identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
933F3CFDC4A05E7DB6895071F4E849CD.text	933F3CFDC4A05E7DB6895071F4E849CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus aipo Balke & Villastrigo & Hájek & Surbakti & Panjaitan & Hendrich 2025	<div><p>Limbodessus aipo sp. nov.</p><p>Figs 1 A, 7 A, 11 A, 13 A, 14, 17 F, G</p><p>Type locality.</p><p>Small peatland pool, along track from Aipomek to Langda, 3,550 –3,600 m [140°05'E, 04°27'S], Papua, Indonesia.</p><p>Type material.</p><p>Holotype, male. “ IRIAN JAYA: Aipomek Area [N Pass] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.08333&amp;materialsCitation.latitude=-4.45" title="Search Plazi for locations around (long 140.08333/lat -4.45)">Aipomek-Langda</a>, 140°05'E 04°27'S, 22. - 24. 8. 1992, 3550–3600 m, leg. M. Balke (20–22) ” [IR 92 # 20-22], “ Holotype Limbodessus aipo sp. nov. Balke et al. 2025 ” [red printed label] (NHMW).</p><p>Paratypes (96): 46 exs., same label data as holotype (NHMW, KSP, ZSM); 37 exs., “ Irian Jaya: Aipomek area, 140°06'E 04°27'S, 27. 8. 1992, 3500 m [S pass] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.1&amp;materialsCitation.latitude=-4.45" title="Search Plazi for locations around (long 140.1/lat -4.45)">Aipomek-Langda</a>, leg. Balke (23) ” ; 13 exs., “ Irian Jaya, ca. 139°47'E 04°19'S [Valley below Mt. Orom, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.78334&amp;materialsCitation.latitude=-4.3166666" title="Search Plazi for locations around (long 139.78334/lat -4.3166666)">trek Kono-Angguruk</a>, 2800 m], 5–6.10. 1993, leg. Balke (29 / 30) ” [IR 93 # 29-30] (NHMW, KSP, ZSM) . All paratypes are provided with a red printed paratype label.</p><p>Measurements.</p><p>Holotype: TL = 1.55 mm; TL-H = 1.50 mm; MW = 0.70 mm. Paratypes: TL = 1.60–1.70 mm; TL-H = 1.50–1.60 mm; MW = 0.65–0.70 mm.</p><p>Description.</p><p>Smaller species, body elongate, appearing somewhat parallel-sided, with slight discontinuity in lateral outline between pronotum and elytron. Cervical or sutural line absent. Pronotal and elytral stria present and distinct, long (Fig. 1 A).</p><p>Color. Body dark brown to blackish. Clypeus and pronotum anteriorly, medially and laterally slightly paler: dark ferruginous to castaneous brown. Antennomeres 1–5 / 6 dark ferruginous, 6–11 darker, blackish; in a few specimens paler however, completely ferruginous. Legs dark ferruginous (Fig. 1 A). Ventral side chiefly black. Prosternum, epipleura and sternite 7 ferruginous.</p><p>Sculpture. Head moderately densely punctate; punctation on clypeus and vertex sparse; with distinct microreticulation of small polygonal meshes. Size of punctures equals diameter of meshes. Pronotum with moderately dense punctation along fore-margin; other parts with sparse punctation. Size of punctures as on head. Elytron with moderately dense and evenly distributed punctation; between punctures with almost invisible larger polygonal meshes. Ventral side with few punctures, shiny; only sternite 7 with distinct microreticulation. Epipleura narrow posteriorly, disappeared at level of sternite 6, sternite 7 with border.</p><p>Structures. Metacoxal lines anteriorly diverging and not reaching posterior margin of metaventrite.</p><p>Male. Pronotum with microreticulation of small polygonal meshes being confined to a medial portion, expanded laterally as far as level of striae. Elytron shiny, no microreticulation visible between punctures but a network of large polygonal meshes. Antenna moniliform due to globular form of antennomeres (Fig. 13 A). Median lobe as in Fig. 7 A, parameres as in Fig. 11 A.</p><p>Female. Pronotum shagreened, dull. Elytron dull, distinctly shagreened between the punctures. Antenna moniliform due to globular form of antennomeres.</p><p>Etymology.</p><p>Dedicated to the people inhabiting parts of the type area, i. e. the Aipo people. The name is a noun in the nominative standing in apposition.</p><p>Distribution.</p><p>Aipomek-Langda and valley below Mt. Orom, trek Kono-Angguruk, in the Central Highlands of Papua (Fig. 14).</p><p>Differential diagnosis.</p><p>Limbodessus aipo sp. nov. belongs to a group of smaller (TL = 1.55–1.90 mm) and almost blackish high alpine species. The new species is close to L. aldente sp. nov., L. elit sp. nov. and L. wilhelmina sp. nov. but can be separated from them by the form of the female antennae, which is moniliform due to globular form of antennomeres (Fig. 13 A), the form of the median lobe (Fig. 7 A, B, C), and the form of the parameres (Fig. 11 A).</p><p>Habitat.</p><p>Small puddles in alpine meadows (Fig. 17 F, G).</p></div>	https://treatment.plazi.org/id/933F3CFDC4A05E7DB6895071F4E849CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
7849C62D7CA05A63BD5D243E08643A99.text	7849C62D7CA05A63BD5D243E08643A99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus aldente Balke & Villastrigo & Hájek & Surbakti & Panjaitan & Hendrich 2025	<div><p>Limbodessus aldente sp. nov.</p><p>Figs 1 C, 12 C, 13 B, 14</p><p>Type locality.</p><p>Small, mossy puddles, Mt. Trikora area, 3,450 m [138°43'E, 04°13'S], West Papua, Indonesia.</p><p>Type material.</p><p>Holotype, female: “ IRIAN JAYA: Trikora Gebiet 19. - 20. 10. 1993, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.71666&amp;materialsCitation.latitude=-4.2166667" title="Search Plazi for locations around (long 138.71666/lat -4.2166667)">Habbema [Lake] and Kali Oue Tal [Valley]</a>, ca. 138°43'E 04°13'S, 3450 m, leg. M. Balke (39) ”, “ Holotype Limbodessus aldente sp. nov. Balke et al. 2025 ” [red printed label] (NHMW).</p><p>Paratypes. 2 females with the same data as the holotype (NHMW, ZSM) . Both paratypes are provided with a red printed paratype label.</p><p>Measurements.</p><p>Holotype: TL = 1.90 mm; TL-H = 1.75 mm; MW = 0.80 mm. Paratypes: TL 1.80–1.90 mm, TL-H = 1.75–1.80 mm; MW = 0.80–0.85 mm.</p><p>Description.</p><p>Rather small, elytra somewhat parallel-sided; body with distinct discontinuity in lateral outline between pronotum and elytron; pronotum cordiform, strongly narrowed posteriorly. Cervical or sutural line absent. Pronotal and elytral stria present and distinct (Fig. 1 C).</p><p>Color. Body chiefly black. Clypeus anteriorly, pronotum anteriorly and laterally slightly paler: dark ferruginous to castaneous brown. Antennomeres 1–5 / 6 dark ferruginous, 6–11 darker, blackish. Legs dark ferruginous (Fig. 1 C). Ventral side chiefly black. Prosternum, epipleura and sternite 7 ferruginous.</p><p>Sculpture. Head densely and evenly punctate; microreticulation indistinct; shiny. Pronotum with moderately dense punctation; discally sparser; microreticulate, also laterally. Elytron with moderately dense and evenly distributed punctation. Ventral side with few punctures, shiny; only sternite 7 with distinct microreticulation.</p><p>Structures. Metacoxal lines anteriorly diverging and not reaching posterior margin of metaventrite.</p><p>Male unknown.</p><p>Female. Elytron dull, with distinct microreticulation between punctures. Antenna stout with some antennomeres being conspicuously modified: 6–8 distinctly dilated, 9–11 slightly dilated (Figs 12 C, 13 B).</p><p>Etymology.</p><p>Named for our friend Alexander Riedel, Munich, Germany, inventor of the “ aldente ” way to prepare keladi, a staple food in West Papua. The name is a noun in the nominative standing in apposition.</p><p>Distribution.</p><p>Only known from the Lake Habbema area, in the Central Highlands of Papua (Fig. 14).</p><p>Differential diagnosis.</p><p>Limbodessus aldente sp. nov. belongs to a group of smaller (TL = 1.55–1.90 mm) and almost blackish high alpine species. The new species is close to L. aipo sp. nov., L. elit sp. nov. and L. wilhelmina sp. nov. but can be separated from them by the form of the female antennae, which is stout with antennomeres being conspicuously modified: 6–8 distinctly dilated, 9–11 slightly dilated (Figs 12 C, 13 B).</p><p>Habitat.</p><p>Small puddles in alpine meadows overgrown with mosses.</p></div>	https://treatment.plazi.org/id/7849C62D7CA05A63BD5D243E08643A99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
D5780EDA3CEF5635B3CC8D9FE3572A9B.text	D5780EDA3CEF5635B3CC8D9FE3572A9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus alexanderi Balke & Hendrich 2015	<div><p>Limbodessus alexanderi Balke &amp; Hendrich, 2015</p><p>Limbodessus alexanderi Balke &amp; Hendrich in Balke et al. 2015: 4 (original description); Hendrich et al. 2020: 590 (key); Surbakti et al. 2023: 417 (descr., key).</p><p>Material examined.</p><p>Paratypes: 10 exs., “ Indonesia, Papua; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.08092&amp;materialsCitation.latitude=-3.6699724" title="Search Plazi for locations around (long 137.08092/lat -3.6699724)">N Sugapa</a> 3000 m, 3°40'11.90"S, 137°4'51.31"E, 27. xii. 1995, Riedel leg. ” ; 4 exs. with green labels: “ M. Balke 4424, 4425, 6423, 6424 ” (MZB, ZSM).</p><p>Morpholog. Habitus: Male (Fig. 1 B) and female (Fig. 2 B).</p><p>Antennae: Male and female (Fig. 13 G).</p><p>Median lobe (Fig. 8 C) and paramere (Figs 10 G, 11 H).</p><p>Distribution.</p><p>Indonesian New Guinea, known only from the type locality at 3,000 m, which is the mountain range north of Sugapa, Papua (Balke et al. 2015) (Fig. 14).</p></div>	https://treatment.plazi.org/id/D5780EDA3CEF5635B3CC8D9FE3572A9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
0535213962545C7D9B737CEF25AEA5BD.text	0535213962545C7D9B737CEF25AEA5BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus baliem Balke & Hendrich 2015	<div><p>Limbodessus baliem Balke &amp; Hendrich, 2015</p><p>Limbodessus alexanderi Balke &amp; Hendrich in Balke et al. 2015: 25 (original description); Hendrich et al. 2020: 590 (key); Surbakti et al. 2023: 420 (key).</p><p>Material examined.</p><p>Paratypes: 15 exs., “ New Guinea <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.96054&amp;materialsCitation.latitude=-4.1046553" title="Search Plazi for locations around (long 138.96054/lat -4.1046553)">Wamena</a> 1600 m; 4°6'16.76"S, 138°57'37.96"E; 31. viii. &amp; 6. ix. 1990; leg. Balke &amp; Hendrich (IR 1) ” (CLH, ZSM) .</p><p>Morphology.</p><p>Habitus: Male (Fig. 5 A) and female (Fig. 5 B).</p><p>Antennae: Male and female (Fig. 12 F, 12 G, 13 I).</p><p>Median lobe (Fig. 9 A) and paramere (Figs 10 H, 11 I).</p><p>Distribution.</p><p>Indonesian New Guinea, known only from the Baliem Valley, which also contains the type locality Wamena (Figs 14, 15E, F).</p></div>	https://treatment.plazi.org/id/0535213962545C7D9B737CEF25AEA5BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
EB7C4B401D005ACAAC1F04C868148A24.text	EB7C4B401D005ACAAC1F04C868148A24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus compactus (Clark 1862)	<div><p>Limbodessus compactus (Clark, 1862)</p><p>Hydroporus compactus Clark, 1862: 421 (original description); Biström 1988: 16 (descr.); Guignot 1939: 53 (N. COMB.).</p><p>Bidessus neoguineensis Régimbart, 1892: 990 (original description); Balke and Satô 1995: 188 (N. SYN.).</p><p>Uvarus tokarensis Satô, 1972: 53 (original description); Balke and Satô 1995: 188 (N. SYN.).</p><p>Limbodessus compactus (Clark, 1862): Hendrich et al. 2020: 590 (key).</p><p>Material examined.</p><p>2 exs., “ Papua New Guina: Central Woitape, 1500 m, i. 2008, 33.178S 15.481E, Posman (PNG 167) ” (ZSM); 5 exs., “ Indonesia: Papua, Merauke, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.6016&amp;materialsCitation.latitude=-14.712466" title="Search Plazi for locations around (long 141.6016/lat -14.712466)">Wasur</a>, pools, 20 m, 15. - 16. x. 2011, 14 42.748S 141 36.096E, UNCEN team ” (Pap 02) (KSP, ZSM) .</p><p>Morphology.</p><p>Habitus: Male (Fig. 5 D).</p><p>Antennae: Male and female (Fig. 13 E).</p><p>Distribution.</p><p>South East Asia and widespread in the Australasian realm. On New Guinea L. compactus is widespread especially in the lowlands, it is absent from the Baliem Valley and altitudes above 2,000 m (Balke et al. 2015).</p></div>	https://treatment.plazi.org/id/EB7C4B401D005ACAAC1F04C868148A24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
4DB4A5C75632581F868EA2B373DBC329.text	4DB4A5C75632581F868EA2B373DBC329.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus elit Balke & Villastrigo & Hájek & Surbakti & Panjaitan & Hendrich 2025	<div><p>Limbodessus elit sp. nov.</p><p>Figs 1 D, 7 B, 10 D, 11 B, 13 C, 14, 17 A – D</p><p>Type locality.</p><p>Mossy puddles in Alpine grassland, Mt. Elit area, Angguruk-Baliem, 3,400 m [139°17'E, 04°14'S], Papua, Indonesia.</p><p>Type material.</p><p>Holotype, male. “ IRIAN JAYA: Angguruk-Baliem, 10. - 11. 10. 1993, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.28334&amp;materialsCitation.latitude=-4.233333" title="Search Plazi for locations around (long 139.28334/lat -4.233333)">Mt. Elit</a>, ca. 139°17'E 04°14'S, 3400 m, leg. M. Balke (33) ”, “ Holotype Limbodessus elit sp. nov. Balke et al. 2025 ” [red printed label] (NHMW).</p><p>Paratypes (30): 30 exs. with the same label data as the holotype (NHMW, KSP, ZSM) . All paratypes are provided with a red printed paratype label.</p><p>Measurements.</p><p>Holotype: TL = 1.80 mm; TL-H = 1.60 mm; MW = 0.80 mm. Paratypes: TL 1.70–1.85 mm; TL-H = 1.60–1.70 mm; MW = 0.80–0.85 mm.</p><p>Description.</p><p>Rather small, body elongate-oval, with distinct discontinuity in lateral outline between pronotum and elytron. Cervical or sutural line absent. Pronotal and elytral stria present and distinct (Fig. 1 D).</p><p>Color. Body dark-brown to blackish. Clypeus anteriorly, pronotum medially and laterally slightly paler: dark ferruginous to castaneous brown. Antennomeres 1–4 dark ferruginous, 5–11 darker, blackish. Legs dark brown; metatibiae and metatarsi blackish, metafemora castaneous (Fig. 1 D). Ventral side chiefly black. Prosternum, epipleura and sternite 7 ferruginous.</p><p>Sculpture. Head moderately densely punctate; punctation on clypeus and apically sparse; with distinct microreticulation. Size of the punctures corresponds with diameter of meshes. Pronotum with moderately dense punctation along fore-margin; other parts with sparse punctation; microreticulation only medially expanded as far laterally as level of striae. Size of punctures as on head, laterally smaller. Elytron with moderately dense and evenly distributed punctation; between punctures with almost invisible larger polygonal meshes; punctures slightly larger than on head and pronotum. Ventral side with few punctures, shiny; only sternite 7 with distinct microreticulation.</p><p>Structures. Metacoxal lines anteriorly diverging and not reaching posterior margin of metaventrite.</p><p>Male. Elytron shiny, without microreticulation between punctures. Antenna stout. Median lobe as in Fig. 7 B and parameres as in Fig. 10 D, B.</p><p>Female. Elytron shiny, without microreticulation between punctures. Antenna stout but without conspicuous modifications (Fig. 13 C).</p><p>Etymology.</p><p>Named after the type locality, Mount Elit, a two-days walk from Wamena in the direction to Pronggoli. The name is a noun in the nominative standing in apposition.</p><p>Distribution.</p><p>Only known from Mount Elit (Fig. 17 A), in the Central Highlands of Papua (Fig. 14).</p><p>Differential diagnosis.</p><p>Limbodessus elit sp. nov. belongs to a group of smaller (TL = 1.55–1.90 mm) and almost blackish high alpine species. The new species is close to L. aipo sp. nov., L. aldente sp. nov. and L. wilhelmina sp. nov. but can be separated from them by the form of the female antennae, which is stout without any conspicuous modifications (Fig. 13 C), the form of the median lobe (Fig. 7 B), and the form of the parameres (Figs 10 D, 11 B).</p><p>Habitat.</p><p>In very shallow water at the edge of small puddles and pools in alpine meadows (Fig. 17 B, C, D).</p></div>	https://treatment.plazi.org/id/4DB4A5C75632581F868EA2B373DBC329	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
FC9DB0CEA58E545484247D29B4FF48A0.text	FC9DB0CEA58E545484247D29B4FF48A0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus habbema Balke & Villastrigo & Hájek & Surbakti & Panjaitan & Hendrich 2025	<div><p>Limbodessus habbema sp. nov.</p><p>Figs 3 A, B, C, 8 D, 10 C, 11 F, 12 D, E, 13 F, 14, 16 C</p><p>Type locality.</p><p>Mossy puddles in Alpine grassland, Lake Habbema area, Angguruk-Baliem, 3,200 m [04 ° 07 ' 77 S, 138 ° 40 ' 77 E], Papua, Indonesia.</p><p>Type material.</p><p>Holotype, male: “ Indonesia, Papua, Lake Habbema, pools, 3200 m, 19. x. 2011, 04.07. 772 S 138.40. 773 E, Balke leg. (PAP 06) ”, “ Holotype Limbodessus habbema sp. nov. Balke et al. 2025 ” [red printed label] (MZB) .</p><p>Paratypes (322): 126 exs., with the same label data as the holotype (NHMW, KSP, ZSM); 70 exs., “ IRIAN JAYA: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.66667&amp;materialsCitation.latitude=-4.15" title="Search Plazi for locations around (long 138.66667/lat -4.15)">Lake Habbema</a>, 17. 10. 1993, 3250 m, ca. 138°40E 04°09'S, leg. M. Balke (37) ” [IR 93 # 37] (NHMW, ZSM) ; 1 ex., “ IRIAN JAYA: Trikora Gebiet 19. - 20. 10. 1993, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.71666&amp;materialsCitation.latitude=-4.2166667" title="Search Plazi for locations around (long 138.71666/lat -4.2166667)">Habbema [Lake] and Kali Oue Tal [Valley]</a>, ca. 138°43'E 04°13'S, 3450 m, leg. M. Balke (39) ” (NHMW) ; 81 exs., “ Papua, Lake Habbema, 3300 m, 24.04, 2006, Tindige leg. ”, 3 specimens with printed green labels “ 3161 ”, “ 3162 ”, “ 3163 ” (ZSM) ; 11 exs., “ Irian Jaya, Wamena E, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.66667&amp;materialsCitation.latitude=-4.116667" title="Search Plazi for locations around (long 138.66667/lat -4.116667)">Habbema lake</a>, 4°07'S 138°40'E, 20. 1. 1999, leg. A. Weigel ” (ZSM) ; 2 exs., “ Indonesia, West Papua, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.64839&amp;materialsCitation.latitude=-4.109111" title="Search Plazi for locations around (long 138.64839/lat -4.109111)">Highland near Lake Habbema</a>, Sphagnum-pools near the road, 3451 m, 4°06'32.8"S 138°38'54.2"E, 09. 06. 2022, N. Wendlandt leg. ” (CNW, ZSM) ; 19 exs., “ Indonesia, West Papua, Lanny Jaya, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.59277&amp;materialsCitation.latitude=-4.0204725" title="Search Plazi for locations around (long 138.59277/lat -4.0204725)">Kecamatan Makki highland</a>, peatland pools, 3013 m, 4°01'13.7"S 138°35'34.0"E, 06. 06. 2022, N. Wendlandt leg. ” (CNW, ZSM) ; 18 exs., “ Indonesia, West Papua, Lanny Jaya, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.6057&amp;materialsCitation.latitude=-4.040278" title="Search Plazi for locations around (long 138.6057/lat -4.040278)">Kecamatan Makki highland</a>, peatland pools, 3170 m, 4°02'25.0"S 138°36'20.5"E, 07. 06. 2022, N. Wendlandt leg. ” (CNW, ZSM) ; 11 exs., “ Indonesia: Papua, Lake Habbema, Phyllodrom leg. 2001 ” (ZSM) ; 48 exs., “ INDONESIA, Papua: Jayawijaya Distr., ca. 30 km W of Wamena, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.67833&amp;materialsCitation.latitude=-4.13" title="Search Plazi for locations around (long 138.67833/lat -4.13)">open wetland nr Habbema lake</a>, 04°07.8'S, 138°40.7'E; 3330 m, J. Hájek &amp; J. Šumpich leg., 5. ii. 2015 ” (NMPC) ; 9 exs., “ Indonesia: Papua, Lake Habbema, 3400 m, viii. 2018, - 4.1460 ° 138.675 °, Sumoked (Pap 77) ” (MZB, ZSM) . All paratypes are provided with a red printed paratype label.</p><p>Measurements.</p><p>Holotype: TL = 2.7 mm; TL-H = 2.5 mm; MW = 1.2 mm. Paratypes: TL 2.25–2.8 mm, TL-H = 2.15–2.65 mm, MW = 1.1–1.2 mm.</p><p>Description.</p><p>Large, body elongate-oval, with distinct discontinuity in lateral outline between pronotum and elytron. Sutural line absent; cervical line absent, but some specimens with a cervical dense row of punctures. Pronotal and elytral stria present and distinct (Fig. 3 A, B, C).</p><p>Color. Head black, clypeus anteriorly paler, dark brown. Pronotum with anterior half, laterally, and basal angles dark ferruginous to dark yellowish, baso-medially darker, dark brown to blackish; in one specimen almost entirely yellowish. Elytron dark brown to dark ferruginous; in some specimens posteriorly with distinct dark yellowish and dark brown color pattern. Antennomeres 1–4 / 5 dark ferruginous, 6–11 darker, blackish. Legs dark ferruginous; mid of metatibiae and metatarsi darker, blackish (Fig. 3 A). Ventral side chiefly black. Prosternum, epipleura and sternite 7 ferruginous.</p><p>Sculpture. Head densely punctate; a cervical dense row of punctures visible, vertex almost impunctate but with distinct microreticulation only. Size of punctures on frons equal of size of meshes of microreticulation but decreases towards clypeus. Pronotum with moderately dense punctation along fore-margin; size of punctures as on frons; otherwise with fine and sparse punctation only. Elytron with moderately dense and evenly distributed punctation that may be faint on antero-sutural part. Ventral side with few punctures, shiny; only sternite 7 with distinct microreticulation.</p><p>Structures. Metacoxal lines anteriorly diverge and do not reach posterior margin of metaventrite.</p><p>Male. Shiny. Distinct microreticulation only posteriorly of cervical row of punctures on head; indistinct microreticulation of that line anteriorly. Antennomeres rather long and narrow; antenna thus not appearing stout (Figs 12 D, E, 13 F). Median lobe as in Fig. 8 D and parameres as in Figs 10 C, 11 F.</p><p>Female. Dull. Head with faint microreticulation, pronotum and elytron dull with distinct microreticulation between punctures. Antenna distinctly clubbed due to lateral expansion of antennomeres 3–11 (Figs 12 D, E, 13 F). Some females with broadly enlarged antennomeres (Fig. 3 C).</p><p>Etymology.</p><p>Named after the Habbema lake, where the species was discovered. The name is a noun in the nominative standing in apposition.</p><p>Distribution.</p><p>Only known from the Lake Habbema area in the central mountain range of Papua (Fig. 14).</p><p>Differential diagnosis.</p><p>The new species belongs to a group of larger (TL = 2.25–3.0 mm) and almost blackish high alpine species. Limbodessus habbema sp. nov. is close to L. alexanderi and L. juliana sp. nov. but can be separated from them by the form of the female antennae, which is distinctly clubbed due to lateral expansion of antennomeres 3–11 (Figs 12 D, E, 13 F), the form of the median lobe (Fig. 8 D), and the form of the parameres (Figs 10 C, 11 F).</p><p>Habitat.</p><p>Small puddles in alpine meadows (Fig. 16 C).</p></div>	https://treatment.plazi.org/id/FC9DB0CEA58E545484247D29B4FF48A0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
62D5454AFE1D5597A65B5693F92DD276.text	62D5454AFE1D5597A65B5693F92DD276.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus juliana Balke & Villastrigo & Hájek & Surbakti & Panjaitan & Hendrich 2025	<div><p>Limbodessus juliana sp. nov.</p><p>Figs 4 A, 7 D, 9 D, 10 E, I, 11 E, 13 H, 14, 17 F, G, 18 A, B</p><p>Type locality.</p><p>Mossy puddles in Alpine grassland, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.28334&amp;materialsCitation.latitude=-4.45" title="Search Plazi for locations around (long 140.28334/lat -4.45)">Mt. Juliana area</a>, Sab-me Valley, 3,400–3,500 m [140°17'E, 04°27'S], Papua, Indonesia.</p><p>Type material.</p><p>Holotype, male: “ IRIAN JAYA, Mt. Juliana Gebiet, 16. - 17. 9. 1993, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.28334&amp;materialsCitation.latitude=-4.45" title="Search Plazi for locations around (long 140.28334/lat -4.45)">Sab-me Tal</a>, 3400–3500 m, ca. 140°17'E 04°27'S, leg. M. Balke (14) ”, “ Holotype Limbodessus juliana sp. nov. Balke et al. 2025 ” [red printed label] (NHMW) .</p><p>Paratypes (573): 169 exs., with the same label data as the holotype; 19 exs., “ IRIAN JAYA: Aipomek Area [N Pass] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.08333&amp;materialsCitation.latitude=-4.45" title="Search Plazi for locations around (long 140.08333/lat -4.45)">Aipomek-Langda</a>, 140°05'E 04°27'S, 22. - 24. 8. 1992, 3550–3600 m, leg. M. Balke (20) ” [IR 92 # 20] (MZB, NHMW, KSP, ZSM) ; 216 exs., “ IRIAN JAYA: Aipomek Area [N Pass] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.1&amp;materialsCitation.latitude=-4.45" title="Search Plazi for locations around (long 140.1/lat -4.45)">Aipomek-Langda</a>, 140°06'E 04°27'S, 22. - 24. 8. 1992, 3550–3600 m, leg. M. Balke (21) ” [IR 92 # 21] (NHMW, KSP, ZSM) ; 136 exs., “ IRIAN JAYA: Aipomek Area [N Pass] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.08333&amp;materialsCitation.latitude=-4.45" title="Search Plazi for locations around (long 140.08333/lat -4.45)">Aipomek-Langda</a>, 140°05'E 04°27'S, 22. - 24. 8. 1992, 3550–3600 m, leg. M. Balke (20–22) ” [IR 92 # 20-22] (NHMW, KSP, ZSM) ; 3 exs., “ IRIAN JAYA: Aipomek Area [South Pass], 140°06'E 04°27'S, 27. 8. 1992, 3500 m, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.1&amp;materialsCitation.latitude=-4.45" title="Search Plazi for locations around (long 140.1/lat -4.45)">Aipomek-Langda</a>, leg. Balke (26) ” (NHMW, ZSM) ; 24 exs., “ IRIAN JAYA: Aipomek Area [N Pass], 140°05'E 04°27'S, 22. - 24. 8. 1992, 3800 m, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.08333&amp;materialsCitation.latitude=-4.45" title="Search Plazi for locations around (long 140.08333/lat -4.45)">Aipomek-Langda</a>, leg. Balke (26) ” [IR 92 # 26] (NHMW, KSP, ZSM) ; 6 exs., “ W. Neuguinea, Pass nach Langda, 3900 m, vi. 1979, P. Blum leg. ” (ZSM) . All paratypes are provided with a red printed paratype label.</p><p>Measurements.</p><p>Holotype: TL = 2.6 mm; TL-H = 2.3 mm; MW = 1.2 mm. Paratypes: TL = 2.6–2.7 mm, TL-H = 2.4–2.5 mm, MW = 1.1–1.5 mm.</p><p>Description.</p><p>Body rather elongate-oval, with distinct discontinuity in lateral outline between pronotum and elytron; apically broadly rounded to truncate. No cervical or sutural line. Pronotal stria present and distinct, no elytral stria visible (Fig. 4 A).</p><p>Color. Body chiefly black. Clypeus anteriorly, pronotum anteriorly and laterally slightly paler: dark castaneous brown. Antenna chiefly blackish. Legs dark castaneous brown (Fig. 4 A). Ventral side chiefly black, sternite 7 dark ferruginous.</p><p>Note: The first author also collected / observed paler specimens which appear mature. The body is then castaneous brown to dark ferruginous, elytron sometimes darker medially.</p><p>Sculpture. Head moderately densely punctate; punctation on clypeus and apically sparser; microreticulate. Punctures of same size as meshes of microreticulation. Pronotum with moderately dense punctation along fore-margin; rest sparsely punctate especially on disc. Size of punctures as on head; discally distinctly smaller. Elytron with moderately dense and evenly distributed punctation; between punctures with almost invisible larger polygonal meshes. Ventral side with few punctures, shiny; only sternite 7 with distinct microreticulation.</p><p>Structures. Metacoxal lines slightly diverge anteriorly and do not reach posterior margin of metaventrite; with concavity between lines.</p><p>Male. Pronotum and elytra shiny, without mircoreticulation. Antenna stout (Fig. 13 H). Median lobe as in Fig. 7 D and parameres as in Figs 10 E, I, 11 E.</p><p>Female. Pronotum and elytron dull, with distinct microreticulation between punctures. Antenna stout but without conspicuous modifications (Fig. 13 H).</p><p>Etymology.</p><p>The name refers to Mount Juliana (Fig. 18 B), one of the localities of this species. The name is a noun in the nominative standing in apposition.</p><p>Distribution.</p><p>Only know from the type locality, Sab-me Valley below Mt. Orom, Papua (Fig. 14).</p><p>Differential diagnosis.</p><p>The new species belongs to a group of larger (TL = 2.25–3.0 mm) and almost blackish high alpine species. L. juliana sp. nov. is close to L. alexanderi and L. habbema sp. nov. but can be separated from them by the form of the female antennae, which is stout without any conspicuous modifications (Fig. 13 H), the form of the median lobe (Fig. 7 D), and the form of the parameres (Figs 10 E, 11 E).</p><p>Habitat.</p><p>Small puddles in alpine meadows, with dense stands of submerged mosses and grasses (Figs 17 F, G, 18 A).</p></div>	https://treatment.plazi.org/id/62D5454AFE1D5597A65B5693F92DD276	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
622FD587242958888A5E80D7FA4DE484.text	622FD587242958888A5E80D7FA4DE484.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus konoensis Balke & Villastrigo & Hájek & Surbakti & Panjaitan & Hendrich 2025	<div><p>Limbodessus konoensis sp. nov.</p><p>Figs 2 C, 8 A, 10 F, 11 D, 13 J, 14, 16 C, 17 B, C, D, 18 D, E</p><p>Type locality.</p><p>Mossy puddles in Alpine grassland, near Lake Habbema, 3,400 m [138 ° 67 ' E, 04°14'S], Papua, Indonesia.</p><p>Type material.</p><p>Holotype, male: “ Indonesia: Papua, Lake Habbema, 3400 m, viii. 2018, - 4.1460 ° 138.675 °, Sumoked (Pap 77) ”, “ Holotype Limbodessus konoensis sp. nov. Balke et al. 2025 ” [red printed label] (MZB) .</p><p>Paratypes (722): 69 exs., with the same label data as the holotype (NWM, ZSM); 72 exs., “ Indonesia, Irian Jaya, 10 ix. 1992, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.78334&amp;materialsCitation.latitude=-4.3166666" title="Search Plazi for locations around (long 139.78334/lat -4.3166666)">Kono-Angguruk</a> 2800 m, 139°47'E 04°19'S, leg. Balke (45) ; 71 exs., “ IRIAN JAYA: 15. 9. 1992, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.28334&amp;materialsCitation.latitude=-4.233333" title="Search Plazi for locations around (long 139.28334/lat -4.233333)">Gunung Elit</a>, 3450 m, 139°17'E 04°14'S, leg. Balke (50) ” [IR 92 # 50] (NWM, ZSM) ; 220 exs., “ IRIAN JAYA: Angguruk-Baliem, 10. - 11. 10. 1993, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.28334&amp;materialsCitation.latitude=-4.233333" title="Search Plazi for locations around (long 139.28334/lat -4.233333)">Mt. Elit</a>, ca. 139°17'E 04°14'S, 3400 m, leg. M. Balke (33) ” [IR 93 # 33] ; 35 exs., “ Irian Jaya: 18. 10. 1993 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.71666&amp;materialsCitation.latitude=-4.2166667" title="Search Plazi for locations around (long 138.71666/lat -4.2166667)">Trikora Area Fuß des Mt. Trikora</a>, ca. 138°43'E 04°13'S, 3200 m, leg. M. Balke (38) ” [IR 93 # 38] (NHMW, ZSM) ; 231 exs., “ IRIAN JAYA: Trikora Gebiet, 19. - 20. 10. 1993, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.71666&amp;materialsCitation.latitude=-4.2166667" title="Search Plazi for locations around (long 138.71666/lat -4.2166667)">Habbema [Lake] and Kali Oue Tal [Valley]</a>, ca. 138°43'E 04°13'S, 3450 m, leg. M. Balke (39) ” (NHMW, ZSM) ; 9 exs., “ IRIAN JAYA: 21. 10. 1993, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.75&amp;materialsCitation.latitude=-4.116667" title="Search Plazi for locations around (long 138.75/lat -4.116667)">Habbema-Gebiet Pass zum Ibele Tal</a> [pass to Ibele Valley], ca. 138°45'E 04°07'S, 3300 m, leg. M. Balke (40) ” [IR 93 # 40] (NHMW, KSP, ZSM) ; 6 exs., “ INDONESIA, Papua: Jayawijaya Distr., ca. 30 km W of Wamena, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.67833&amp;materialsCitation.latitude=-4.13" title="Search Plazi for locations around (long 138.67833/lat -4.13)">open wetland nr Habbema lake</a>, 04°07.8'S, 138°40.7'E; 3330 m, J. Hájek &amp; J. Šumpich leg., 5. ii. 2015 ” (NMPC) ; 9 exs., “ Irian Jaya, Wamena, Jayawijaya Prov., W. Gunung Elit, 3300 m, 15. IX. 1991, A. Riedel leg. ” (CLH, ZSM) . All paratypes are provided with a red printed paratype label.</p><p>Measurements.</p><p>Holotype: TL = 2.3 mm; TL-H = 2.1 mm; MW = 1.1 mm. Paratypes: TL 2.35–2.5 mm, TL-H = 2.1–2.2 mm; MW = 1.0– 1.1 mm.</p><p>Description.</p><p>Medium sized, body elongate-oval; apically slightly drop-shaped; with distinct discontinuity in lateral outline between pronotum and elytron. Cervical or sutural line absent. Pronotal and elytral stria present and distinct (Fig. 2 C).</p><p>Color. Body chiefly black. Clypeus anteriorly, pronotum medially and laterally slightly paler: dark ferruginous. Antenna chiefly blackish. Legs dark ferruginous (Fig. 2 C). Ventral side chiefly black. Prosternum, epipleura and sternite 7 ferruginous.</p><p>Note: The first author frequently collected / observed paler specimens that appear mature: head dark castaneous; pronotum anteriorly dark castaneous, rest dark yellow; elytron anterior half dark yellow, posterior half dark castaneous.</p><p>Sculpture. Head moderately densely punctate, densest on frons; punctation on clypeus and vertex sparse; with indistinct microreticulation of small polygonal meshes. Punctures slightly larger than meshes. Pronotum with moderately dense punctation along fore-margin; rest sparsely punctate especially on disc; an indistinct microreticulation visible discally. Size of punctures as on head, discally distinctly smaller. Elytron with moderately dense and evenly distributed punctation; between punctures with hardly visible larger polygonal meshes. Ventral side with few punctures, shiny; only sternite 7 with distinct microreticulation.</p><p>Structures. Metacoxal lines anteriorly diverge and do not reach posterior margin of metaventrite.</p><p>Male. Elytron rather shiny, sometimes with antero-sutural microreticulation. Antenna stout (Fig. 13 J). Median lobe as in Fig. 8 A and parameres as in Figs 10 F, 11 D.</p><p>Female. Generally identical to male in habitus; elytron sometimes dull, with distinct microreticulation between punctures. The corresponding author has seen 4 specimens with strong pronotal and elytral microreticulation from Mount Elit (IR 92 # 50). Antenna stout but without conspicuous modifications.</p><p>Etymology.</p><p>Named after Kono village, situated at the easternmost limit of the species’ range. The name is an adjective in the nominative singular.</p><p>Distribution.</p><p>From the valley below Mt. Orom, Mount Elit, Lake Habbema and Kali Oue Valley around Mt. Trikora (= Mt. Wilhelmina) (Fig. 14).</p><p>Differential diagnosis.</p><p>L. konoensis sp. nov. is a medium sized (TL = 2.3–2.5 mm) and almost blackish high alpine species. The new species is close to L. alexanderi, L. juliana sp. nov., L. moni and L. habbema sp. nov. but can be separated from them by its smaller size, and the form of the female antennae (female with regularly enlarged antennomeres in L. moni), which is stout but without any conspicuous modifications (Fig. 2 C), the form of the median lobe (Fig. 8 A), and the form of the parameres (Figs 10 F, 11 D).</p><p>Habitat.</p><p>Small puddles in alpine meadows (Figs 16 C, 17 B, C, D, 18 D, E).</p></div>	https://treatment.plazi.org/id/622FD587242958888A5E80D7FA4DE484	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
B8F8AA12639F5626BE5D963FBCE01FBB.text	B8F8AA12639F5626BE5D963FBCE01FBB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus moni Surbakti, Balke & Hendrich 2023	<div><p>Limbodessus moni Surbakti, Balke &amp; Hendrich, 2023</p><p>Limbodessus moni Surbakti, Balke &amp; Hendrich, 2023: 414 (original description).</p><p>Type material.</p><p>Holotype: Male, “ Indonesia: Papua, Grasberg, 3,970, viii. 2018, - 4.0447 ° 137.167 °, Surbakti ”, “ HOLOTYPE / Limbodessus moni sp. nov. / Surbakti, Balke &amp; Hendrich det. 2023 ” [red printed label] (MZB) . Paratypes: Three males and one female with the same data as the holotype (MZB, KSP) . 9 males and 2 females: “ Indonesia, Irian Jaya, Lake Anderson [3,555 m] nr Grasberg, 23. iii. 1997, D. Polhemus ” (USNM, ZSM) .</p><p>Additional material examined.</p><p>1 female that, based on the overall morphology, appears to also belong to L. moni . “ Indonesia, Papua; N Sugapa 3000 m, 3°40'11S, 137°4'51E, 27. xii. 1995, Riedel leg. ”</p><p>Morphology.</p><p>Habitus: Male (Fig. 4 C) and female (Fig. 4 B).</p><p>Antennae: Male and female (Fig. 13 K).</p><p>Median lobe (Fig. 8 B) and paramere (Fig. 11 G).</p><p>Distribution.</p><p>Known only from two localities around the Grasberg Mine and Anderson Lake (Carstenz Pyramid area), between 3,555 and 3,970 m, in Papua (Fig. 14).</p></div>	https://treatment.plazi.org/id/B8F8AA12639F5626BE5D963FBCE01FBB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
0291C5DDBB375C169C3480116D8B3B36.text	0291C5DDBB375C169C3480116D8B3B36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus paniai Balke & Villastrigo & Hájek & Surbakti & Panjaitan & Hendrich 2025	<div><p>Limbodessus paniai sp. nov.</p><p>Figs 5 C, 9 B, 11 J, 13 L, 14, 15 C</p><p>Type locality.</p><p>Mats of floating grasses, Enarotali, Lake Paniai, 1,900 m [3°55'16S, 136°22'33E], Papua, Indonesia.</p><p>Type material.</p><p>Holotype, male. “ IR 25 - W. New Guinea, Enarotali [Bobairo], Danau Paniai, 1900 m, 29. vii. - 4. viii. 1991, Balke &amp; Hendrich leg. ” [IR 91 # 25], “ Holotype Limbodessus paniai sp. nov. Balke et al. 2025 ” [red printed label] (NHMW) .</p><p>Paratype: 1 female with the same label data as the holotype (ZSM) . The paratype is provided with a red printed paratype label.</p><p>Measurements.</p><p>Holotype: TL = 2.1 mm; TL-H = 2.0 mm; MW = 1.0 mm. Paratype: TL = 2.05 mm; TL-H = 2.0 mm; MW = 1.0 mm.</p><p>Description.</p><p>Small, body oblong-oval, somewhat parallel-sided, with almost no discontinuity in lateral outline between pronotum and elytron. A faint cervical line visible; sutural line absent. Pronotal and elytral stria present and distinct, both straight; elytral stria short (Fig. 5 C).</p><p>Color. Head dark brown, clypeus paler, ferruginous. Pronotum yellow. Elytron darker yellowish. Body appendages yellow (Fig. 5 C). Prosternum, epipleura and ventrites posteriorly yellow. Mesoventrite dark ferruginous to blackish.</p><p>Sculpture. Head with sparse, coarse punctation; posterior of cervical line impunctate but with microreticulation. Pronotum with moderately dense punctation along anterior margin; otherwise faint and sparse, almost impunctate; punctures however rather shallow giving the pronotum an exceptionally smooth and shiny impression. Elytron with moderately dense and evenly distributed punctation; punctures rather shallow rendering elytron comparably smooth. Ventral side with few punctures only.</p><p>Structures. Metacoxal lines anteriorly diverge and do not reach posterior margin of metaventrite.</p><p>Male. Elytron shiny, without visible microreticulation between punctures. Antenna stout (Fig. 13 L). Median lobe as in Fig. 9 B and parameres as in Fig. 11 J.</p><p>Female. Elytron dull, with distinct microreticulation between punctures. Antenna stout but without conspicuous modifications.</p><p>Etymology.</p><p>Named after the type locality, Lake Paniai (= Wisselmeren). The name is a noun in the nominative standing in apposition.</p><p>Distribution.</p><p>Only known from the type locality (Fig. 14).</p><p>Differential diagnosis.</p><p>The new species can be easily separated from most of the other New Guinean Limbodessus by its yellowish dorsal surface without any dark markings (Fig. 5 C). Limbodessus paniai sp. nov. is close to L. baliem (Fig. 5 A, B) but can be separated by its smaller size ( L. paniai sp. nov. = 2.0– 2.1 mm; L. baliem = 2.6–3.0 mm], the form of the median lobe (Fig. 9 B), and the form of the parameres (Figs 10 H, 11 I, J).</p><p>Habitat.</p><p>Both specimens were collected in very shallow water, among sun exposed floating mats of grasses, at the edge of the large Paniai lake near the village Enarotali (Fig. 15 A – D). The species was associated with numerous Rhantus ekari Balke &amp; Hendrich, 1992, the most common diving beetle in that habitat (Balke and Hendrich 1992).</p></div>	https://treatment.plazi.org/id/0291C5DDBB375C169C3480116D8B3B36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
E4E475F7C04557808E490B4E94707654.text	E4E475F7C04557808E490B4E94707654.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus skalei Hendrich, Surbakti & Balke 2020	<div><p>Limbodessus skalei Hendrich, Surbakti &amp; Balke, 2020</p><p>Limbodessus skalei Hendrich, Surbakti &amp; Balke, 2020: 588 (original description); Surbakti et al. 2023: 420 (key).</p><p>Type material.</p><p>Holotype: Male, “ Indonesia, W-Papua, Raja Ampat Prov., Waigeo, Werabiai River, 197 m, - 0.258519 ° 130.985685 °, 11. 1. 2004 ”, “ HOLOTYPE Limbodessus skalei sp. nov. Hendrich &amp; Balke det. 2019 ” [red printed label] (MZB) . Paratypes: One male and one female with the same data as the holotype (ZSM) .</p><p>Additional material examined.</p><p>1 ex., “ Indonesia: West Papua, Kebar valley, 596 m, 6. v. 2015, - 0, 8406 133,268 2, Panjaitan et al. (BH 059) ” (KSP). First record of the species for New Guinea .</p><p>Morphology.</p><p>Habitus: Male (Fig. 6 A) and female (Fig. 6 B).</p><p>Antennae: Male and female (Fig. 13 M).</p><p>Median lobe (Fig. 9 C).</p><p>Distribution.</p><p>Known from the type locality on Waigeo Island (Hendrich et al. 2020) and Bird’s Head Peninsula (Vogelkop) of New Guinea (Fig. 14).</p></div>	https://treatment.plazi.org/id/E4E475F7C04557808E490B4E94707654	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
460BBBAFB21D5AF4823C8CF0FE8FBA48.text	460BBBAFB21D5AF4823C8CF0FE8FBA48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limbodessus wilhelmina Balke & Villastrigo & Hájek & Surbakti & Panjaitan & Hendrich 2025	<div><p>Limbodessus wilhelmina sp. nov.</p><p>Figs 2 A, 7 C, 10 A, B, 11 C, 12 A, B, 13 D, 14, 16 C</p><p>Type locality.</p><p>Mossy puddles in Alpine grassland, Lake Habbema, 3,200 m [4°07'77S, 138°40'77E], Papua, Indonesia.</p><p>Type material.</p><p>Holotype, male: “ Indonesia, Papua, Lake Habbema, pools, 3200 m, 19. x. 2011, 04.07. 772 S 138.40. 773 E, team UNCEN leg. (PAP 06) ”, “ Holotype Limbodessus wilhelmina sp. nov. Balke et al. 2025 ” [red printed label] (MZB) .</p><p>Paratypes (632): 182 exs., with the same label data as the holotype (MZB, KSP, ZSM); 395 exs. “ IRIAN JAYA: Trikora Gebiet 19. - 20. 10. 1993, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.71666&amp;materialsCitation.latitude=-4.2166667" title="Search Plazi for locations around (long 138.71666/lat -4.2166667)">Habbema [Lake] and Kali Oue Tal [Valley]</a>, ca. 138°43'E 04°13'S, 3450 m, leg. M. Balke (39) ” [IR 93 # 39] (MZB, NHMW, KSP, ZSM) ; 22 exs., “ Irian Jaya: 18. 10. 1993 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.71666&amp;materialsCitation.latitude=-4.2166667" title="Search Plazi for locations around (long 138.71666/lat -4.2166667)">Trikora Area Fuß des Mt. Trikora</a>, ca. 138°43'E 04°13'S, 3200 m, leg. M. Balke (38) ” [IR 93 # 38] (NHMW, ZSM) ; 3 exs., “ Irian Jaya, Wamena E, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.66667&amp;materialsCitation.latitude=-4.116667" title="Search Plazi for locations around (long 138.66667/lat -4.116667)">Habbema lake</a>, 4°07'S 138°40'E, 20. 1. 1999, leg. A. Weigel ” (ZSM) ; 27 exs., “ INDONESIA, Papua: Jayawijaya Distr., ca. 30 km W of Wamena, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.67833&amp;materialsCitation.latitude=-4.13" title="Search Plazi for locations around (long 138.67833/lat -4.13)">open wetland nr Habbema lake</a>, 0 4°07.8'S, 138°40.7'E; 3330 m, J. Hájek &amp; J. Šumpich leg., 5. ii. 2015 ” (NMPC) ; 2 exs., “ Indonesia, West Papua, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.64839&amp;materialsCitation.latitude=-4.109111" title="Search Plazi for locations around (long 138.64839/lat -4.109111)">Highland near Lake Habbema</a>, Sphagnum-pools near the road, 3451 m, 4°06'32.8"S 138°38'54.2"E, 09. 06. 2022, N. Wendlandt leg. ” (CNW, ZSM) . All paratypes are provided with a red printed paratype label.</p><p>Measurements.</p><p>Holotype: TL = 1.8 mm; TL-H = 1.65 mm; MW = 0.85 mm. Paratypes: TL 1.7–1.9 mm, TL-H = 1.6–1.75 mm; MW = 0.85–0.9 mm.</p><p>Description.</p><p>Small, body oblong-oval, with distinct discontinuity in lateral outline between pronotum and elytron. Cervical or sutural line absent. Pronotal and elytral stria present and distinct (Fig. 2 A).</p><p>Color. Body chiefly black. Clypeus anteriorly, pronotum laterally slightly paler, castaneous brown. Antenna almost completely blackish. Legs dark ferruginous to blackish (Fig. 2 A). Ventral side chiefly black. Prosternum, epipleura and sternite 7 ferruginous.</p><p>Sculpture. Head comparably densely punctate; punctation apically sparse; with indistinct microreticulation of small polygonal meshes. Size of punctures corresponds to diameter of meshes. Pronotum shiny; rather sparse and fine punctation; slightly denser and coarser along fore-margin; without microreticulation. Elytron with moderately dense and evenly distributed punctation; size of punctures as on head or slightly larger; between punctures with large, indistinct polygonal meshes. Ventral side with few punctures, shiny; only sternite 7 with distinct microreticulation.</p><p>Structures. Metacoxal lines anteriorly diverge and do not reach posterior margin of metaventrite.</p><p>Male. Elytron shiny, without visible microreticulation between punctures. Antenna stout (Fig. 13 D). Median lobe as in Fig. 7 C and parameres as in Figs 10 A, B, 11 C.</p><p>Female. Elytron shiny, without visible microreticulation between punctures. Antenna stout but without conspicuous modifications (Fig. 12 A, B).</p><p>Etymology.</p><p>Named after the highest peak in the type area, Mount Wilhelmina (the historic name of what is nowadays known as Mt. Trikora). The name is a noun in the nominative standing in apposition.</p><p>Distribution.</p><p>Only known from the Lake Habbema area, in the Central Highlands of Papua (Fig. 14).</p><p>Differential diagnosis.</p><p>Limbodessus wilhelmina sp. nov. belongs to a group of smaller (TL = 1.55–1.9 mm) and almost blackish high alpine species. The new species is close to L. aipo sp. nov., L. aldente sp. nov. and L. elit sp. nov. but can be separated from them by the form of the female antennae, which is stout without any conspicuous modifications (Fig. 12 A, B), the form of the median lobe (Fig. 7 C), and the form of the parameres (Figs 10 A, B, 11 C).</p><p>Habitat.</p><p>Small puddles in alpine meadows (Fig. 16 C).</p></div>	https://treatment.plazi.org/id/460BBBAFB21D5AF4823C8CF0FE8FBA48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Villastrigo, Adrián;Hájek, Jiří;Surbakti, Suriani;Panjaitan, Rawati;Hendrich, Lars	Balke, Michael, Villastrigo, Adrián, Hájek, Jiří, Surbakti, Suriani, Panjaitan, Rawati, Hendrich, Lars (2025): A review of Limbodessus Guignot, 1939 from New Guinea, with description of eight new tropical-alpine species (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 9: 5-28, DOI: 10.3897/alpento.9.147831
