taxonID	type	description	language	source
9C1F2009FD3C4D230013FBDFB0BED68B.taxon	description	corresponding author e-mail: b. dewilde-duyfjes @ naturalis. nl.	en	Wilde, W. J. J. O. de, Duyfjes, B. E. E. (2019): Three new species of Lythraceae and one new species of Stemonaceae described from Indochina. Blumea 64 (2): 177-182, DOI: 10.3767/blumea.2019.64.02.08, URL: https://doi.org/10.3767/blumea.2019.64.02.08
9C1F2009FD3C4D22035CFCEFB193D961.taxon	etymology	Etymology. The specific epithet honours Eugène Poilane, 1888 – 1964, French botanist and coffee planter in Indochina, who made one of the most valuable and largest botanical collections in that area. Two varieties are recognised. var. poilanei Shrub or treelet 2.5 – 6 m tall, bark peeling (?). Leaves: petiole glabrous, 9 – 12 mm long; lamina thin, glabrous, elliptic, small, 3 – 6 cm long, base short-cuneate, apex subacute or bluntish; intercostal venation reticulate. Inflorescences terminal, paniculate, 3 – 6 - flowered. Flowers: buds ± obconical to obovoid, (sub) glabrous; calyx tube c. 5 mm long, c. 5 mm diam at apex, 6 - ridged, ridges sharp, not winged, running onto the pseudopedicel; pseudopedicel 1 – 2 mm long; auricles absent; calyx lobes 6, reflexed at anthesis, long-triangular, c. 3 by 2 mm, (densely) fine hairy at apex within; petals pinkish, c. 15 mm long including c. 3 mm long claw, margin short crispate-dentate; stamens numerous, radial symmetric, presumably dimorphic (outer stamens longer than numerous inner ones); ovary hairy. Capsules dark brown or black, smooth, not shagreen, glabrous or minutely hairy at apex, 5 - valved, c. 10 mm long, c. 9 mm diam, apex rounded, calyx 6 - ridged, ridges low, running onto the c. 2 mm long fruiting pseudopedicel; calyx lobes glabrous within, finally reflexed. Distribution — Southern Vietnam: Nhatrang, Phanrang. Habitat & Ecology — On poor sandy or rocky soil. Specimens studied. VIETNAM, Khanh Hoa Prov., Poilane 2858, Isl. Tre near Nhatrang, 30 Mar. 1922, fr.; Poilane 3111, Isl. Tre near Nhatrang (Moî village), 22 Apr. 1922, fr. (P 05115298); Poilane 4654, between Nhatrang and Ninh Hoa, 7 km N of Nhatrang, 21 Sept. 1922, fl.; Poilane 4848, Nui Han Heo near Nhatrang, 3 Oct. 1922, fl.; Poilane 6148, Hone Cohé near Ninh Hoa, Nhatrang, 30 Apr. 1923, fr.; Poilane 6836 B, Annam, Peninsula Nui Han Heo, 13 June 1923, fl. (P 05114584); Poilane 6839, Annam, Peninsula Nui Han Heo, 12 June 1936, fr., type; Poilane 9751, Ba Ran, 27 Feb. 1924, fr.; Poilane 9753, Ba Ran, 27 Feb. 1924, fl.; Schmid s. n., Ba Ngoi, 22 Nov. 1961. fr. – Ninh Thuân Prov., Poilane 5586, Annam, Ca-Na, 5 Mar. 1923, fl. Note — With capsule is meant the visible part of the fruit above the calyx tube, the latter hiding the base of the capsule. The pseudopedicel is the narrowed pedicelliform basal part below the calyx tube and belongs to the flower; at base it ar- ticulates with the true pedicel. var. grandis W. J. de Wilde & Duyfjes, var. nov. Differs from the type-variety var. poilanei in larger fruit (visible part of capsule) 14 – 15 mm long vs fruit 8 – 10 mm. — Type: Poilane 8541 (holo P P 04711372), Annam, Prov. Phanrang, 12 Feb. 1923, fr. Etymology. The variety epithet grandis (Latin, grandis = big) refers to the larger fruits. Tree 6 – 12 m tall, bark not recorded. Leaves: petiole glabrous, 10 – 12 mm long; lamina thin, glabrous, (ovate-) elliptic, c. 8 cm long, base (broadly) rounded or short cuneate, apex bluntish; intercostal venation reticulate. Inflorescences (infructescences) terminal, paniculate, 5 – 10 - flowered. Flowers or flower buds unknown. Capsules black, smooth, not shagreen, glabrous (sparse weak minute hairs less than 0.1 mm long excepted), 5 - valved, 14 – 15 mm long, c. 10 mm diam, apex rounded with short beak, calyx 6 - ridged, glabrous, ridges low, hardly 1 mm high, shortly running onto the 2 – 3 mm long pseudopedicel; calyx lobes glabrous within, somewhat reflexed. Distribution — Southern Vietnam (Annam): Nhatrang, Phanrang. Habitat & Ecology — Rocky soil; fruiting in February and September. Specimen studied. VIETNAM, Khang Hoa Prov., Nhatrang, near Banghoï, Fleury in Chevalier 39029 (P P 05114593), 27 – 29 Sept. 1918, fr. Rotala The two new Rotala species concern one species endemic in Laos with yellow flowers, a unique character in the genus, and one species discovered among the material of the widespread Rotala cordata. In the very variable and widespread R. indica some deviating specimens are here described as a new variety of R. indica.	en	Wilde, W. J. J. O. de, Duyfjes, B. E. E. (2019): Three new species of Lythraceae and one new species of Stemonaceae described from Indochina. Blumea 64 (2): 177-182, DOI: 10.3767/blumea.2019.64.02.08, URL: https://doi.org/10.3767/blumea.2019.64.02.08
9C1F2009FD3F4D200013FF56B0D7D2FE.taxon	etymology	Etymology. The variety epithet minima (Latin, minimus = small, very little) refers to the tiny habit of the plant. Plants herbaceous, to 10 cm tall, erect or with few spreading or decumbent branches, not rooting at the nodes. Leaves 4 – 6 mm long, with cartilaginous margin. Inflorescences terminal, spike-like, the flowers solitary in the axils of bracts, as long as the bracts; bracts broadly elliptic, more or less imbricate, c. 3 mm long, about as long as the flowers, at apex broadly rounded or short-acute. Flowers as in the type variety. Capsules opening by 2 valves. Distribution — Vietnam (Annam): Khang Hoa Prov., Nhatrang and vicinity. Habitat & Ecology — Flowering and fruiting in March. Specimens studied. VIETNAM, Khang Hoa Prov., Nhatrang and vicinity, Pham Hoang Ho 5082, 1960, fl., fr.; Robinson 1512, Robinson 1235, Robinson 1530, type, all 11 – 26 Mar. 1911, fl., fr. Note — The flowers are recorded as white (Robinson 1235).	en	Wilde, W. J. J. O. de, Duyfjes, B. E. E. (2019): Three new species of Lythraceae and one new species of Stemonaceae described from Indochina. Blumea 64 (2): 177-182, DOI: 10.3767/blumea.2019.64.02.08, URL: https://doi.org/10.3767/blumea.2019.64.02.08
9C1F2009FD3F4D20035CFED6B1B7D4F1.taxon	etymology	Etymology. The specific epithet protracta (Latin, protractus = drawn out, lengthened) refers to the protruding stamens. Amphibious or terrestrial annual herb, erect or at base decumbent and rooting, 20 – 30 cm long, stem broadly 4 - winged. Leaves decussate, sessile; lamina (narrowly) ovate-elliptic or elliptic or oblong, c. 1 by c. 0.5 cm, base narrowly cordate, apex more-or-less obtuse. Flowers solitary in leaf axils, towards apex of branches forming spikes, with the bracts about twice as long as the flowers, bract apex subacute. Flowers (4 –) 5 - merous, sessile, bracteoles scarious, minute; calyx campanulate, tube c. 1.5 mm long, lobes c. 0.3 mm long, calyx appendages absent; petals rose-purple, persistent, 1 – 1.5 mm long, obtuse; stamens (4 –) 5, inserted near the base of the calyx (but filaments about halfway adherent to the calyx tube when young), filaments long, c. 4 mm long, c. twice the length of the calyx, carrying the anthers far outside the flower; ovary about half as long as the calyx tube, style as long as ovary. Capsule globose, as long as calyx tube, opening by 3 valves. Seeds c. 0.5 mm long. Distribution — Cambodia, Laos and Vietnam. Habitat & Ecology — Moist places; on basalt bedrock; 100 – 500 m altitude; flowering and fruiting in October and November. Specimens studied. CAMBODIA, Kompong Cham Prov., Chup near Kompong Cham, Evrard 720, Nov. 1921 (P 05134365, P 05134366, P 05134367). – LAOS, Prov. Champassak, Poilane 16051, Bassac, near Kuang si, 100 m, 20 Oct. 1928 (P 05134533); Thorel s. n., Expédition du Mékong, Sedone, 1866 – 1868 (P 05134537). – VIETNAM, Cân Tho prov., Vu Van Cuong 1028, 1966 (P 05134526); Daclac Prov., Ban Brieng, Schmid 981, 450 – 500 m, Nov. 1949, type (P 05103027, P 05103028). Note — Rotala protracta deceptively resembles R. cordata in general appearance, both species share conspicuously 4 - winged branches, and most specimens assigned to R. protracta were formerly determined as R. cordata. Cook (1979) apparently did not examine Indochinese specimens of R. cordata in P.	en	Wilde, W. J. J. O. de, Duyfjes, B. E. E. (2019): Three new species of Lythraceae and one new species of Stemonaceae described from Indochina. Blumea 64 (2): 177-182, DOI: 10.3767/blumea.2019.64.02.08, URL: https://doi.org/10.3767/blumea.2019.64.02.08
9C1F2009FD3F4D200013FC06B644D1C8.taxon	etymology	Etymology. The specific epithet saxatilis refers to the locality, as it was found on rocks. Perennial herbs, 5 – 10 cm tall, possibly gregarious, but not tufted or cushion-forming; stems erect, purplish, 4 - angled, solitary, usually unbranched, growing from short, entangled somewhat woody short rhizomes c. 0.5 cm long. Leaves decussate, sessile; lamina with minute cystoliths, (broadly) ovate, 2 – 5 by 1 – 3 mm, base obtuse or cuneate, apex obtuse. Inflorescence a rather lax terminal spike, 1 – 1.5 (– 2) cm long, not or hardly pedunculate, the lower flowers in bracts approaching the size of upper leaves; bracts leaf-like, smaller than foliage leaves, as long as or longer than the flowers, base obtuse; bracteoles 2, lanceolate, c. 1 mm long. Flowers: pedicel 1.5 – 3 mm long; c alyx tube somewhat urceolate, c. 2 mm long; calyx lobes 4, triangular, c. 0.8 mm long; calyx appendages absent; petals 4, yellow (according to the field label, see note), persisting in fruit, obovate, 1 – 1.2 mm long; stamens 4; filaments inserted c. halfway up the calyx tube; anthers c. 0.3 by c. 0.4 mm, included, level or somewhat above the apex of the calyx tube; ovary ovoid, c. 1.2 mm long, style short, 0.2 – 0.3 mm long. Capsules narrowly ellipsoid, c. 2 by c. 1 mm, opening by 2 valves. Seeds oblong, c. 0.5 mm long. Distribution — Laos, Luang Prabang Province, between Dauh Quau and Luong Prabang, where known only from the type. Habitat & Ecology — Growing on rocks along river at 200 m altitude. Note — The flower colour, recorded as yellow, is remarkable as to date only pink, lilac or white flowers have been observed for Rotala species (Cook 1979).	en	Wilde, W. J. J. O. de, Duyfjes, B. E. E. (2019): Three new species of Lythraceae and one new species of Stemonaceae described from Indochina. Blumea 64 (2): 177-182, DOI: 10.3767/blumea.2019.64.02.08, URL: https://doi.org/10.3767/blumea.2019.64.02.08
9C1F2009FD3F4D26035CF99AB1E8D020.taxon	etymology	Etymology. The specific epithet hirtella (Latin, hirtus = hairy. With smallish hairs) refers to the finely hairiness of the plant. Erect, little-branched herb 30 – 40 cm tall (known from one col- lection only), very finely hairy almost all over, except adaxial lamina, hairs c. 0.1 mm long. Leaves alternate; petiole 3.5 – 7.5 cm long; lamina ovate, 11 – 16 by 5.5 – 9 cm, base cordate, apex acute-acuminate; 9 – 13 - veined. Inflorescences axillary, 1 – 4 - flowered, sessile (peduncle absent); bracts c. 5 mm long. Flowers: pedicel 12 – 16 mm long; tepals narrowly ovate to lanceolate, gradually narrowed into acute apex, 10 – 11 by (2 –) 3 mm, (sparingly) hairy abaxially, glabrous within, margin without hairs; stamens c. 12 mm long, c. 0.8 mm wide, abaxially conspicuously squamulose (bran-like) - rugose, filaments c. 1 mm long, anthers c. 8 mm long, ridge separating the thecae c. 0.5 mm high, petaloid outgrowth of the connective c. 5 mm long, additional appendage c. 1 mm long; ovary minute, c. 1.5 mm long. Fruits and seeds not seen. Distribution — Vietnam, Cuc Phuong National Park, only known from the type. Notes — 1. The plant is annotated as being a liana of 60 cm long (but this does not show on the herbarium specimen); the flowers are recorded as violet with light white calyx and the fruits as 8 mm long; additional data about habitat and ecology are not known. 2. The minutely squamulose-rugose surface of the abaxial side of the (dried) stamens is also present in some other species (Inthachub 2008), e. g., in Stemona kerrii, however, less pronounced. Acknowledgement We thank the curators of L and P for allowing us to study their collections. The authors are indebted to Dr Sovanmoly Hul of the Muséum national d’Histoire naturelle, Paris, for her kind help and mediation in funding our study in the Paris museum. Jan van Os (L) prepared the beautiful drawings.	en	Wilde, W. J. J. O. de, Duyfjes, B. E. E. (2019): Three new species of Lythraceae and one new species of Stemonaceae described from Indochina. Blumea 64 (2): 177-182, DOI: 10.3767/blumea.2019.64.02.08, URL: https://doi.org/10.3767/blumea.2019.64.02.08
