identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A70CDA7B4D189F5BFCB8AA421BB3FE70.text	A70CDA7B4D189F5BFCB8AA421BB3FE70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ancyrocephalidae Bychowsky 1937	<div><p>Ancyrocephalidae Bychowsky, 1937</p><p>The anatomy of the new species described herein corresponds to the diagnosis of Quadriacanthus Paperna, 1961 given by PAPERNA (1961), amended by KRITSKY &amp; KULO (1988) and used by NACK et al. (2015).</p></div>	https://treatment.plazi.org/id/A70CDA7B4D189F5BFCB8AA421BB3FE70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bahanak, Dieu ne dort;Nack, Jacques;Pariselle, Antoine;Bilong Bilong, Charles F.	Bahanak, Dieu ne dort, Nack, Jacques, Pariselle, Antoine, Bilong Bilong, Charles F. (2016): Description of three new species of Quadriacanthus (Monogenea: Ancyrocephalidae) gill parasites of Clarias submarginatus (Siluriformes: Clariidae) from Lake Ossa (Littoral region, Cameroon). Zoologia (e 20160044) 33 (4): 1-8, DOI: 10.1590/S1984-4689zool-20160044, URL: http://dx.doi.org/10.1590/s1984-4689zool-20160044
A70CDA7B4D189F58FCFDA9021DC0FA2B.text	A70CDA7B4D189F58FCFDA9021DC0FA2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quadriacanthus macruncus Bahanak, Nack & Pariselle 2016	<div><p>Quadriacanthus macruncus Bahanak, Nack &amp; Pariselle, sp. nov.</p><p>Fig. 3</p><p>urn:lsid:zoobank.org:act: 2FE39BE9-F936-4BE4-A907-F8BDB8FEA566</p><p>Description (based on 9 specimens, all mounted in GAP): adults measure 428.3 (209.7-737.6) long, 83.4 (47.8-101.5) wide at level of ovary. Pharynx circular is 26. 9 (25.8-27.9) wide. Straight tubular penis widened at basal extremity and tapering at distal one, Pe = 28.7 (25.1-31.5) long. Accessory piece pointed, slightly curved and wider at median level, equipped with two subterminal, symmetrical and similar spines, Ap = 21.6 (18.1- 23.3). Vagina not observed. Hooklets pair I = 12.9 (12.2-13.7), II = 12.9 (11.8-13.8), III = 13.6 (12.9-13.9), IV = 25.6 (24.9-26.7), V = 13.4 (12.4-14.3), VI = 14 (13.1-15.9), VII = 13.5 (12.9-14.1). Dorsal bar composed of developed rectangular centre, Ct = 24.7 (24.1-25.7), w = 9.1 5 (6.5-14.2) wide, with one median triangular process posteriorly directed, h = 10.2 (8.6-11.6), and two lateral expansions, x = 35.6 (33.6-37.1). Dorsal anchor with curved blade, reduced guard and short point, a = 38.3 (36.8-41.6), ba = 10.6 (9.3-11.7), e = 2.5 (1.9-3.1). Dorsal triangular cuneus (or patch), i = 3.6 (2.1-4.6), j = 9 (6.1-12.5). Ventral bar V-shaped, made up of two branches medially articulated, x = 46.3 (41.9- 49.5) long, w = 6.5 (5.5-8) wide. Ventral anchor quite similar to dorsal one, a = 39.7 (37.3-41.5), ba = 9.2 (8.8-9.9), short point e = 3.9 (3.2-4.3). Slightly curved ventral cuneus with bilobed proximal extremity, i = 2.9 (2.2-3.9), j = 6.1 (4-7.5).</p><p>Type host. Clarias submarginatus Peter, 1882 .</p><p>Site. Gills.</p><p>Type locality. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=10.05&amp;materialsCitation.latitude=3.85" title="Search Plazi for locations around (long 10.05/lat 3.85)">Lake Ossa</a>, Cameroon (3°45’- 3°51’N, 9°58’- 10°03’E) .</p><p>Type specimens. Holotype deposited at the Royal Museum for Central Africa (Tervuren): #37935 . Paratype deposited at the Royal Museum for Central Africa (Tervuren): #37936 .</p><p>Parasitic indices. Prevalence = 18.2%, mean abundance = 0.2 ± 0.4.</p><p>Etymology. The specific name, macruncus, refers to the large size of the anchors of this species.</p><p>Remarks. The morphology of the male copulatory complex of Q. macruncus resembles Q. levequei Birgi, 1988 and Q. euzeti Nack, Pariselle &amp; Bilong Bilong, 2015; Q. macruncus) can be distinguished by its smaller size: (Pe = 25.1-31.5 vs 45-50, 36-40; Ap = 18.1-23.3 vs 30-35, 25-28), and by the morphology of the median process, directed posteriorly to the dorsal bar, triangular. This structure is in the shape of a stick in Q. levequei and a funnel in Q. euzeti .</p></div>	https://treatment.plazi.org/id/A70CDA7B4D189F58FCFDA9021DC0FA2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bahanak, Dieu ne dort;Nack, Jacques;Pariselle, Antoine;Bilong Bilong, Charles F.	Bahanak, Dieu ne dort, Nack, Jacques, Pariselle, Antoine, Bilong Bilong, Charles F. (2016): Description of three new species of Quadriacanthus (Monogenea: Ancyrocephalidae) gill parasites of Clarias submarginatus (Siluriformes: Clariidae) from Lake Ossa (Littoral region, Cameroon). Zoologia (e 20160044) 33 (4): 1-8, DOI: 10.1590/S1984-4689zool-20160044, URL: http://dx.doi.org/10.1590/s1984-4689zool-20160044
A70CDA7B4D1B9F5EFB86AEE01B4AFA07.text	A70CDA7B4D1B9F5EFB86AEE01B4AFA07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quadriacanthus ossaensis Bahanak, Nack & Pariselle 2016	<div><p>Quadriacanthus ossaensis Bahanak, Nack &amp; Pariselle, sp. nov.</p><p>Fig. 4</p><p>urn:lsid:zoobank.org:act: 81837D50-AFB3-4C9A-AE61-073D2B42DE1A</p><p>Description (based on 33 specimens, all mounted in GAP): adults measure: 368 ± 86.8 (188.9-597.7) long, 65. 5 ± 16. 1 (33.7- 98.9) wide at level of ovary). Pharynx circular is 21 (18-24) wide. Winding tubular penis, widened at basal extremity, regularly tapering to the distal one Pe = 28.2 ± 3.1 (23.9-32.4). Accessory piece, tubular, slightly arched and ending with two expansions: one fork shaped, the other one chevron shaped, Ap = 26.5 ± 2.9 (22-30.1). Tubular vagina, Vg = 24.9 ± 5 (20.9-30.6). Hooklets pair I = 13.5 ± 0.9 (11.7-15.4), II = 13.3 ± 0.8 (12-15.6), III = 14.3 ± 0.6 (12.1-15.7), IV = 24.8 ± 0.5 (23.8-25.7), V = 12.9 ± 0.7 (11.7- 14.3), VI = 13.2 ± 0.8 (11.9-15), VII = 12.7 ± 0.8 (11-14.2). Dorsal bar composed of a developed trapezoidal centre, Ct = 17.4 ± 2.7 (14.5-21.9), w = 9.7 ± 0.8 (8.2-10.7) wide, with one posteriorly directed rectangular process, h = 8.6 ± 1.4 (7.1-9.9), and two lateral expansions, x = 20.4 ± 1.2 (18.9-22). Dorsal anchor without shaft, but with curved blade and short point, a = 28.3 ± 1.2 (26.9-30.8), ba = 9.1 ± 0.6 (8.1-9.8), e = 6.2 ± 1.2 (5.2-8.6), triangular dorsal cuneus (or patch) slightly curved, i = 5.2 ± 0.5 (4.8-5.8), j = 15.7 ± 0.6 (15-16.8). Ventral bar V-shaped, made up of two branches medially articulated, x = 32.6 ± 1.3 (30.4-36.2), w = 4.3 ± 0.5 (3.3-5.3), ventral anchor quite similar to dorsal one, a = 19.9 ± 1 (18.6-21.4), ba = 4.8 ± 0.3 (4.1-7.7), e = 9.8 ± 1.2 (7.7-12.1), triangular ventral cuneus, i = 2.4 ± 0.4 (1.6-3.3), j = 4.1 ± 0.7 (3.5-5.8).</p><p>Type host. Clarias submarginatus Peter, 1882 .</p><p>Site. Gills.</p><p>Type locality. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=10.05&amp;materialsCitation.latitude=3.85" title="Search Plazi for locations around (long 10.05/lat 3.85)">Lake Ossa</a>, Cameroon (3°45’- 3°51’N, 9°58’- 10°03’E).</p><p>Type specimens. Holotype deposited at the Royal Museum for Central Africa (Tervuren): #37933 . Paratype deposited at the Royal Museum for Central Africa (Tervuren): #37934 .</p><p>Materials studied. 43 host individuals and 33 monogeneans.</p><p>Parasitic indices. Prevalence = 81.8%, mean abundance = 4.4 ± 5.8.</p><p>Etymology. The specific name, ossaensis, refers to the type locality of this species.</p><p>Remarks. The morphologies of the penis and of the accessory piece of Q. ossaensis sp. nov. are similar to Quadriacanthus ayameensis N’Douba, Lambert &amp; Euzet, 1999, a gill parasite of Heterobranchus longifilis (Valenciennes, 1840) and Heterobranchus isopterus Bleeker, 1863 in the rivers Bia, Agnéby and in Lake Ayamé (Ivory Coast). Quadriacanthus ossaensis can be distinguished from Q. ayameensis species by having one tip of the distal end of the accessory piece shaped as a fork, by the size of the dorsal cuneus (patch) (i = 2.5-7 vs 7-10, j = 13.8-18 vs 8-11), the trapezoid shape of the central sclerite of the dorsal bar, and the shape (thin and twisted) of the branches of the ventral bars.</p></div>	https://treatment.plazi.org/id/A70CDA7B4D1B9F5EFB86AEE01B4AFA07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bahanak, Dieu ne dort;Nack, Jacques;Pariselle, Antoine;Bilong Bilong, Charles F.	Bahanak, Dieu ne dort, Nack, Jacques, Pariselle, Antoine, Bilong Bilong, Charles F. (2016): Description of three new species of Quadriacanthus (Monogenea: Ancyrocephalidae) gill parasites of Clarias submarginatus (Siluriformes: Clariidae) from Lake Ossa (Littoral region, Cameroon). Zoologia (e 20160044) 33 (4): 1-8, DOI: 10.1590/S1984-4689zool-20160044, URL: http://dx.doi.org/10.1590/s1984-4689zool-20160044
A70CDA7B4D1D9F5FFEF3AEB11CC6F9D1.text	A70CDA7B4D1D9F5FFEF3AEB11CC6F9D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quadriacanthus submarginati Bahanak, Nack & Pariselle 2016	<div><p>Quadriacanthus submarginati Bahanak, Nack &amp; Pariselle sp. nov.</p><p>Fig. 5</p><p>urn:lsid:zoobank.org:act: 423B9D06-68FA-4424-86F6-4D532D7075F4</p><p>Description (based on 22 specimens, all mounted in GAP): adults measure 399.6 (250.6-572.3) length, 63.8 (11.6-113.6) wide at level of ovary. Pharynx circular is 21.7 (20.2-21.6) wide. Penis wide-opened at basal extremity, tapering, bent and forked at the distal extremity, Pe = 29.7 (25.8-31.6). Accessory piece tubular, slightly curved, with a complex structure made up of one long tip flanked by a short tip and by a bulb, Ap = 21.8 (19.2-25.8). Tubular curved vagina, slim and slightly sclerotized at basal zone, and connected to a large seminal receptacle which opens near vaginal pore, Vg = 26.9 (25.3-32.1). Hooklets pair I = 13.7 (12.7-14.7), II = 14 (12.9-14.7), III = 14.3 (12.5-15.7), IV = 21.1 (19.7-24.3), V = 13.6 (12.3-14.8), VI = 14.1(12.5-15.1), VII = 14.1 (11.7-15.7). Dorsal bar with a rectangular centre, Ct = 19.1 (16.7-23.3), w = 8.5 (6-11.5) wide, a median posteriorly directed triangular process, h = 5.3 (4-6.5), and two lateral expansions, x = 29.2 (26.4-32.9), dorsal anchor lacking shaft, but with reduced guard, curved blade at distal region and mean point, a = 32.7 (30-34.7), ba = 9.1 (7.7-10), e = 8.4 (6.1-9.7), triangular and slightly curved dorsal cuneus (or patch), i = 3.5 (2.1-3.1), j = 6.3 (5.1-7.1), ventral bar made up of two branches medially articulated, x = 43 (37.9-47.7), w = 5.5 (3.5-7.6), ventral anchor without shaft, but with reduced guard, curved blade, and with long point, a = 24.7 (22.8-26.9), ba = 5.8 (5.1-6.3), e = 10.8 (8-11.7), ventral cuneus (or patch) smaller than dorsal one, i = 1.7(1.2-2), j = 3.8 (4.6-4.9).</p><p>Type host. Clarias submarginatus Peter, 1882 .</p><p>Site. Gills.</p><p>Type locality. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=10.05&amp;materialsCitation.latitude=3.85" title="Search Plazi for locations around (long 10.05/lat 3.85)">Lake Ossa</a>, Cameroon (3°45’- 3°51’N, 9°58’- 10°03’E).</p><p>Type specimens. Holotype deposited at the Royal Museum for Central Africa (Tervuren): #37937 . Paratype deposited at the Royal Museum for Central Africa (Tervuren): #37938 .</p><p>Materials studied. 43 host individuals and 22 monogeneans.</p><p>Parasitic indices. Prevalence = 45.5%, mean abundance = 2.8 ± 3.8.</p><p>Etymology. The specific name, submarginati refers to specific name of the host.</p><p>Remarks. This species is close to Q. macruncus sp. nov. in the morphology of the dorsal and ventral bars, dorsal and ventral anchors; it differs from it in the morphology of the copulatory complex and the size of the point of its dorsal (e = 6.1-9.7 vs 1.9-3.1) and ventral (e = 8-11.7 vs 3.2-4.3) anchors.</p></div>	https://treatment.plazi.org/id/A70CDA7B4D1D9F5FFEF3AEB11CC6F9D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bahanak, Dieu ne dort;Nack, Jacques;Pariselle, Antoine;Bilong Bilong, Charles F.	Bahanak, Dieu ne dort, Nack, Jacques, Pariselle, Antoine, Bilong Bilong, Charles F. (2016): Description of three new species of Quadriacanthus (Monogenea: Ancyrocephalidae) gill parasites of Clarias submarginatus (Siluriformes: Clariidae) from Lake Ossa (Littoral region, Cameroon). Zoologia (e 20160044) 33 (4): 1-8, DOI: 10.1590/S1984-4689zool-20160044, URL: http://dx.doi.org/10.1590/s1984-4689zool-20160044
