identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9D794D0FE553C67137E3FB6A5469F9B5.text	9D794D0FE553C67137E3FB6A5469F9B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aneflomorpha spinifera Heffern & Santos-Silva 2025	<div><p>Aneflomorpha spinifera sp. nov.</p><p>(Figs 1–10)</p><p>Description. Holotype female. Frons, vertex, area behind upper eye lobes, and antennal tubercles both light brown and orangish brown; area behind lower eye lobes and sides of gulamentum between eyes and prothorax light brown; genae, remaining surface of gulamentum orangish except brown anterior margins; ventral mouthparts orangish with irregular brownish areas, except light brown anterior region of mentum and whitish apex of palpomeres; postclypeus mostly light brown laterally, blackish centrally; anteclypeus mostly brownish centrally, dark brown laterally; labrum dark brown anterocentrally, orangish-brown on remaining surface; mandibles mostly orangish brown, gradually reddish brown toward apex and inner surface of dorsal surface, except dark brown innermost region of dorsal surface, inner surface, and apical region; scape and pedicel mostly orangish brown; antennomeres III–XI orangish, slightly darker toward apex of antenna. Pronotum brown, slightly darker anteriorly and posteriorly. Sides of prothorax brown except transverse orangish brown band on anterior third, this band gradually narrowed toward pronotum. Prosternum orangish brown with irregular brownish areas, except brown anterior margin and margins close to procoxal cavities; prosternal process orangish brown except brown margins on anterior half. Ventral surface of mesothorax mostly brown on anterior half, orangish on posterior half, except brown margins of mesoventrite and mesoventral process. Ventral surface of mesothorax mostly orangish brown with large, irregular brownish areas interspersed. Scutellum brown. Elytra mostly light brown on wide inner dorsal surface, orangish on remaining surface, orangish area gradually lighter toward apex. Coxae and trochanters mostly orangish brown. Femora orangish on peduncle and basal region of club, brown on remaining lateral and ventral surfaces and reddish brown on remaining dorsal surface. Tibiae orangish. Tarsomeres I–II orangish except reddish-brown posterocentral region; antennomere III, IV, and basal 2/3 of V reddish brown; remaining surface of V brown. Abdominal ventrites orangish with large, irregular reddish-brown areas.</p><p>Head. Wide central area of frons longitudinally striate-punctate, except almost smooth anterior plate; sides abundantly, moderately coarsely punctate, except subsmooth area close to striate-punctate region; with moderately sparse, bristly pale-yellow setae, except almost glabrous anterior plate and glabrous area close to median groove. Vertex moderately coarsely and abundantly punctate; with moderately abundant, bristly pale-yellow setae not obscuring integument, setae slightly denser close to eyes and between antennal tubercles. Area behind upper eye lobes abundantly, coarsely punctate, except smooth area close to eye; with abundant pale-yellow pubescence not obscuring integument, except glabrous smooth area. Area behind lower eye lobes somewhat rugose, glabrous. Genae moderately abundantly and coarsely punctate, except smooth apex; with pale-yellow pubescence and long, suberect setae of same color interspersed, except smooth, glabrous anterior half. Wide central area of postclypeus abundantly, coarsely punctate; with a few bristly setae, setae longer, yellowish-white centrally, more abundant, pale-yellow laterally. Sides of postclypeus densely microsculptured, glabrous. Labrum sparsely, coarsely punctate posterocentrally, moderately abundantly and finely punctate anterocentrally, smooth on remaining surface; with sparse, long, erect yellowish-brown setae posterocentrally, moderately abundant and short yellowish-brown setae anterocentrally, glabrous on remaining surface. Outer and dorsal surface of mandibles densely, coarsely punctate, except smooth innermost area of dorsal surface, apical quarter, and inferior margin of outer surface; punctate area with sparse whitish pubescence, with one long, erect white seta basally and sparse tuft of long, erect yellowish-brown setae close to smooth apical area; remaining surface glabrous; basal 2/3 of inner surface abundantly, finely, transversely striate, striae more conspicuous on left mandible. Gulamentum smooth, glabrous on posterior third; anterior 2/3 rugose-punctate, with sparse, bristly pale-yellow setae, more abundant, shorter, and decumbent on intermaxillary process, and long, erect setae of same color interspersed. Maxillary palpomere IV and labial palpomere III cuneiform. Apex of antennal tubercles rounded. Distance between upper eye lobes 0.35 times distance between outer margins of eyes; in ventral view, distance between lower eye lobes 0.52 times distance between outer margins of eyes. Antennae 1.1 times elytral length, almost reaching posterior sixth of elytra. Scape moderately abundantly and coarsely punctate, except smooth dorsal apex; with moderately sparse yellowish-white pubescence, pubescence sparser ventrally, except glabrous smooth area; with a few long, erect yellowish-white setae interspersed on apical region of ventral surface. Pedicel with sparse yellowish-white pubescence, almost absent ventrally, and long, erect setae of same color interspersed throughout, erect setae longer ventrally. Antennomeres III–XI with abundant white pubescence not obscuring integument, pubescence shorter and denser toward XI; ventral surface of III–VIII with long, erect whitish setae ventrally, setae gradually sparser toward VIII; dorsal apex of III–X with long, erect whitish setae, setae gradually shorter toward X; inner apex of III (Fig. 4) with long spine with acute apex, as long as 1.40 times apical diameter of antennomere; inner apex of IV (Fig. 7) with long spine with acute apex, as long as 1.25 times apical diameter of antennomere; inner apex of V (Fig. 8) with long spine with acute apex, as long as 0.95 times apical diameter of antennomere; inner apex of VI (Fig. 9) with long spine with acute apex, as long as 0.85 times apical diameter of antennomere; inner apex of VII (Fig. 10) with long spine with acute apex, as long as 0.40 times apical diameter of antennomere; dorsal surface of V–VII slightly longitudinally carinate. Antennal formula (ratio) based on length of antennomere III (excluding apical spines): scape = 0.83; pedicel = 0.19; IV = 0.90; V = 0.95; VI = 0.92; VII = 0.90; VIII = 0.85; IX = 0.85; X = 0.78; XI = 0.95.</p><p>Thorax. Prothorax slightly longer than wide; anterior and posterior constrictions narrow, well marked; sides slightly divergent from anterior constriction to after middle, then slightly convergent toward posterior constriction. Pronotum somewhat rugose, abundantly, coarsely punctate; with moderately sparse pale-yellow pubescence on wide central area, except glabrous anterocentral quarter and narrow, longitudinal central area from about middle to posterior quarter, and abundant pubescence not obscuring integument on posterior fifth; sides with abundant light yellowish-brown pubescence, denser on posterior third; with a few long, erect whitish setae interspersed laterally. Sides of prothorax abundantly, coarsely punctate, with abundant light yellowish-brown pubescence not obscuring integument, pubescence sparser on light integumental area. Prosternum slightly transversely striate; moderately abundantly, finely punctate on posterior 2/3, sparsely punctate on anterior third, especially close to posterior third; posterior 2/3 with abundant yellowish pubescence not obscuring integument, and long, erect whitish setae interspersed; anterior third with sparse yellowish-white pubescence, especially close to posterior 2/3, and long, erect whitish setae interspersed. Prosternal process with sparse whitish pubescence on center of anterior half, pubescence denser on sides of anterior half and center of posterior half, absent on sides of posterior fifth; narrowest area 0.14 times procoxal width. Ventral surface of meso- and metathorax with abundant whitish pubescence not obscuring integument, pubescence denser, more yellowish on mesanepisterna and mesepimeron, except glabrous area of mesanepisterna close to mesoventrite, and sparser close to metathoracic discrimen; with long erect whitish setae interspersed on metanepisterna and sides of metaventrite. Scutellum with dense yellow pubescence, pubescence more yellowish brown close to margins. Elytra. Abundantly, coarsely punctate on anterior third, punctures gradually, finer, sparser, shallower toward apex on remaining surface; with moderately abundant, thick yellowish pubescence not obscuring integument, and long, erect setae of same color interspersed on sides of posterior half and posterior quarter of dorsal surface; apex bispinose, sutural spine longer than outer spine. Legs. Femora with sparse whitish pubescence except abundant pubescence not obscuring integument on apical third and ventral surface of peduncle; with long, erect setae of same color interspersed; apex of sides with rounded projection, projections gradually longer toward metafemora. Tibiae with sparse whitish pubescence, except denser pubescence on apical third of dorsal and inner surface of protibiae and abundant, most yellowish pubescence on almost entire ventral surface of all tibiae; with long, erect whitish setae interspersed; basal region of dorsal surface of protibia longitudinally carinate, carina slightly conspicuous depending on viewing angle. Tarsi with abundant yellowish pubescence not obscuring integument, and long setae of same color directed backward; metatarsomere I slightly shorter than II–III together.</p><p>Abdomen. Ventrites with abundant whitish pubescence not obscuring integument, with long, erect setae of same color interspersed, except glabrous apex of ventrites 1–4; apex of ventrite 5 slightly rounded.</p><p>Dimensions (mm). Total length, 19.80; prothoracic length, 2.80; anterior prothoracic width, 2.25; posterior prothoracic width, 2.35; maximum prothoracic width, 2.60; humeral width, 3.85; elytral length, 14.90.</p><p>Type material. Holotype female from MEXICO, Jalisco: Nevado de Colima, Sayulapa, 2410 m, 21.VII.2012, Cunningham &amp; Nogueira leg. (TAMU, formerly DHCO).</p><p>Etymology. Latin, “spinifera ” (thorny, prickly, spiny); alluding to the spines on the basal antennomeres and on the elytral apex.</p><p>Remarks. The key by Linsley (1963) to species from the United States of America leads Aneflomorpha spinifera sp. nov. to the alternative of couplet “3” with A. aculeata (LeConte, 1873) (see photographs on Bezark (2024) and Lingafelter (2022)). The new species differs from A. aculeata especially by the elytra distinctly longer, slightly longer than 3.5 times the length of the head and prothorax together. In A. aculeata, the elytra are distinctly shorter, about 2.5 times the length of the head and prothorax together.</p><p>The key by Lingafelter (2022) leads Aneflomorpha spinifera to the alternative of couplet “18” with A. yumae Giesbert &amp; Hovore, 1976 (see photographs on Bezark (2024) and Lingafelter (2022)), when the base of the dorsal surface of the protibiae is considered carinate. However, the new species differs by the flagellomeres not distinctly carinate dorsally (carinae well marked in A. yumae), and elytra slightly longer than 5.0 times the prothoracic length (distinctly shorter than 5.0 times in A. yumae). If the base of the dorsal surface of the protibiae is considered not carinate, the key leads the new species to the alternative of couplet “15” with A. seminuda Casey, 1912 (see photographs on Bezark 2024; Lingafelter et al. 2014; and Lingafelter 2022), but they differ especially by the elytra proportionally distinctly shorter in the second.</p><p>The key by Chemsak &amp; Noguera (2005) leads Aneflomorpha spinifera to the alternative of couplet “11” with A. longitudinis Chemsak &amp; Noguera, 2005 (see photographs on Bezark (2024) and in the original description) and A. exilis Chemsak &amp; Noguera, 2005 (see photographs on Bezark (2024) and in the original description). The new species differs from A. longitudinis by the body lighter, especially the antennae (darker in A. longitudinis), prothorax distinctly shorter than the basal width of elytra and about as long as wide (slightly shorter than the basal width of elytra and longer than wide in A. longitudinis). Aneflomorpha spinifera differs from A. exilis especially by the apical spine of the antennomere III almost twice as long as the pedicel (about as long in A. exilis) and the apical spine of the antennomere IV about as wide as the apical width of the antennomere (much shorter in A. exilis). Although A. longitudinis and A. exilis are previously known only by males, usually the differences between males and females in Aneflomorpha are few, only the length of the antennae being more conspicuous.</p><p>Aneflomorpha spinifera is also similar to A. crinita Chemsak &amp; Linsley, 1975 (see photographs on Bezark 2024), but differs by the spine of the antennomere III distinctly longer, and antennomere IV–VI with distinct spine. According to Chemsak &amp; Linsley (1975), in A. crinita the “third segment moderately spined, fourth with a short spine, segments five and six with minute spines.” Finally, it differs from A. rectilinea Casey, 1924 (see photographs on Bezark 2024; Lingafelter et al. 2014; and Lingafelter 2022) especially by the apical spine of the antennomeres V–VI moderately long (very short on V, usually absent on VI in A. rectilinea).</p></div>	https://treatment.plazi.org/id/9D794D0FE553C67137E3FB6A5469F9B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Heffern, Daniel;Santos-Silva, Antonio	Heffern, Daniel, Santos-Silva, Antonio (2025): New species of Elaphidiini (Coleoptera, Cerambycidae, Cerambycinae) from Mexico and Central America, and new records in Cerambycidae and Disteniidae. Zootaxa 5569 (2): 231-252, DOI: 10.11646/zootaxa.5569.2.2, URL: https://doi.org/10.11646/zootaxa.5569.2.2
9D794D0FE556C67237E3F97E5168F81D.text	9D794D0FE556C67237E3F97E5168F81D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aneflomorpha makra Heffern & Santos-Silva 2025	<div><p>Aneflomorpha makra sp. nov.</p><p>(Figs 11–20)</p><p>Description. Holotype male. Head capsule reddish brown, darker centrally from frontal plate to area between antennal tubercles, narrow area close upper eye lobes, area behind lower eye lobes, and apex of antennal tubercles, except orangish-brown ventral surface; postclypeus reddish brown; anteclypeus reddish brown except brown sides and narrow central area close to postclypeus; labrum reddish brown except orangish brown anterocentral region and narrow dark-brown oblique band on each side of anterior 2/3; scape, pedicel, and antennomeres III–IV reddish brown; antennomeres V–XI orangish brown; pronotum and sides of prothorax reddish brown, darker close to anterior and posterior margins; prosternum light reddish brown on posterior third, except dark-brown area close to procoxal cavities, dark reddish brown on anterior fifth, orangish brown on remaining surface. Prosternal process dark brown, except reddish-brown sides on posterior fifth. Ventral surface of meso- and metathorax mostly dark reddish brown except irregular orangish areas and dark-brown area close to coxal cavities. Scutellum dark reddish brown. Elytra orangish brown, gradually lighter toward apex, except narrow dark-brown sutural band on anterior quarter. Femora orangish brown except reddish brown apex. Tibiae orangish except reddish brown base. Tarsi orangish except posterocentral area of tarsomeres I–II reddish brown and tarsomeres V mostly reddish brown. Abdominal ventrites dark reddish brown except some irregular light reddish brown areas.</p><p>Head. Frons abundantly, coarsely punctate, central area somewhat rugose, except frontal plate with moderately fine punctures centrally and smooth laterally; with moderately abundant yellowish pubescence not obscuring integument, except glabrous median groove and sides of frontal plate. Vertex abundantly, coarsely punctate, punctures denser between antennal tubercles and dense and shallower close to prothorax; with moderately abundant yellowish pubescence not obscuring integument; with a few long, erect yellowish setae close to eyes. Area behind upper eye lobes abundantly, coarsely punctate, punctures partially confluent, except smooth area close to eye; with sparse yellowish pubescence superiorly, absent inferiorly and close to eye. Area behind lower eye lobes shallowly rugose-punctate, except smooth area close to eye, glabrous. Genae moderately finely punctate near eye, moderately coarsely punctate on wide central area, smooth apically; with abundant yellowish pubescence not obscuring integument, except glabrous apex. Gulamentum smooth, glabrous on posterior half; anterior half coarsely rugose-punctate, with sparse yellowish pubescence and long, erect setae of same color interspersed, except glabrous and smooth intermaxillary process. Antennal tubercles abundantly, moderately coarsely punctate, except smooth apical area; apex elevated, subacute. Maxillary palpomeres IV and labial palpomeres III cuneiform. Basal half of inner surface of mandibles transversely striate, striae moderately well marked. Distance between upper eye lobes 0.31 times distance between outer margins of eyes; in ventral view, distance between lower eye lobes 0.48 times distance between outer margins of eyes. Antennae 1.45 times elytral length, reaching elytral apex at base of antennomere XI. Scape abundantly, coarsely punctate except smooth dorsal apex; with moderately abundant yellowish pubescence not obscuring integument, except glabrous smooth area, and long, erect setae of same color interspersed, especially ventrally. Pedicel with abundant yellowish pubescence not obscuring integument and long, erect setae of same color interspersed, erect setae more abundant ventrally. Antennomeres III–IV with abundant yellowish pubescence not obscuring integument, except glabrous apex, apical spine, and longitudinal carina; with long, erect yellowish setae interspersed ventral and dorsal area near apex, erect setae distinctly more abundant ventrally; longitudinal carina moderately well marked, from base to apical quarter; inner apex of III (Fig. 14) with long spine with acute apex, as long as 1.35 times apical diameter of antennomere; inner apex of IV (Fig. 17) with long spine with acute apex, as long as 1.2 times apical diameter of antennomere. Antennomeres V–XI with yellowish pubescence not obscuring integument, denser than on III–IV, except glabrous longitudinal carina on V–VIII; V–VI with long, erect yellowish setae interspersed ventrally; V–X with a few long, erect yellowish setae on dorsal apex; inner apex of V (Fig. 18) with moderately long spine with acute apex, as long as 0.68 times apical diameter of antennomere; inner apex of VI (Fig. 19) with moderately long spine with acute apex, as long as 0.72 times apical diameter of antennomere; inner apex of VII (Fig. 20) with short spine with acute apex, 0.34 times apical diameter of antennomere; dorsal longitudinal carina on V–VIII slightly elevated, well marked. Antennal formula (ratio) based on length of antennomere III (excluding apical spines): scape = 0.79; pedicel = 0.18; IV = 0.86; V = 0.90; VI = 1.00; VII = 1.04; VIII = 1.04; IX = 1.04; X = 1.00; XI = 1.44.</p><p>Thorax. Anterior and posterior constriction well marked; sides distinctly widened close to anterior constriction, parallel-sided toward posterior third, distinctly narrowed on beginning of posterior third, then slightly widened toward posterolateral angles. Pronotum coarsely rugose-punctate, except smooth central area from about middle to posterior quarter; with abundant yellowish pubescence not obscuring integument, except dense pubescence close to anterior margin and on posterior sixth, especially laterally, and glabrous anterior sulcus and smooth central area; with a few long, erect yellowish setae laterally. Sides of prothorax abundantly rugose-punctate; with abundant yellowish pubescence not obscuring integument, except glabrous anterior sulcus; with a few long, erect yellowish setae interspersed, erect setae distinctly more abundant near anterior margin, especially close to prosternum. Posterior 2/3 of prosternum moderately abundantly, coarsely punctate, with abundant yellowish pubescence not obscuring integument, pubescence gradually sparser toward center, and a few long, erect setae of same color interspersed; anterior third moderately abundantly and finely punctate close to anterior margin, finely, transversely striate-punctate close to posterior 2/3; with sparse yellowish pubescence, sparser on striate area, and a few long, erect yellowish setae interspersed. Prosternal process with dense yellowish-white pubescence laterally on anterior 3/4, glabrous centrally from base to apex, glabrous laterally on posterior quarter, and with dense yellowish-white pubescence between glabrous areas on posterior quarter; narrowest area 0.19 times procoxal width. Ventral surface of meso- and metathorax with abundant yellowish pubescence, pubescence distinctly denser on mesanepisterna, mesepimera, metanepisterna, and large C-shaped macula on sides of posterior half of metaventrite, except glabrous area close to metathoracic discrimen; with long, erect yellowish setae interspersed on metaventrite. Sides of mesoventral process convergent from base to middle, slightly, gradually widened toward posterior fifth, then abruptly, strongly widened; posterior margin strongly notched centrally; narrowest area 0.34 times mesocoxal width. Scutellum with dense pale-yellow pubescence. Elytra. Abundantly, coarsely punctate on anterior third, punctures gradually finer, sparser toward apex on posterior 2/3; with abundant, slender yellowish pubescence not obscuring integument; with long, erect yellowish setae near epipleural margin on posterior third and dorsal surface on posterior seventh; apex with two long spines with similar length. Legs. Femora with moderately abundant yellowish pubescence not obscuring integument, pubescence sparser on inner surface of basal third, and long, erect setae of same color interspersed; inner and outer apices of profemora with rounded projection; inner and outer apices of meso- and metafemora with triangular projection. Tibiae longitudinally carinate laterally, carinae gradually becoming absent toward apex; base of dorsal surface of protibiae longitudinally carinate; protibiae almost glabrous basal third on inner surface, with sparse yellowish pubescence on dorsal and outer surface, sparse pubescence on central third of inner surface, and abundant yellowish pubescence on apical third of inner surface and almost entire ventral surface, pubescence bristly ventrally, with long, erect yellowish setae interspersed, especially on apical half of dorsal surface and entire outer surface; meso- and metatibiae with sparse yellowish pubescence, except denser, bristly pubescence on apical third of ventral surface, with long, erect yellowish setae interspersed. Tarsi with moderately abundant, both short and long, suberect yellowish setae; Metatarsomere I slightly shorter than II–III together.</p><p>Abdomen. Ventrites with abundant yellowish pubescence not obscuring integument, more yellowish-white centrally, pubescence denser laterally on ventrites 1–4, except glabrous apex on ventrites 1–4; with a few long, erect yellowish setae interspersed throughout; apex of ventrite 5 subtruncate.</p><p>Dimensions in mm. Total length, 22.10; prothoracic length, 2.85; anterior prothoracic width, 2.40; posterior prothoracic width, 2.65; maximum prothoracic width, 2.70; humeral width, 4.50; elytral length, 16.45.</p><p>Type material. Holotype male from MEXICO, Oaxaca: Comaltepec, 2650 m, 16.V.1996, G. Nogueira leg. (TAMU, formerly DHCO).</p><p>Etymology. Greek, “makra,” meaning long; alluding to the shape of the body.</p><p>Remarks. The key to species from North America by Linsley (1963) leads Aneflomorpha makra sp. nov. to the alternative of couplet “3” with A. aculeata (LeConte, 1873) (see photographs on Bezark (2024) and Lingafelter (2022)). However, the new species has the elytra notably longer, almost six times prothoracic length (about 4.0 times in A. aculeata).</p><p>The key to species from the United States of America by Lingafelter (2022) leads the new species to the alternative of couplet “19” with A. linearis (LeConte, 1859) (see photographs on Bezark (2024) and Lingafelter (2022)) and A. paralinearis Lingafelter, 2022 (see photographs on Bezark (2024) and Lingafelter (2022)). Aneflomorpha makra differs from both by the elytra proportionally much longer (at most, 5.0 times prothoracic length in A. lineraris and slightly longer than 4.0 times in A. paralinearis), and with apex with two long spines (at most, with short projections in A. linearis and A. paralinearis).</p><p>The key to species from Mexico by Chemsak &amp; Noguera (2005) leads the new species to the alternative of couplet “9” with A. preclara Chemsak &amp; Linsley, 1975 (see photographs on Bezark (2024) and CNC (2024)). However, A. makra differs by the pronotum without very sparse pubescence on wide posterocentral area (very sparse in A. preclara), pedicel and antennomeres with light integumental color (dark in A. preclara), apical spine of the antennomere III distinctly longer than pedicel (shorter in A. preclara), and elytral apex bispinose (not bispinose in A. preclara).</p><p>Aneflomorpha makra sp. nov. is similar to A. spinifera sp. nov. but differs as follows: apex of antennal tubercles subacute; antennomeres III–IV with longitudinal carina more distinct and glabrous, pronotum and elytra lighter; elytral setae shorter and slender; and femoral peduncle almost entirely light in color. In A. spinifera, the apex of antennal tubercles are rounded, antennomeres III–IV with longitudinal carina almost indistinct and not glabrous, elytral setae are distinctly longer and thicker, pronotum and elytra are mostly darker, and the femoral peduncles are mostly dark. The new species differs from A. crinita Chemsak &amp; Linsley, 1975 (see photographs on Bezark 2024) by the pronotal pubescence not dense throughout (dense in A. crinita), and apex of antennomeres III–IV with long spine (both shorter than pedicel in A. crinita, especially on IV). It is also similar to A. yumae Giesbert &amp; Hovore, 1976 (see photographs on Bezark (2024) and Lingafelter (2022)) but differs by the elytra proportionally longer (shorter than 4.0 times the prothoracic length in A. yumae), and elytral apex with two long spines (at most with short spine or dentate in A. yumae); from A. wappesi Chemsak &amp; Noguera, 2005 (see photographs on Bezark 2024 and in the original description) it differs especially by the acute apex of the spine of the antennomere III (blunt in A. wappesi).</p></div>	https://treatment.plazi.org/id/9D794D0FE556C67237E3F97E5168F81D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Heffern, Daniel;Santos-Silva, Antonio	Heffern, Daniel, Santos-Silva, Antonio (2025): New species of Elaphidiini (Coleoptera, Cerambycidae, Cerambycinae) from Mexico and Central America, and new records in Cerambycidae and Disteniidae. Zootaxa 5569 (2): 231-252, DOI: 10.11646/zootaxa.5569.2.2, URL: https://doi.org/10.11646/zootaxa.5569.2.2
9D794D0FE55AC67E37E3FF575163FBD1.text	9D794D0FE55AC67E37E3FF575163FBD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aneflomorpha guatemalana Heffern & Santos-Silva 2025	<div><p>Aneflomorpha guatemalana sp. nov.</p><p>(Figs 21–29)</p><p>Description. Holotype female. Head capsule brown dorsally between apex of antennal tubercles and prothorax and laterally, light reddish brown on frons and ventrally, except orangish brown part of antennal tubercles and genae, area close to clypeus, and irregular, fragmented oblique band on each side of ventral surface; ventral mouthparts mostly dark yellowish brown with irregular brownish areas, except maxillary palpomere IV and labial palpomere III light brown with blackish outer apex; clypeus reddish brown; labrum reddish brown close to anteclypeus, dark brown on margins, orangish brown centrally; mandibles mostly orangish except black apex; scape reddish brown with large, irregular orangish-brown areas interspersed; pedicel reddish brown basally, gradually lighter toward apex; antennomeres III–XI orangish dorsally and laterally, except reddish-brown longitudinal carina, reddish brown ventrally. Pronotum mostly reddish brown with irregular light reddish brown areas. Sides of prothorax reddish brown except orangish-brown area close to anterior constriction, area near posterior margin, and dark-brown anterior margin. Prosternum brown on posterior half, except orangish posterocentral quarter, dark brown on anterior margin, dark reddish brown close to anterior margin, dark orangish on remaining anterior half, except orangish central area close anterior dark reddish brown area. Anterior half of prosternal process brown laterally, orangish centrally; posterior half light yellowish brown. Mesoventrite brown centrally, orangish brown on sides. Mesanepisterna and mesepimeron reddish brown except orangish-brown area close to mesoventrite. Mesoventral process orange, except brown margins. Metanepisterna and sides of metaventrite black; remaining surface of metaventrite brown with irregular light areas. Scutellum brown basally, gradually reddish brown toward apex. Elytra orangish brown, slightly lighter toward apex. Femora reddish brown with irregular orangish-brown areas. Tibiae reddish brown basally, orangish brown on remaining surface. Tarsi reddish brown with irregular yellowish-brown areas. Abdominal ventrites 1–4 black, except dark orangish-brown intercoxal process, reddish brown apex, dark orangish-brown area close to reddish-brown apex on ventrite 1, and orangish area close to reddish-brown apex on ventrites 2–4; ventrite 5 mostly orangish brown on basal quarter, reddish brown on posterior half, and dark brown on remaining surface.</p><p>Head. Frons longitudinally rugose centrally, with moderately abundant minute punctures interspersed, abundantly, finely punctate laterally; with abundant whitish pubescence not obscuring integument, except glabrous median groove; frontal plate transversely rugose-punctate, with moderately sparse whitish pubescence laterally, glabrous centrally. Area between antennal tubercles and eyes moderately abundantly, finely punctate, with abundant whitish pubescence not obscuring integument and a few long, erect setae of same color near eyes; remaining surface of vertex abundantly, moderately coarsely punctate, with sparse, short whitish pubescence. Area behind upper eye lobes smooth on a narrow area close to eye, somewhat rugose-punctate on remaining surface; with sparse whitish pubescence superiorly near eye, glabrous on remaining surface. Area behind lower eye lobes glabrous, mostly smooth. Genae subsmooth close to eye, moderately abundantly, finely punctate on remaining surface, except smooth apex; with moderately sparse whitish pubescence, except glabrous apex and area close to eye. Wide central area of postclypeus with moderately abundant whitish pubescence centrally close to frontal plate, sparse yellowish pubescence centrally close to anteclypeus, glabrous laterally; with one long, erect yellowish-brown seta on each side. Sides of postclypeus glabrous. Labrum with tuft of long, erect yellowish-brown setae on each side close to anteclypeus, sparse, both short and long, yellowish-brown setae centrally on posterior quarter, fringe of moderately short yellowish-brown setae anterocentrally, glabrous on remaining surface. Outer and dorsal surfaces of mandibles densely, coarsely punctate, except smooth apical quarter; punctate area with moderately abundant whitish pubescence not obscuring integument, sparse, long, erect whitish setae interspersed, and tuft of long, erect yellowish-brown setae close to smooth area. Gulamentum smooth, glabrous on posterior 2/3; anterior third transversely striate-punctate, with short and long brownish setae and short and long whitish setae not obscuring integument. Maxillary palpomere IV and labial palpomere III securiform. Antennal tubercles abundantly, finely punctate, except smooth apex; apex moderately elevated, rounded; with abundant whitish pubescence not obscuring integument, except glabrous apex. Distance between upper eye lobes 0.54 times distance between outer margins of eyes; in ventral view, distance between lower eye lobes 0.73 times distance between outer margins of eyes. Antennae 1.05 times elytral length, reaching posterior quarter of elytra. Scape abundantly, coarsely punctate, smoother punctures ventrally, except smooth apex of dorsal surface; with moderately sparser whitish pubescence dorsally and laterally and sparser yellowish pubescence ventrally; with long, erect yellowish-brown setae on posterior third of ventral surface. Pedicel moderately abundantly and finely punctate on basal 3/4, smooth on apical quarter; punctate area with sparse white pubescence and long, erect, both white and yellowish-white setae throughout. Antennomeres III–XI with abundant white pubescence not obscuring integument, pubescence denser from V, absent on longitudinal carina; III–X with long, erect yellowish-white setae interspersed ventrally, setae gradually sparser toward X, present only on apex from IX; dorsal apex of III–IV with a few long, erect yellowish-white setae; inner apex of III (Fig. 26) with moderately long spine with acute apex, as long as 0.90 times apical diameter of antennomere; inner apex of IV (Fig. 27) with short spine with acute apex, as long as 0.60 times apical diameter of antennomere; inner apex of V (Fig. 28) with short spine with acute apex, as long as 0.50 times apical diameter of antennomere; inner apex of VI (Fig. 29) with short spine with acute apex, as long as 0.35 times apical diameter of antennomere; dorsal surface of III–VII longitudinally carinate. Antennal formula (ratio) based on length of antennomere III (excluding apical spines): scape = 0.85; pedicel = 0.25; IV = 0.64; V = 0.68; VI = 0.67; VII = 0.72; VIII = 0.71; IX = 0.71; X = 0.64; XI = 0.81.</p><p>Thorax. Prothorax about as long as wide; anterior and posterior constrictions narrow, well marked; sides rounded, moderately abruptly widened close to anterior constriction, subparallel-sided on remaining anterior quarter, divergent on remaining anterior third, roundly widened between anterior third and posterior quarter, then distinctly constricted toward posterior fifth, which is slightly roundly widened. Pronotum with slightly elevated gibbosity on each side of anterior third, distinctly inclined from posterior fifth; anterior third and sides densely, minutely punctate; posterior fifth abundantly, finely punctate; central area between anterior third and posterior quarter smooth; remaining surface sparsely, finely punctate; with dense whitish pubescence laterally and on posterior fifth, this pubescent area distinctly narrowed centrally; remaining surface with a few short, decumbent whitish setae, except glabrous smooth central area; with a few long, erect yellowish-white setae on sides of smooth central area. Sides of prothorax with abundant whitish pubescence partially obscuring integument on wide central area, subglabrous close to posterior margin, with sparse whitish pubescence on anterior orange area, and moderately sparse, bristly yellowish pubescence close to anterior margin; with long, erect yellowish setae interspersed close to anterior margin. Posterior half of prosternum abundantly, finely punctate, except almost smooth orangish central area, with abundant yellowish-white pubescence not obscuring integument, except glabrous orangish central area; anterior quarter somewhat finely, rugose-punctate, with moderately abundant yellowish pubescence not obscuring integument, with long, erect setae of same color interspersed, except glabrous narrow area close to anterior margin; remaining surface of anterior half mostly smooth, with a few short, decumbent whitish setae. Prosternal process smooth, subglabrous, with slightly rounded apex; narrowest area 0.39 times procoxal width. Mesoventrite with abundant whitish pubescence not obscuring integument, except glabrous area close to mesoventral process. Sides of mesoventral process convergent from base to posterior third, then strongly roundly widened; apex strongly emarginated; with moderately abundant white pubescence not obscuring integument laterally and sparse whitish pubescence centrally; narrowest area 0.63 times mesocoxal width. Mesanepisterna, mesepimera, metanepisterna, and sides and posterior region of metaventrite with dense yellowish-white pubescence; central area of metaventrite with sparse both short and long yellowish-white setae, except glabrous area close to metathoracic discrimen and oblique area on each side of posterior quarter. Scutellum with dense pale-yellow pubescence. Elytra. Moderately abundantly and coarsely punctate on anterior third, punctures gradually finer toward apex on posterior 2/3; with moderately sparse white pubescence; with a few long, erect whitish setae close to epipleural margin and moderately abundant, long, erect whitish setae close to apex; apex truncate, wide, with rounded outer angle and short triangular projection on sutural angle. Legs. Femora with sparse yellowish-white pubescence, denser ventrally on basal fifth; with a few long, erect yellowish setae interspersed; apices of protibiae with rounded projection; apices of meso- and metafemora with subtriangular projection. Tibiae with moderately sparse, both short and long white setae dorsally and laterally, except sides of apical quarter of protibiae with abundant, bristly yellowish-brown pubescence not obscuring integument; almost entire ventral surface of protibiae with dense, bristly yellowish-brown pubescence; almost entire ventral surface of meso- and metatibiae with abundant yellowish-brown pubescence not obscuring integument, pubescence denser apically; dorsal surface of protibiae longitudinally carinate basally, carina slightly conspicuous depending on viewing angle. Tarsi with moderately sparse, both short and long whitish setae, setae more yellowish white on tarsomeres I; metatarsomere I slightly shorter than II–III together.</p><p>Abdomen. Sides of ventrites with dense yellowish-white pubescence; central area with moderately abundant yellowish-white pubescence not obscuring integument, sparser on ventrite 1, except glabrous apex of ventrites 1–4; with sparse, long, erect yellowish setae interspersed, setae more abundant and yellower on ventrite 5; apex of ventrite 5 subtruncate.</p><p>Dimensions (mm). Total length, 19.00; prothoracic length, 2.90; anterior prothoracic width, 2.45; posterior prothoracic width, 2.65; maximum prothoracic width, 3.10; humeral width, 3.80; elytral length, 13.40.</p><p>Type material. Holotype female from GUATEMALA, Santa Rosa: Finca Miramundo, 1.VII.2000, J. Monzon leg. (TAMU, formerly DHCO).</p><p>Etymology. Named for the country of Guatemala where the type was collected.</p><p>Remarks. The key by Linsley (1963) to species from the United States of America leads Aneflomorpha guatemalana sp. nov. to the alternative of couplet “14” with A. subpubescens (LeConte, 1862) (see photographs on Bezark (2024) and Lingafelter (2022)). The new species differs from A. subpubescens by the pronotum not abundantly and coarsely punctate (abundantly and coarsely punctate in A. subpubescens), and the elytral apex wider and not spiniform on outer angle (narrower and spiniform on outer angle in A. subpubescens).</p><p>The key by Lingafelter (2022) leads Aneflomorpha guatemalana to the alternative of couplet “19,” when considering the basal area of the dorsal surface of the protibiae with a longitudinal carina, with A. linearis (LeConte, 1859) (see photographs on Bezark (2024) and Lingafelter (2022)) and A. paralinearis Lingafelter, 1922 (see photographs on Bezark (2024) and Lingafelter (2022)). The new species differs from A. linearis by the pronotum with very sparse pubescence on the wide central area (moderately abundant in A. linearis), outer apical angle of the elytra not dentate (dentate or somewhat spiniform in A. linearis), and the elytral pubescence shorter (longer in A. linearis). Aneflomorpha guatemalana differs from A. paralinearis by the prothorax shorter and distinctly wider (longer and wider in A. paralinearis), wide central area of the pronotum with very sparse pubescence (moderately abundant in A. paralinearis), and the outer apical angle of the elytra not dentate (dentate in A. paralinearis). Considering that there is no longitudinal carina on the base of the dorsal surface of the protibiae, the key leads the new species to the alternative of couplet “17” with A. minuta Chemsak, 1962 and A. crypt a Lingafelter, 2022. However, A. guatemalana differs notably from both by the prothorax distinctly wider (narrower in these two species).</p><p>The key by Chemsak &amp; Noguera (2005) leads Aneflomorpha guatemalana to the alternative of couplet “9” with A. preclara Chemsak &amp; Linsley, 1975 (see photographs on Bezark (2024) and CNC (2024)), and this is the most similar species of Aneflomorpha . The new species differs as follows: head and pronotum distinctly darker than elytra; scape not darkened apically; pedicel and flagellomeres light reddish brown; legs without black areas; and elytral punctures not dense. In A. preclara, the head, pronotum, and elytra have similar color, scape darkened apically, pedicel and flagellomeres blackish, apex of the femora, tibiae, and tarsi blackish, and the elytral punctures are dense.</p></div>	https://treatment.plazi.org/id/9D794D0FE55AC67E37E3FF575163FBD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Heffern, Daniel;Santos-Silva, Antonio	Heffern, Daniel, Santos-Silva, Antonio (2025): New species of Elaphidiini (Coleoptera, Cerambycidae, Cerambycinae) from Mexico and Central America, and new records in Cerambycidae and Disteniidae. Zootaxa 5569 (2): 231-252, DOI: 10.11646/zootaxa.5569.2.2, URL: https://doi.org/10.11646/zootaxa.5569.2.2
9D794D0FE559C67B37E3FBA2549BFA95.text	9D794D0FE559C67B37E3FBA2549BFA95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aneflomorpha gracilenta Heffern & Santos-Silva 2025	<div><p>Aneflomorpha gracilenta sp. nov.</p><p>(Figs 30–40)</p><p>Description. Holotype female (Figs 30–38). Head capsule dark brown, except dark reddish-brown posterior half of ventral surface; anteclypeus dark yellowish brown with irregular dark brown maculae; labrum brown on posterior half, yellowish brown on anterior half; mentum dark brown; remaining surface of ventral mouthparts orangish brown except palpomeres brownish with orangish apex; scape dark brown; pedicel dark brown except orangish-brown apex; antennomeres III–XI brown, slightly darker before V; outer and dorsal surfaces of mandibles dark reddish brown except black inferior margin of outer surface, longitudinal band on outer margin of dorsal surface, and entire apical third. Prothorax dark brown except blackish anterior margin and area of prosternum close to procoxal cavities. Ventral surface of meso- and metathorax dark brown with irregular dark reddish-brown areas. Scutellum dark brown. Elytra brown except dark-brown sutural band and apical spines. Legs mostly brown with irregular dark-brown areas. Abdominal ventrites mostly brown.</p><p>Head. Frons somewhat rugose, abundantly, coarsely, partially confluently punctate, except frontal plate moderately abundantly and coarsely punctate centrally, mostly smooth laterally; with abundant yellowish pubescence not obscuring integument, pubescence sparser centrally, except glabrous smooth areas of frontal plate. Vertex abundantly, coarsely punctate; with abundant yellowish pubescence not obscuring integument; with a few long, erect yellowish setae interspersed. Area behind upper eye lobes superiorly with sculpturing and pubescence as on vertex, except glabrous area close to eye; remaining surface somewhat finely punctate with minute yellowish pubescence not obscuring integument. Area behind lower eye lobes with abundantly yellowish pubescence not obscuring integument close to eye, glabrous on remaining surface. Genae moderately abundant and finely punctate except smooth apex; with moderately abundant yellowish pubescence not obscuring integument, except glabrous apex. Gulamentum smooth, glabrous on posterior half; anterior half coarsely, transversely rugose, with abundant yellowish pubescence not obscuring integument and long, erect setae of same color interspersed.Antennal tubercles abundantly, finely punctate, except smooth apex; apex moderately elevated, rounded; with abundant yellowish pubescence partially obscuring integument, except glabrous apex. Maxillary palpomeres IV and labial palpomeres III securiform. Distance between upper eye lobes 0.32 times distance between outer margins of eyes; in ventral view, distance between lower eye lobes 0.53 times distance between outer margins of eyes. Antennae 1.2 times elytral length, reaching about posterior ninth. Scape abundantly, coarsely punctate except smooth apex; with abundant yellowish pubescence not obscuring integument; with long, erect yellowish-white setae interspersed, setae slightly more abundant ventrally on apical third. Pedicel with abundant yellowish pubescence not obscuring integument, except glabrous apex; with long, erect yellowish setae interspersed on pubescent area. Antennomeres III–XI with abundant yellowish pubescence not obscuring integument, pubescence denser from IV; III–XI with long, erect yellowish setae ventrally, setae gradually sparser toward X; dorsal apex of III–X with long, erect yellowish setae; III–XI without distinct longitudinal carina dorsally; inner apex of III with short spine (Fig. 33) with acute apex, as long as 0.5 times apical diameter of antennomere; inner apex of IV with short spine (Fig. 36) with acute apex, as long as 0.42 times apical diameter of antennomere; inner apex of V (Fig. 37) with spicule with acute apex, as long as 0.29 times apical diameter of antennomere; inner apex of VI (Fig. 38) with spicule with acute apex, as long as 0.13 times apical diameter of antennomere. Antennal formula (ratio) based on length of antennomere III (excluding apical spines): scape = 0.97; pedicel = 0.23; IV = 0.85; V = 1.00; VI = 1.03; VII = 1.09; VIII = 1.06; IX = 1.09; X = 1.03; XI = 1.35.</p><p>Thorax. Prothorax distinctly longer than wide; anterior constriction narrow, well marked; subparallel-sided, slightly roundly widened centrally. Pronotum abundantly, coarsely, confluently punctate on anterior third; remaining surface abundantly, coarsely punctate, with somewhat rugose general appearance, except moderately narrow smooth central region; with longitudinal, dense yellowish-brown pubescent band centrally on posterior third, abundantly, bristly yellowish-brown pubescence close to posterior margin, and longitudinal, moderately dense dull yellowish pubescence band on sides, from posterior margin to anterior margin, pubescent band distinctly widened on posterior third and pubescence gradually sparser toward anterior margin; remaining posterior third with moderately abundant, dull yellowish pubescence not obscuring integument; remaining surface of anterior 2/3 with sparse dull yellowish pubescence, except glabrous smooth central area; with a few long, erect dull yellowish setae throughout. Sides of prothorax abundantly, coarsely, partially confluent punctate, except transversely striate anterior constriction; with moderately abundant yellowish pubescence not obscuring integument, except glabrous anterior constriction. Posterior 2/3 of prosternum with moderately sparse asperate punctures, with abundant yellowish pubescence not obscuring integument, pubescence mostly absent close to procoxal cavities, and long, erect setae of same color interspersed; anterior third finely striate-punctate, with sparse yellowish pubescence, pubescence almost absent close to anterior margin, and with a few long, erect setae of same color interspersed. Sides of prosternal process convergent on anterior 2/3, widened on posterior third; with sparse yellowish pubescence on anterior third and moderately abundant yellowish-brown pubescence on apical third; narrowest area 0.23 times procoxal width. Mesoventrite with abundant yellowish pubescence not obscuring integument, pubescence denser laterally. Mesanepisterna, mesepimera, and most of metanepisterna with abundant yellowish-brown pubescence partially obscuring integument. Sides of metaventrite, anterior quarter, and area close to metacoxae with dense yellowish-brown pubescence, pubescence lighter on anterior quarter; remaining surface with moderately sparse yellowish pubescence; with long, erect yellowish setae interspersed. Sides of mesoventral process slightly divergent on anterior 3/4, distinctly widened on posterior quarter; posterior margin distinctly notched centrally; with abundant yellowish-brown pubescence not obscuring integument; narrowest area 0.48 times mesocoxal width. Scutellum with dense yellowish-brown pubescence. Elytra. Moderately sparsely, coarsely punctate on anterior quarter, punctures gradually finer, sparser toward apex on posterior 3/4; with abundant yellowish pubescence not obscuring integument; with sparse, long, erect, moderately thick setae interspersed, setae yellowish brown basally, gradually paler toward apex, and more abundant on posterior quarter; area close to epipleural margin with long, erect, slender yellowish setae; apex bispinose, outer spine longer than sutural spine. Legs. Femora and tibiae with abundant yellowish pubescence not obscuring integument and long, erect setae of same color interspersed, except ventral surface of protibiae and apical quarter of meso- and metatibiae with dense, bristly yellowish-brown pubescence; base of dorsal surface of protibiae not longitudinally carinate; sides longitudinally carinate from base to about apical quarter, carina on inner surface slightly distinct. Tarsi with moderately abundant, both short and long yellowish setae not obscuring integument, setae more yellowish-brown on protarsomeres; metatarsomere I slightly shorter than II–III together.</p><p>Abdomen. Ventrites with abundant yellowish pubescence not obscuring integument, except glabrous apex on ventrites 1–4; with long, erect yellowish setae interspersed throughout; apex of ventrite 5 subtruncate.</p><p>Male (Figs 39–40). Similar to females, differing: antennae slightly longer, 1.3 times elytral length, almost reaching elytral apex; and prosternum coarsely, abundantly punctate.</p><p>Variation. Antennae mostly dark orangish brown; prothorax with irregular orangish brown areas; elytra mostly orangish brown; ventral surface of prosternum mostly dark orangish brown with irregular blackish areas; ventral surface of meso- and metathorax and abdominal ventrites mostly orangish brown; legs mostly orangish brown.</p><p>Dimensions in mm (holotype female; paratype male; paratype female). Total length, 23.30/20.20/23.60; prothoracic length, 3.70/2.30/3.65; anterior prothoracic width, 2.30/2.10/2.35; posterior prothoracic width, 2.90/2.55/2.90; maximum prothoracic width, 3.15/2.90/3.10; humeral width, 4.60/3.90/4.50; elytral length, 16.30/13.80/16.40.</p><p>Type material. Holotype female from MEXICO, Jalisco: Mixtlan, 1842 m, 22.VII.2012, G. Nogueira leg. (TAMU, formerly DHCO) . Paratypes — 1 male (DHCO), 1 female (MZSP), same data as holotype .</p><p>Etymology. Latin, “gracilenta ” (slender, thin); alluding to the shape of the body.</p><p>Remarks. The key to species from North America by Linsley (1963) leads Aneflomorpha gracilenta sp. nov. to the alternative of couplet “13” with A. seminuda Casey, 1912 . The key to species from the United States of America by Lingafelter (2022) leads the new species to the alternative of couplet “15” also with A. seminuda . However, the new species has the elytra proportionally longer, about 4.5 times prothoracic length (about 4.0 times in A. seminuda), elytra with long, thick erect setae (erect thick setae absent in A. seminuda), elytral punctures abundant (very abundant in A. seminuda), and elytral apex bispinose (bidentate in A. seminuda).</p><p>The key to species from Mexico by Chemsak &amp; Noguera (2005) separate the species into two groups from the alternative of couplet “6:” “Elytra and usually pronotum pale brownish… 7;” “Elytra and pronotum black to dark brown to dark reddish brown… 12.” As the color of elytra and prothorax are variable in A. gracilenta, we followed both alternatives. The first alternative leads the new species to the alternative of couplet “10” but the two options do not allow including the new species. Following the second alternative of “6,” the key leads A. gracilenta to the alternative of couplet “16” with A. puebla Chemsak &amp; Noguera, 2005 . However, the new species differs by the antennae in male not surpassing the elytral apex (distinctly surpassing in A. puebla), and the elytral apex bispinose (oblique, making the sutural angle acute in A. puebla).</p><p>The general appearance of A. gracilenta resembles that in Aneflus poriferus Giesbert, 1993, especially by the presence of contrasting setose punctures on elytra. Aneflus gracilenta differs by the elytral apex bispinose (sutural angle spinose and outer angle unarmed in A. poriferus), and metaventrite without numerous small, round, glabrous spots (present in A. poriferus). Using the key to Mexican species of Aneflus from Chemsak &amp; Linsley (1963), the new species would be included in the alternative of couplet “9” with Aneflus protensus (LeConte, 1858) . However, the prothorax in A. protensus is about as long as wide (distinctly longer than wide in A. gracilenta).</p></div>	https://treatment.plazi.org/id/9D794D0FE559C67B37E3FBA2549BFA95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Heffern, Daniel;Santos-Silva, Antonio	Heffern, Daniel, Santos-Silva, Antonio (2025): New species of Elaphidiini (Coleoptera, Cerambycidae, Cerambycinae) from Mexico and Central America, and new records in Cerambycidae and Disteniidae. Zootaxa 5569 (2): 231-252, DOI: 10.11646/zootaxa.5569.2.2, URL: https://doi.org/10.11646/zootaxa.5569.2.2
9D794D0FE55CC66437E3FA665282F8B1.text	9D794D0FE55CC66437E3FA665282F8B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aneflus macraei Heffern & Santos-Silva 2025	<div><p>Aneflus macraei sp. nov.</p><p>(Figs 41–50)</p><p>Description. Holotype male. Integument mostly dark brown; ventral mouthparts brown except yellowish-brown apex of palpomeres; anterior region of anteclypeus brownish; sides of labrum brownish; antennae dark reddish brown; elytra gradually dark reddish brown toward apex; apex of abdominal ventrites 1–4 brownish, and area close to brownish apex of ventrite 4 orangish brown. The general whitish pubescence appears to be whiter depending on light intensity.</p><p>Head. Frons somewhat rugose, moderately abundantly, finely punctate, with coarser punctures close to clypeus; with abundant whitish pubescence not obscuring integument, except glabrous median groove; frontal plate somewhat rugose-punctate, with moderately sparse whitish pubescence. Area between antennal tubercles and posterior margin of eyes abundantly, shallowly punctate, punctures partially confluent, except deeper and coarser punctures close to eyes, and with abundant whitish pubescence not obscuring integument, except glabrous median groove; area between posterior margin of eyes and prothorax elevated, abundantly, shallowly punctate, punctures mostly anastomosed, and with sparse whitish pubescence. Area behind upper eye lobes with sculpturing and pubescence as on posterior region of vertex, except smooth, glabrous narrow area close to eye. Area behind lower eye lobes smooth, glabrous superiorly and close to eyes, transversely rugose-punctate, with a few minutes whitish setae. Genae abundantly, finely punctate, except smooth apex; with abundant whitish pubescence not obscuring integument, except glabrous smooth area. Gulamentum smooth, glabrous on posterior half; anterior half transversely, coarsely, striate-punctate, with whitish pubescence not obscuring integument and a few long, erect setae of same color interspersed, except smooth, glabrous anterior area of intermaxillary process. Antennal tubercles moderately elevated with apex forming an obtuse angle; abundantly, finely punctate, with coarser punctures interspersed, except smooth apex; with moderately abundant whitish pubescence not obscuring integument, except smooth apex. Maxillary palpomeres IV and labial palpomeres III securiform. Wide central area of postclypeus abundantly, coarsely punctate; with moderately sparse whitish pubescence; with one long, erect yellowish-white seta on each side. Sides of postclypeus smooth, glabrous. Labrum smooth, glabrous close to anteclypeus and anterior margin; central area abundantly, coarsely punctate, with yellowish-white pubescence not obscuring integument centrally and tuft of long, erect yellowish setae laterally; anterior margin with fringe of golden setae. Distance between upper eye lobes 0.37 times distance between outer margins of eyes; in ventral view, distance between lower eye lobes 0.59 times distance between outer margins of eyes. Antennae 1.4 times elytral length, slightly surpassing elytral apex. Scape densely, finely punctate except smooth apex of dorsal surface; with abundant whitish pubescence not obscuring integument, except glabrous smooth area. Pedicel with abundant whitish pubescence not obscuring integument and long, erect yellowish-white setae interspersed on posterior half. Antennomere III with abundant whitish pubescence not obscuring integument, pubescence gradually yellowish toward apex; apical quarter with long, erect yellowish setae interspersed throughout; inner apex (Fig. 46) with long spine with acute apex, about as long as apical diameter of antennomere.Antennomere IV with abundant yellowish pubescence not obscuring integument; ventral surface with long, erect yellowish setae, setae longer apically; apex of dorsal and lateral surfaces with long, erect yellowish setae interspersed; inner apex (Fig. 47) with moderately long spine with acute apex, as long as 0.67 times apical diameter of antennomere. Antennomeres V–XI with abundant yellowish pubescence not obscuring integument; outer surface distinctly flattened; with distinct longitudinal carina dorsally from VII; apex of V–X with short yellowish setae directed backward throughout; inner apex of V (Fig. 48) with moderately short spine with acute apex, as long as 0.43 times apical diameter of antennomere; inner apex of VI (Fig. 49) with short spine with acute apex, as long as 0.35 times apical diameter of antennomere; inner apex of VII (Fig. 50) with spicule with acute apex. Antennal formula (ratio) based on length of antennomere III (excluding apical spines): scape = 0.91; pedicel = 0.27; IV = 0.82; V = 0.95; VI = 1.13; VII = 1.13; VIII = 1.09; IX = 1.09; X = 1.04; XI = 1.29.</p><p>Thorax. Prothorax longer than wide; anterior constriction narrow, well marked; sides slightly rounded from anterior constriction to near posterior sixth, then subparallel-sided toward posterolateral angles. Pronotum densely rugose-punctate except smooth anterior sulcus, narrow area close to posterior margin, and smooth, longitudinal central area from about middle to posterior sixth; sides with dense whitish pubescence, pubescence slightly sparser and yellowish centrally; anterior sulcus and smooth central area glabrous; remaining central region of anterior third with moderately abundant whitish pubescence not obscuring integument; central area of posterior third with U-shaped whitish pubescent macula, remaining surface of central area on posterior third with abundant whitish pubescence not obscuring integument close to posterior margin; remaining surface with sparse whitish pubescence; with a few long, erect yellowish setae interspersed throughout. Sides of prothorax moderately abundantly, coarsely punctate except striate-punctate area close to anterior margin and posterior margins; with abundant whitish pubescence not obscuring integument, pubescence sparser anteriorly, especially close to prosternum, except glabrous anterior sulcus; with long, erect yellowish setae interspersed on superior region of area close to anterior margin. Prosternum transversely striate on anterior third; anterior third with sparse whitish pubescence; with a few long, erect yellowish setae throughout; coarsely rugose-punctate on posterior 2/3, with abundant whitish pubescence not obscuring integument on posterior 2/3, pubescence denser laterally and close to procoxal cavities. Prosternal process with abundant whitish pubescence except glabrous sides of apex; distinctly narrowed centrally, with narrowest area 0.13 times procoxal width. Mesoventrite with moderately abundant whitish pubescence not obscuring integument, pubescence denser on apex of sides, partially absent close to mesanepisterna. Mesanepisterna and mesepimera with abundant yellowish-white pubescence partially obscuring integument on some areas. Sides of mesoventral process convergent from base to posterior, then distinctly widened; apex oblique laterally and central area distinctly notched; sides with abundant whitish pubescence on anterior 3/4; remaining surface with sparse whitish pubescence; narrowest area 0.5 times mesocoxal width. Metanepisterna with abundant whitish pubescence not obscuring integument. Metaventrite with dense whitish pubescence laterally and near metacoxal cavities, abundant whitish pubescence not obscuring integument on center of anterior third; remaining surface with very sparse whitish pubescence, except glabrous area close to metathoracic discrimen on posterior third; with long, suberect yellowish setae interspersed on pubescent region. Scutellum with dense pale-yellow pubescence. Elytra. Abundantly, coarsely punctate on anterior half, punctures gradually finer and sparser toward apex on posterior half; apex bispinose, outer spine longer and curved inward; with abundant whitish pubescence not obscuring integument, pubescence slightly sparser on irregular areas, except sparse white pubescence on oblique, wide band located on dorsal surface of anterior third, starting on humerus and not reaching suture; with sparse, long, erect, thick yellowish-brown setae throughout. Legs. Femora densely, shallowly, coarsely punctate except smooth apex; with abundant whitish pubescence not obscuring integument and sparse, long, erect yellowish-brown setae interspersed. Protibiae with moderately sparse whitish pubescence on basal third of dorsal and lateral surfaces, moderately abundant yellowish-brown pubescence on remaining dorsal and outer surface, abundant, bristly yellowish-brown pubescence on remaining inner surface, and dense, bristly yellowish-brown pubescence on ventral surface; with long, erect yellowish-brown setae interspersed dorsally. Dorsal and lateral surfaces of meso- and metatibiae with moderately sparse whitish pubescence basally, pubescence gradually yellowish brown and slightly denser toward apex; ventral surface with bristly yellowish-brown pubescence, denser apically; with long, erect, thick yellowish-brown setae interspersed. Dorsal surface of tarsi with moderately abundant, both whitish and yellowish-brown pubescence not obscuring integument and long, erect yellowish-brown setae interspersed; Metatarsomere I shorter than II–III together.</p><p>Abdomen. Ventrites with dense whitish pubescence laterally and entire anterior quarter, pubescence distinctly sparser on remaining central area, except glabrous apex of ventrites 1–4; with both short and long, suberect yellowish-brown setae throughout, except glabrous areas on ventrites 1–4, suberect setae more abundant on ventrite 5. Apex of ventrite 5 slightly concave.</p><p>Dimensions in mm. Total length, 20.60; prothoracic length, 3.60; anterior prothoracic width, 2.65; posterior prothoracic width, 3.00; maximum prothoracic width, 3.40; humeral width, 4.20; elytral length, 14.00.</p><p>Type material. Holotype male from MEXICO, Oaxaca: 16 km E Tehuantepec, 12.VII.1992, T.C. MacRae leg. (TAMU, formerly DHCO).</p><p>Etymology. This new species is dedicated to Ted MacRae, friend of the first author and collector of the holotype.</p><p>Remarks. The key by Linsley (1936) leads Aneflus macraei sp. nov. to the alternative of couplet “3” with A. prolixus and A. protensus, currently, both with subspecies: A. prolixus prolixus LeConte, 1873; A. prolixus fisheri Knull, 1934; A. prolixus insoletus Chemsak &amp; Linsley, 1963; A. protensus protensus (LeConte, 1858); and A. protensus pallidus Tyson, 1968 . Aneflus macraei differs from A. prolixus prolixus, A. p. fisheri, and A. p. insoletus especially by the inner apical spine of the antennomere III about as wide as the maximum diameter of the antennomere (distinctly shorter in these three subspecies). It differs from A. protensus protensus by the slender body (stouter in A. p. protensus), and the distance between the upper eye lobes distinctly wider than twice the maximum diameter of the scape (at most as wide as twice the maximum diameter of the scape in A. p. protensus); it differs from A. p. pallidus especially by the longer apical spine of the antennomere III (distinctly shorter than twice the maximum diameter of the scape in A. p. pallidus).</p><p>The key by Chemsak &amp; Linsley (1963) leads Aneflus macraei to the alternative of couplet “13” with A. rugicollis Linsley, 1935 . However, the new species differs from A. rugicollis by the smaller size (about 20 mm) (larger specimens, about 30 mm in A. rugicollis), and the elytral pubescence not forming patches (forming patches in A. rugicollis).</p><p>Aneflus macraei is similar to A. poriferus Giesbert, 1993, but differs as follows: apical spine of the antennomere III is about as long as the maximum diameter of the antennomere; and the outer apical angle of the elytra spiniform. In A. poriferus, the apical spine of the antennomere III is shorter than the maximum diameter of the antennomere, and the outer apical angle of the elytra is unarmed.</p><p>Aneflus macraei also resembles Aneflomorpha sinaloae (Linsley, 1935), a species that apparently would be better placed in Aneflus, but differs by the apical spine of the antennomere III shorter and straight (longer and arched in A. sinaloae), glabrous central area of the pronotum somewhat rugose (smooth in A. sinaloae), and the elytral pubescence denser (distinctly sparser in A. sinaloae).</p></div>	https://treatment.plazi.org/id/9D794D0FE55CC66437E3FA665282F8B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Heffern, Daniel;Santos-Silva, Antonio	Heffern, Daniel, Santos-Silva, Antonio (2025): New species of Elaphidiini (Coleoptera, Cerambycidae, Cerambycinae) from Mexico and Central America, and new records in Cerambycidae and Disteniidae. Zootaxa 5569 (2): 231-252, DOI: 10.11646/zootaxa.5569.2.2, URL: https://doi.org/10.11646/zootaxa.5569.2.2
9D794D0FE540C66737E3FB6554CDFA3F.text	9D794D0FE540C66737E3FB6554CDFA3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ameriphoderes cribricollis (Bates 1892)	<div><p>Ameriphoderes cribricollis (Bates, 1892)</p><p>(Fig. 55)</p><p>Acyphoderes cribricollis Bates, 1892: 160 .</p><p>Remarks. This species was described based on male syntypes from Mexico (Durango). Currently, it is known from Mexico (Sinaloa, Durango, Jalisco, Guerrero, Nayarit, Veracruz, Puebla, Morelos, Oaxaca, and Colima), Guatemala, and Costa Rica (Monné 2024a; Tavakilian &amp; Chevillotte 2023).</p><p>Material examined. NICARAGUA (new country record), Nueva Segovia: Cerro Jesus, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-86.41&amp;materialsCitation.latitude=13.66" title="Search Plazi for locations around (long -86.41/lat 13.66)">Hwy</a> 29, abt 2 km E Mozonte, 13.66°N 86.41°W, 650 m, 1 male, 19–21.V.2017, E. van den Berghe leg. (DHCO) .</p></div>	https://treatment.plazi.org/id/9D794D0FE540C66737E3FB6554CDFA3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Heffern, Daniel;Santos-Silva, Antonio	Heffern, Daniel, Santos-Silva, Antonio (2025): New species of Elaphidiini (Coleoptera, Cerambycidae, Cerambycinae) from Mexico and Central America, and new records in Cerambycidae and Disteniidae. Zootaxa 5569 (2): 231-252, DOI: 10.11646/zootaxa.5569.2.2, URL: https://doi.org/10.11646/zootaxa.5569.2.2
9D794D0FE540C66737E3FE3F51CBFC04.text	9D794D0FE540C66737E3FE3F51CBFC04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Susuacanga marcelae Botero 2015	<div><p>Susuacanga marcelae Botero, 2015</p><p>(Figs 51–54)</p><p>Susuacanga marcelae Botero, 2015: 11 .</p><p>Remarks. This species was described from a single male from Mexico, with no further details. The female differs from male especially by the shape of the scape: elongate, slightly and gradually widened from base to apex, not strongly rugose and not depressed dorsally in female; strongly widened basally and gradually narrowed toward apex and strongly rugose and depressed dorsally in males. Additionally, the antennae are distinctly shorter in the female and the antennomere XI is distinctly shorter than the humeral width (much longer, with the antennomere XI distinctly longer than the humeral width in the male (see photographs on Bezark 2024 and original description)).</p><p>Material examined. MEXICO, Oaxaca (new state record): Metates, 800 m, 1 female, 12–13.III.1992, G. Nogueira leg. (DHCO) .</p></div>	https://treatment.plazi.org/id/9D794D0FE540C66737E3FE3F51CBFC04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Heffern, Daniel;Santos-Silva, Antonio	Heffern, Daniel, Santos-Silva, Antonio (2025): New species of Elaphidiini (Coleoptera, Cerambycidae, Cerambycinae) from Mexico and Central America, and new records in Cerambycidae and Disteniidae. Zootaxa 5569 (2): 231-252, DOI: 10.11646/zootaxa.5569.2.2, URL: https://doi.org/10.11646/zootaxa.5569.2.2
9D794D0FE540C66137E3F8E85333FE81.text	9D794D0FE540C66137E3F8E85333FE81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spinestoloides hefferni Santos-Silva, Wappes & Galileo 2018	<div><p>Spinestoloides hefferni Santos-Silva, Wappes &amp; Galileo, 2018</p><p>(Fig. 56)</p><p>Spinestoloides hefferni Santos-Silva, Wappes &amp; Galileo, 2018: 474 .</p><p>Remarks. This species was described based on a single female from Panama (Chiriquí). The male differs from female by the elytra proportionally shorter, abdominal ventrite 5 not inclined posterocentrally and with the apex distinctly notched centrally.</p><p>Material examined. PANAMA, Bocas del Toro (new department record): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.1909&amp;materialsCitation.latitude=8.7814" title="Search Plazi for locations around (long -82.1909/lat 8.7814)">Fortuna Cabins</a>, 08.7814°N 82.1909°W, 1 male, 27.IV–4.V.2023, B.T. Raber leg. (DHCO) .</p></div>	https://treatment.plazi.org/id/9D794D0FE540C66137E3F8E85333FE81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Heffern, Daniel;Santos-Silva, Antonio	Heffern, Daniel, Santos-Silva, Antonio (2025): New species of Elaphidiini (Coleoptera, Cerambycidae, Cerambycinae) from Mexico and Central America, and new records in Cerambycidae and Disteniidae. Zootaxa 5569 (2): 231-252, DOI: 10.11646/zootaxa.5569.2.2, URL: https://doi.org/10.11646/zootaxa.5569.2.2
9D794D0FE546C66137E3FDAB52B4FC5F.text	9D794D0FE546C66137E3FDAB52B4FC5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Novantinoe solisi Santos-Silva & Hovore 2007	<div><p>Novantinoe solisi Santos-Silva &amp; Hovore, 2007</p><p>(Fig. 57)</p><p>Novantinoe solisi Santos-Silva &amp; Hovore, 2007: 585 .</p><p>Remarks. This species was described from a male and female from Costa Rica. Currently, it is known from Nicaragua and Costa Rica (Monné 2024c; Tavakilian &amp; Chevillotte 2023).</p><p>Material examined. PANAMA (new country record), Bocas del Toro: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.1909&amp;materialsCitation.latitude=8.7814" title="Search Plazi for locations around (long -82.1909/lat 8.7814)">Fortuna Cabins</a>, 08.7814°N 82.1909°W, 1 female, 26.IV–5.V.2023, E.G. Riley leg. (DHCO) .</p></div>	https://treatment.plazi.org/id/9D794D0FE546C66137E3FDAB52B4FC5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Heffern, Daniel;Santos-Silva, Antonio	Heffern, Daniel, Santos-Silva, Antonio (2025): New species of Elaphidiini (Coleoptera, Cerambycidae, Cerambycinae) from Mexico and Central America, and new records in Cerambycidae and Disteniidae. Zootaxa 5569 (2): 231-252, DOI: 10.11646/zootaxa.5569.2.2, URL: https://doi.org/10.11646/zootaxa.5569.2.2
