ogasawara Isls., Chichijima Is., Tokoyo Falls – Tastumizaki; 29 Jun. 1994; Y. Kaneko leg.; referring to Souma and Kamitani (2021); TUA • 1 ♀; Ogasawara Isls., Chichijima Is., Mt. Chuosan; 30 Jun. 1994; Y. Kaneko leg.; referring to Souma and Kamitani (2021); TUA • 1 ♂ 4 ♀♀; same locality data as for preceding; 27 Jun. 2001; K. Matsumoto leg.; 1 ♂ 2 ♀♀ referring to Souma and Kamitani (2021); TUA • 1 ♀; same locality data as for preceding; 22 Jun. 2024; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls., Chichijima Is., Mt. Tsutsuji; 28 Jul. 1996; T. Kishimoto leg.; referring to Souma (2022 a); NSMT • 1 ♂ 1 ♀; Ogasawara Isls., Chichijima Is., Mt. Yoake; 14 Jun. 1999; K. Morimoto leg.; referring to Souma and Kamitani (2021); ELKU • 1 ♀; Ogasawara Isls., Anijima Is., “ ヤギ 柵手前 ” [= Front of goat fence located at Central plateau]; 8 Jul. 2009; Japan Forest Technology Association leg.; referring to Souma (2022 a); KPMNH • 1 ♂; Ogasawara Isls., Anijima Is., Mt. Mikaeri, Anbu; Malaise trap; 4 Jul. 2014; D. Watabiki leg.; referring to Souma and Kamitani (2021); TUA • 1 ♀; same locality, trap, and collector data as for preceding; 28 Jul. 2014; referring to Souma and Kamitani (2021); TUA • 1 ♀; Ogasawara Isls., Anijima Is., Mt. Omaru; 25 Jun. 2023; Y. Hisasue leg.; SIHU • 1 ♂; Ogasawara Isls., Anijima Is., Mt. Maruyama; 2 Jul. 2024; Y. Hisasue leg.; SIHU • 1 ♂ 1 ♀; Ogasawara Isls., Anijima Is., Southwest foot of Mt. Token; 8 Jul. 2024; N. Tsuji leg.; SIHU • 1 ♂; Ogasawara Isls., Chichijima Is., Mt. Tsuitate; 19 Jul. 2024; Y. Hisasue leg.; SIHU • 1 ♀; same data as for holotype; SIHU • 3 ♂♂ 4 ♀♀; Ogasawara Isls., Anijima Is., Tamana Beach – Mt. Mikaeri; Machilus kobu; 24 Sep. 2024; J. Souma leg.; SIHU .</p><p>Additional material examined.</p><p>Non-types, Japan • 1 fifth instar nymph 1 fourth instar nymph; Ogasawara Isls., Anijima Is., Tamana Beach – Mt. Mikaeri; Machilus kobu; 24 Sep. 2024; J. Souma leg.; SIHU. The two nymphs recorded above are in poor condition and are thus not described in the present study .</p><p>Diagnosis.</p><p>Omoplax kobugashi sp. nov. is recognized among the other Omoplax species based on a combination of the following characteristics: rostrum reaching posterior margin of mesosternum (Fig. 11 F); pronotal disc black (Figs 3 F, 4 F, 5 F, 6 F); hood more than 0.5 times as wide as maximum width of head across compound eyes, not reaching apex of clypeus (Fig. 14 F); paranotum without areolae in middle part, with areolae in remaining parts; anterior margin of hemelytron weakly curved downward in apical half (Figs 7 F, 8 F, 9 F, 10 F); subcostal and discoidal areas of hemelytron not united; costal area narrower than combined width of subcostal and discoidal areas; Sc (subcosta) vein of hemelytron distinct in apical part of dorsal view; R + M (fused radius and media) vein of hemelytron indistinct, not carinate; and ventral surface of body dark brown to black (Figs 12 F, 13 F).</p><p>Description.</p><p>Male. Markings on dorsum, head, hood, median carina of pronotum, paranotum, calli, posterior process, hemelytron, and ventral surface of body dark brown to black; antenna and legs in various shades of brown; pronotal disc black; compound eye dark red; areolae of pronotum and hemelytron transparent; pubescence on body yellowish (Figs 1 F, 3 F, 5 F, 7 F, 9 F, 11 F, 12 F).</p><p>Body ovate; pubescence on body shorter than radius of compound eye (Figs 1 F, 12 F). Head (Figs 3 F, 5 F) glabrous; pair of frontal spines separated from each other at apices, not reaching apex of clypeus, occasionally reduced; median spine not reaching bases of frontal spines, occasionally reduced; pair of occipital spines not reaching anterior margin of compound eyes, occasionally reduced; antenniferous tubercle obtuse, curved inward, longer than frontal spines; vertex and clypeus smooth. Compound eye round in dorsal view. Antenna densely covered with minute pubescence on segments I to III and long pubescence on segment IV; pubescence on segment IV longer than pubescence on other parts of body; segment I cylindrical, shorter than segment IV; segment II cylindrical, shortest among antennal segments; segment III linear, longest amongst antennal segments; segment IV fusiform. Bucculae closed at anterior ends, with 3 rows of areolae at highest part. Rostrum (Fig. 11 F) reaching posterior margin of mesosternum.</p><p>Pronotum (Figs 1 F, 3 F, 5 F) glabrous. Pronotal disc coarsely punctate. Hood shorter than median carina of pronotum, higher than median carina, with 4–5 rows of areolae at highest part, more than 0.5 times as wide as maximum width of head across compound eyes, not reaching apex of clypeus, without robust denticles throughout its length; dorsal margin arched; posterior margin extending to anterior part of pronotal disc. Collar not covering compound eye. Median carina straight, extending to apex of posterior process, with 2 rows of areolae at highest part, without robust denticles throughout its length; dorsal margin arched. Calli smooth. Paranotum subvertical, widened posteriad, with a single row of areolae in anterior part and 1–2 rows in posterior part, without areolae in middle part; outer margin gently curved outward in posterior part and straight in remaining parts, without robust denticles throughout its length. Posterior process triangular; apex rounded.</p><p>Hemelytron (Figs 7 F, 9 F) glabrous, extending beyond apex of abdomen; anterior margin weakly curved downward in apical half; apices close to each other at rest; subcostal and discoidal areas not united; costal area narrower than combined width of subcostal and discoidal areas, with 4–5 rows of areolae at widest part; subcostal area with 3 rows of areolae at widest part; discoidal area with 4 rows of areolae at widest part; sutural area with 5 rows of areolae at widest part; hypocostal lamina with a single row of areolae throughout its length; Sc (subcostal) and Hc (hypocosta) veins distinct throughout their respective length; R + M (fused radius and media) and Cu (cubitus) veins indistinct throughout their respective length, not carinate; Sc and R + M veins without robust denticles throughout their respective length; Sc vein distinct in apical part of dorsal view.</p><p>Thoracic pleura smooth in anterior part, coarsely punctate in posterior part. Ostiolar peritreme oblong. Sternal laminae (Fig. 11 F) lower than bucculae; pro- and mesosternal laminae open at both anterior and posterior ends; metasternal laminae as high as mesosternal laminae, open at anterior ends, fused with each other at posterior ends. Legs (Fig. 1 F) smooth, covered with pubescence; femora thickest at middle. Abdomen ellipsoidal. Terminalia (Fig. 12 F) pentagonal in ventral view, covered with pubescence.</p><p>Measurements (n = 13). Body length with hemelytra 2.60–2.80 mm; maximum width of body across hemelytra 1.20–1.35 mm; length of antennal segments I to IV 0.20 mm, 0.10 mm, 1.10 mm, and 0.60 mm, respectively; pronotal length 1.15–1.25 mm; pronotal width across paranota 0.70–0.75 mm; hemelytral length 1.95–2.15 mm; maximum width of hemelytron 0.75–0.80 mm.</p><p>Female. General habitus very similar to that of male (Figs 2 F, 4 F, 6 F, 8 F, 10 F, 13 F) except for the following characters: body usually longer and wider than in male; antennal segments III and IV shorter than in male; hemelytron usually longer and wider than in male; and apical part of abdomen pentagonal in ventral view.</p><p>Measurements (n = 21). Body length with hemelytra 2.70–3.05 mm; maximum width of body across hemelytra 1.40–1.50 mm; length of antennal segments I to IV 0.20 mm, 0.10 mm, 1.00 mm, and 0.50 mm, respectively; pronotal length 1.15–1.30 mm; pronotal width across paranota 0.70–0.80 mm; hemelytral length 2.00– 2.25 mm; maximum width of hemelytron 0.80–0.85 mm.</p><p>Remarks.</p><p>In previous studies (Souma and Kamitani 2021; Souma 2022 a), Omoplax kobugashi sp. nov. was identified as O. majorcarinae, but the former differs from the original description and the illustrations of the latter (Guilbert 2001) in the following characters: body length with hemelytra 2.60–3.05 mm (3.45 mm in type material) (Fig. 1 F, 2 F); maximum width of body across hemelytra 1.20–1.50 mm (1.85 mm in type material); rostrum reaching posterior margin of mesosternum (Fig. 11 F); pronotal disc black (Figs 3 F, 4 F, 5 F, 6 F); anterior margin of hemelytron weakly curved downward in apical half (Figs 7 F, 8 F, 9 F, 10 F); and Sc (subcosta) vein of hemelytron distinct in apical part of dorsal view. These features were considered an intraspecific variation of O. majorcarinae in previous studies (cf. Souma and Kamitani 2021; Souma 2022 a), but constitute interspecific variation in the present study based on the examination of dozens of specimens which show two morphological species, each respectively collected from different plant species.</p><p>In general appearance, Omoplax kobugashi sp. nov. is very similar to O. desecta, whose distribution range in Chichijima Group is consistent with that of O. kobugashi sp. nov. (Souma and Kamitani 2021; Souma 2022 a), but the former can be distinguished from the latter based on the rostrum reaching the posterior margin of the mesosternum (reaching the posterior margin of the metasternum in O. desecta) and the black pronotum (pale brown in O. desecta) (Figs 3 B, F, 4 B, F, 5 B, F, 6 B, 11 B, F). Morphological differences between O. kobugashi sp. nov. and the five other Omoplax species are presented in the identification key below.</p><p>Distribution.</p><p>Japan: Ogasawara Islands: Chichijima Group (Anijima Island, Chichijima Island) (Fig. 19) (Souma and Kamitani 2021; Souma 2022 a). Omoplax kobugashi sp. nov. is endemic to Chichijima Group.</p><p>Etymology.</p><p>The specific epithet is the Japanese plant name “ Kobugashi ” [= Machilus kobu], referring to the host plant of the new species; a noun in apposition.</p><p>Host plant.</p><p>Only Machilus kobu ( Lauraceae) (Fig. 17 H), which is also known as “ Kobugashi ”, was confirmed as a host plant for Omoplax kobugashi sp. nov. by the field and captive observations of adults and nymphs, suggesting the possibility of monophagy for this lace bug species.</p><p>Bionomics.</p><p>Omoplax kobugashi sp. nov. inhabits an evergreen broad-leaved forest with a subtropical climate in the Ogasawara Islands (Souma and Kamitani 2021), and sucks sap on the abaxial side of the leaves of Machilus kobu, causing irregular yellowing on the adaxial side (Fig. 17 H). Adults were collected in September and from May to July (Souma and Kamitani 2021; Souma 2022 a); nymphs were collected in September.</p></div>	https://treatment.plazi.org/id/718C9CE1C7BF5E0FA66F9390989F29BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Souma, Jun	Souma, Jun (2025): An illustrated key to the lace bugs (Hemiptera, Heteroptera, Tingidae) from “ Oriental Galapagos ” (the Ogasawara Islands, Japan), with descriptions of three new species of the endemic genus Omoplax Horváth, 1912. ZooKeys 1250: 243-284, DOI: 10.3897/zookeys.1250.160064
F67163155CF95BB9AC49DCA87A0CFF90.text	F67163155CF95BB9AC49DCA87A0CFF90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Omoplax majorcarinae Guilbert 2001	<div><p>Omoplax majorcarinae Guilbert, 2001</p><p>Figs 1 G, 2 G, 3 G, 4 G, 5 G, 6 G, 7 G, 8 G, 9 G, 10 G, 11 G, 12 G, 13 G, 14 G, 16 G – I</p><p>Omoplax majorcarinae Guilbert, 2001: 551. Holotype: ♂; type locality: Japan • Bonin Islands, Chichijima, Chuo san [= Ogasawara Islands, Chichijima Group, Chichijima Island, Mt. Chuosan]; Bernice P. Bishop Museum, Honolulu, Hawaii, USA.</p><p>Omoplax desecta (Horváth, 1912): Souma and Kamitani (2021: 8) (distribution: part); Souma (2022 a: 125) (distribution: part); Shimamoto and Ishikawa (2023: 94) (catalog: part). Misidentifications.</p><p>References.</p><p>Yamada and Tomokuni (2012: 198) (monograph); Yamada and Ishikawa (2016: 432) (checklist: Japan); Shimamoto and Ishikawa (2023: 94) (catalog: part).</p><p>Material examined.</p><p>Non-types, Japan • 1 ♂; Ogasawara Isls., Ototojima Is.; 8 Jul. 1994; Y. Kaneko leg.; referring to Souma and Kamitani (2021); TUA • 3 ♂♂; Ogasawara Isls., Ototojima Is.; 2 Aug. 1996; T. Matsumoto leg.; referring to Souma (2022 a); NSMT • 1 ♂; Ogasawara Isls., Anijima Is., Mt. Togari; 29 Jul. 2021; T. Matsumoto &amp; S. Shimamoto leg.; SIHU • 1 ♀; Ogasawara Isls., Chichijima Is., Tsurihama; 10 Feb. 2024; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls., Chichijima Is., Shigureyama; 9 Mar. 2024; Y. Hisasue leg.; SIHU • 2 ♂♂ 1 ♀; Ogasawara Isls., Chichijima Is., Mt. Mikazuki; 5 May 2024; N. Tsuji leg.; SIHU • 1 ♂; Ogasawara Isls, Chichijima Is., Mt. Mikazuki; 17 May 2024; Y. Hisasue leg.; SIHU • 1 ♀; same locality data as for preceding; 20 Aug. 2024; Y. Hisasue leg.; SIHU • 2 ♀♀; Ogasawara Isls., Ototojima Is., Ainosawa; 3–4 Jul. 2024; N. Tsuji leg.; SIHU • 1 ♂; Ogasawara Isls., Anijima Is., Mt. Maruyama; 9 Jul. 2024; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls., Ototojima Is., Mt. Sokuryogatake; 12 Jul. 2024; Y. Uehara leg.; SIHU • 1 ♀; Ogasawara Isls., Anijima Is., Mt. Omaru; 13 Jul. 2024; Y. Hisasue leg.; SIHU • 2 ♀♀; Ogasawara Isls., Chichijima Is., Hatsuneura; 28 Jul. 2024; N. Tsuji leg.; SIHU • 1 ♂; Ogasawara Isls., Chichijima Is., Mt. Yoake; 4 Aug. 2024; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls., Chichijima Is., Mt. Nyuto; 10 Aug. 2024; Y. Hisasue leg.; SIHU • 2 ♀♀ 1 fifth instar nymph; Ogasawara Isls., Ototojima Is., Kurohama – Ichinotani; Neolitsea sericea var. aurata; 23 Sep. 2024; J. Souma leg.; SIHU • 3 ♂♂ 2 ♀♀; same locality, host plant, and collector data as for preceding; 5 Oct. 2024; SIHU • 2 ♂♂ 2 ♀♀; Ogasawara Isls., Anijima Is., Tamana Beach – Mt. Mikaeri; Neolitsea sericea var. aurata; 24 Sep. 2024; J. Souma leg.; SIHU • 2 ♂♂ 4 ♀♀; Ogasawara Isls, Ototojima Is., Shikahama – Mt. Hirone; 27 May 2025; S. Shimamoto leg.; SIHU • 5 ♂♂ 2 ♀♀; Ogasawara Isls, Nishijima Is.; 7 Jun. 2025; Y. Hisasue leg.; SIHU • 1 ♀; Ogasawara Isls, Ototojima Is., Ichinotani; 9 Jun. 2025; Y. Hisasue leg.; SIHU . The single nymph recorded above is in poor condition and is thus not described in the present study.</p><p>Diagnosis.</p><p>Omoplax majorcarinae is recognized among the other Omoplax species based on a combination of the following characteristics: rostrum reaching middle part of mesosternum (Fig. 11 G); pronotal disc pale brown (Figs 3 G, 4 G, 5 G, 6 G); hood more than 0.5 times as wide as maximum width of head across compound eyes, not reaching apex of clypeus (Fig. 14 G); paranotum without areolae in middle part, with areolae in remaining parts; anterior margin of hemelytron strongly curved downward in apical half (Figs 7 G, 8 G, 9 G, 10 G); subcostal and discoidal areas of hemelytron united; costal area narrower than fused subcostal and discoidal areas; Sc (subcosta) vein of hemelytron indistinct in apical part of dorsal view; R + M (fused radius and media) vein of hemelytron indistinct, not carinate; and ventral surface of body dark brown to black (Figs 12 G, 13 G).</p><p>Remarks.</p><p>The above specimens matched well with the original description and the illustrations of Omoplax majorcarinae (Guilbert 2001) in terms of their morphological characteristics, especially body size, coloration, rostral length, and the shape of the paranotum and hemelytron, which are not consistent with the specimens recorded as O. majorcarinae in the previous studies (Souma and Kamitani 2021; Souma 2022 a): body length with hemelytra 3.10–3.45 mm (3.45 mm in type material) (Figs 1 G, 2 G); maximum width of body across hemelytra 1.65–1.95 mm (1.85 mm in type material); rostrum reaching middle part of mesosternum (Fig. 11 G); pronotal disc pale brown (Figs 3 G, 4 G, 5 G, 6 G); paranotum without areolae in middle part, with areolae in remaining parts (Fig. 14 G); anterior margin of hemelytron strongly curved downward in apical half (Figs 7 G, 8 G, 9 G, 10 G); and Sc (subcosta) vein of hemelytron indistinct in apical part of dorsal view. Therefore, the examined specimens were identified as O. majorcarinae . Morphological differences between O. majorcarinae and the six other Omoplax species are presented in the identification key below.</p><p>Distribution.</p><p>Japan: Ogasawara Islands: Chichijima Group (Anijima Island, Chichijima Island, Nishijima Island, Ototojima Island) (Fig. 19) (Guilbert, 2001; Souma and Kamitani 2021; Souma 2022 a). Omoplax majorcarinae is endemic to Chichijima Group and is newly recorded from Anijima and Nishijima islands.</p><p>Host plant.</p><p>Only Neolitsea sericea var. aurata ( Lauraceae) (Fig. 17 I), which is also known as “ Kinshokudamo ”, was confirmed as a host plant for Omoplax majorcarinae by the field and captive observations of adults and nymphs, suggesting the possibility of monophagy for this lace bug species. However, no feeding behavior of O. majorcarinae was observed on Cinnamomum sp. ( Lauraceae) or Ligustrum sp. ( Oleaceae), from which only a single adult was collected in a previous study (Guilbert 2001). Therefore, these two tree species do not appear to be host plants for this lace bug species.</p><p>Bionomics.</p><p>Omoplax majorcarinae inhabits an evergreen broad-leaved forest with a subtropical climate in the Ogasawara Islands (Souma and Kamitani 2021) and sucks sap on the abaxial side of the leaves of Neolitsea sericea var. aurata, causing irregular yellowing on the adaxial side (Fig. 17 I). Adults were collected in February, March, and from May to October (Guilbert 2001; Souma and Kamitani 2021; Souma 2022 a); a single nymph was collected in September.</p></div>	https://treatment.plazi.org/id/F67163155CF95BB9AC49DCA87A0CFF90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Souma, Jun	Souma, Jun (2025): An illustrated key to the lace bugs (Hemiptera, Heteroptera, Tingidae) from “ Oriental Galapagos ” (the Ogasawara Islands, Japan), with descriptions of three new species of the endemic genus Omoplax Horváth, 1912. ZooKeys 1250: 243-284, DOI: 10.3897/zookeys.1250.160064
5189B30990465C1396EACE59DA41E20B.text	5189B30990465C1396EACE59DA41E20B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Omoplax mukojimensis Souma 2022	<div><p>Omoplax mukojimensis Souma, 2022</p><p>Figs 2 H, 4 H, 6 H, 8 H, 10 H, 11 H, 13 H, 14 H</p><p>Omoplax mukojimensis Souma, 2022 a: 122 . Holotype: ♀; type locality: Japan • “ 東京都小笠原村聟島南部 ” [= Ogasawara Islands, Mukojima Group, Mukojima Island, Southern part]; KPMNH.</p><p>References.</p><p>Shimamoto and Ishikawa (2023: 94) (catalog); Souma (2023: 9) (monograph).</p><p>Material examined.</p><p>No additional specimens have been collected since the original description (Souma 2022 a).</p><p>Diagnosis.</p><p>Omoplax mukojimensis is recognized among the other Omoplax species based on a combination of the following characteristics: rostrum reaching middle part of mesosternum (Fig. 11 H); pronotal disc pale brown (Figs 4 H, 6 H); hood more than 0.5 times as wide as maximum width of head across compound eyes, not reaching apex of clypeus (Fig. 14 H); paranotum with areolae throughout its length; anterior margin of hemelytron strongly curved downward in apical half (Figs 8 H, 10 H); subcostal and discoidal areas of hemelytron united; costal area narrower than fused subcostal and discoidal areas; Sc (subcosta) vein of hemelytron indistinct in apical part of dorsal view; R + M (fused radius and media) vein of hemelytron indistinct, not carinate; and ventral surface of body dark brown (Figs 13 H).</p><p>Remarks.</p><p>Among all the Omoplax species, O. mukojimensis is most similar to O. majorcarinae in terms of its general habitus; however, the former can be distinguished from the latter based on the paranotum with areolae throughout its length (without areolae in the middle part and with areolae in the remaining parts in O. majorcarinae) (Figs 3 G, 4 G, H, 5 G, 6 G, H, 14 G, H). Morphological differences between O. mukojimensis and the five other Omoplax species are presented in the identification key below.</p><p>Distribution.</p><p>Japan: Ogasawara Islands: Mukojima Group (Mukojima Island) (Fig. 19) (Souma 2022 a). Omoplax mukojimensis is endemic to Mukojima Island.</p><p>Host plant.</p><p>Unknown (Souma 2022 a).</p><p>Bionomics.</p><p>Omoplax mukojimensis inhabits an evergreen broad-leaved forest with a subtropical climate in the Ogasawara Islands (Souma 2022 a). A single adult was collected in April; nymphs are unknown (Souma 2022 a).</p></div>	https://treatment.plazi.org/id/5189B30990465C1396EACE59DA41E20B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Souma, Jun	Souma, Jun (2025): An illustrated key to the lace bugs (Hemiptera, Heteroptera, Tingidae) from “ Oriental Galapagos ” (the Ogasawara Islands, Japan), with descriptions of three new species of the endemic genus Omoplax Horváth, 1912. ZooKeys 1250: 243-284, DOI: 10.3897/zookeys.1250.160064
