taxonID	type	description	language	source
A91987BEFFEECA2CFD8D23FFFB2BDFA5.taxon	description	Specimens examined – Holotype female, University of Agriculture, Faisalabad, Punjab, Pakistan, 31 ° 26 ' N, 73 ° 04 ' E, 20 August 2014, soil core. Paratypes: 3 females, same data as holotype except 21 October 2014; 1 female, Dhannot, District Lodhran, Punjab, Pakistan, 29 ° 36 ' N, 71 ° 45 ' E, 21 June 2014, soil core; 1 female, Dhudial, Chakwal, Pakistan, 33 ° 03 ' N, 72 ° 57 ' E, 19 February 2014, soil core. Female Dorsal idiosoma (Fig. 1) – Female with doral shield tapering posteriorly, lightly sclerotised, pale yellow, not completely covering dorsal idiosoma; length 506 µm, width 294 µm, surface with faint polygonal ornamentation thoughout, with adhering fragments of soil and debris, and approximately 12 pairs of minute circular pores and two pairs of lyrifissures. Podonotal region with 23 pairs of setae, opisthonotal region with 17 pairs of setae including Zx 1 and Zx 2 between the J and Z setae. Three of the six specimens examined have single unpaired Jx seta between J 3 and J 4. Seta z 1 fine, pointed, pilose, J 5 pointed, lightly pilose, all other setae distally expanded and pilose (Fig. 2). Setae j 3, j 4, r 3 longest (40 – 50 μm), z 1 and J 5 shortest (20 – 30 μm), others intermediate (30 – 40 μm). Opisthonotal soft cuticle with seven pairs of setae, three anterior pairs fine, four posterior pairs heavy and strongly pilose. Ventral idiosoma (Fig. 3) – Tritosterum with trapezoidal base, pre-sternal area with transverse lineate ornamentation. Sternal shield with curved anterior margin and weakly concave posterior margin, with weak polygonal ornamentation, lateral corners fused with endopodal plates II / III, with three pairs of subequal smooth needle-like setae, st 1 on anterior margin of shield, and two pairs of lyrifissures. Metasternal setae st 4 and metasternal pore inserted in soft integument. Endopodal plates III / IV long, narrow, crescentic. Genital shield elongate, narrowest between coxa IV, weakly ornamented laterally and posteriorly, with one pair of smooth pointed setae. Anal shield longer than wide, with weak ornamentation laterally and smooth posteriorly, cribrum extending anteriorly past post-anal seta, post-anal seta thick and distally pilose, para-anal setae smooth and pointed; pore gv 3 on edges of shield, at posterior level of anus. Peritrematal shield very long and narrow except for a short section outside the peritreme opposite coxae II – III bearing a gland pore and a poroid; anterior end reaching z 1, post-stigmatal section of peritrematal shield narrow, reaching mid-level of coxa IV; with one pre-stigmatic pore and two post-stigmatic pores. Opisthogastric integument with one pair of elongate metapodal plates and eight pairs of distally pilose setae, their length, thickness, and pilosity increasing posteriorly; opisthogastric pores not visible. Gnathosoma – Hypostomal setae normal, deutosternal groove with six transverse rows of teeth, each row with about 20 fine teeth (Fig. 4), corniculi robust and horn-like, palp trochanter with two long pointed setae, other features of palp normal for genus. Movable digit of chelicera with two teeth, arthrodial brush with a circular membrane and several long bristles; fixed digit with six small proximal teeth, three distal teeth, and one large tooth near pilus dentilus, pilus dentilis fine and pointed (Fig. 5); epistome semi-circular, serrated, with a long underlying bifurcate process (Fig. 6). Legs (Figs. 7 – 10) – Chaetotaxy typical for free-living Laelapidae as reported by Evans (1963): Leg I: coxa 0 0 / 1 0 / 1 0, trochanter 1 0 / 1 1 / 2 1, femur 2 3 / 1 2 / 3 2, genu 2 3 / 2 3 / 1 2, tibia 2 3 / 2 3 / 1 2. Leg II: coxa 0 0 / 1 0 / 1 0, trochanter 1 0 / 2 0 / 1 1, femur 2 3 / 2 2 / 1 1, genu 2 3 / 1 2 / 1 2, tibia 2 2 / 1 2 / 1 2, tarsus 3 3 / 2 3 / 2 3 + mv, md. Leg III: coxa 0 0 / 1 0 / 1 0, trochanter 1 0 / 2 0 / 1 1, femur 1 2 / 1 1 / 0 1, genu 2 2 / 1 2 / 1 1, tibia 2 1 / 1 2 / 1 1, tarsus 3 3 / 2 3 / 2 3 + mv, md. Leg IV: coxa 0 0 / 1 0 / 0 0, trochanter 1 1 / 0 0 / 2 1, femur 1 2 / 1 1 / 0 1, genu 2 2 / 1 3 / 0 1, tibia 2 1 / 1 3 / 1 2, tarsus 3 3 / 2 3 / 2 3 + mv, md. Most dorsal and ventral setae thick, distally expanded and pilose; seta pv 1 on femur II short, thick, spine-like; pl 1 and pl 2 on tarsus II very thick and conspicuous. Insemination structures – Not visible in the available specimens. Etymology The name lutosus (dirty) refers to the fragments of debris adhering to most of the specimens. Notes The genus Cosmolaelaps includes 135 species (Moraes et al. 2022). Attempts to develop a subgenus or species-group classification for these species have been unsatisfactory (for example Karg, 1988). We have checked the descriptions and illustrations of all the species of Cosmolaelaps listed by Moraes et al. (2022), with the exception of a few Berlese species that cannot be recognised. The dorsal shield setae in Cosmolaelaps are modified in various ways – expanded and leaf-like, scimitar-shaped with a basal swelling, terminally trifurcate, or with a longitudinal rib, but only a few species have distally pilose brush-like setae. A few pairs of opisthonotal setae are distally pilose in C. bipennata (Karg, 2003) and C. brevilingua (Karg, 2006) but most of the idiosomal setae in these species are smooth and pointed. The only other species that has distal pilosity on most of the dorsal shield setae is C. kassaii (Van Aswegen & Loots, 1970). However, C. kassaii has strongly developed exopodal plates, which are absent in C. lutosus. The anal shield of C. kassaii is almost circular, with a small cribrum confined to its posterior margin. In C. lutosus the anal shield is elongate and produced posteriorly, and the arms of the cribrum extend anteriorly beyond the post-anal seta.	en	Khan, Ahmad Kamran, Halliday, Bruce, Bashir, Muhammad Hamid (2024): New records of soil mites from Pakistan, with description of a new species (Acari: Laelapidae). Persian Journal of Acarology 13: 719-732, DOI: 10.22073/pja.v13i4.84483
A91987BEFFEBCA2DFE7424B2FB77DD7D.taxon	description	One female, Rajhana, Toba Tek Singh, Pakistan, 20 April 2014, soil core, undisturbed soil, A. K. Khan coll.; 1 female, Basti Budhan, Pakistan, 28 April 2014, soil core, undisturbed soil, A. K. Khan coll. Notes Bregetova (1977) placed this species in Hypoaspis (Euandrolaelaps) Bregetova, 1977. Karg (1979, 1993) synonymised Euandrolaelaps under Alloparasitus Berlese, 1920, and transferred L. karawaiewi to Hypoaspis (Pneumolaelaps) Berlese, 1920. However, it cannot be placed in Pneumolaelaps as defined by Evans and Till (1966) because the peritreme is of normal width and genu IV has only one ventral seta. We agree with Moraes et al. (2022) in placing this species in Euandrolaelaps until the whole group can be thoroughly revised. The diagnostic character states we used to identify the species are the modifications of leg II of the female, and the very short cheliceral digits. Femur II has a large ventral cuticular spur and a thickened ventral spine-like seta, there is a very large spine-like ventral seta on genu II, and thickened ventral setae on tibia and tarsus II (Costa 1968, his Figure 15). Our specimens also show the other distinctive character states described by Costa (1968) – movable digit of chelicera with a single tooth, corniculi elongate, and extreme differences in the lengths of the hypostomal setae. The species has been recorded from Israel (Costa 1968), Iran (Kazemi and Rajaei 2013; Hasanvand et al. 2014), Russia (Berlese 1904 b; Bregetova 1956, 1977), Bulgaria (Koyumdjieva 1981), Slovakia (Mrciak 1963), Latvia (Salmane 2001), China (Ma 1995) and USA (Walter 1988; Walter et al. 1988; Farrierand Hennessey 1993). It is now recorded from Pakistan for the first time.	en	Khan, Ahmad Kamran, Halliday, Bruce, Bashir, Muhammad Hamid (2024): New records of soil mites from Pakistan, with description of a new species (Acari: Laelapidae). Persian Journal of Acarology 13: 719-732, DOI: 10.22073/pja.v13i4.84483
A91987BEFFEACA2DFDB226DAFDD4DA14.taxon	description	Two females, Jhangiwala, Bahawalpur, Punjab, Pakistan, 29 ° 25 ' N, 71 ° 45 ' E, 30 June 2014, soil core, undisturbed soil. Notes Specimens of Gaeolaelaps minor are very small and lightly sclerotised, with a narrow and elongate dorsal shield. The epigynal shield is elongate behind setae st 5, but does not reach the anal shield. The peritremes reach to the posterior level of coxa I. The anal shield is semi-triangular, widest in the anterior half. The ventral setae on tibia IV and genu IV are longer and thicker than the other leg setae. This species is unusual in the genus Gaeolaelaps in that the anterior margin of the sternal shield is strongly desclerotised, so that setae st 1 are inserted in soft granular skin in the pre-sternal area. Beaulieu (2009) reported only two species with this character state – G. minor and G. franzi (Van Aswegen and Loots 1970). Our specimens agree with the description of G. minor, with thesternal shield smooth, the sternal setae short, and hypostomal seta h 3 short. In G. franzi the sternalshield is ornamented, the sternal setae are much longer, and hypostomal seta h 3 is very long. Geolaelaps minor has been reported from Israel (Costa 1968) and India (Pramanik and Raychaudhuri 1978). Bregetova (1977) provisionally proposed that G. minor could be a junior synonym of Laelaps (Hypoaspis) expolitus Berlese, 1904 c, which was described from Italy. Berlese's illustrations of L. expolitus appears to show the unusual desclerotised anterior margin of the sternal shield. However, the types of L. expolitus are missing or lost (Castagnoli and Pegazzano 1985), and no informationabout it has been published since the original description, so the proposed synonymy cannot beconfirmed.	en	Khan, Ahmad Kamran, Halliday, Bruce, Bashir, Muhammad Hamid (2024): New records of soil mites from Pakistan, with description of a new species (Acari: Laelapidae). Persian Journal of Acarology 13: 719-732, DOI: 10.22073/pja.v13i4.84483
A91987BEFFEACA2EFE7B2141FB6ADE35.taxon	description	One female, Dhannot, District Lodhran, Punjab, Pakistan, 29 ° 36 ' N, 71 ° 45 ' E, 3 April 2014; 2 males, same data except 27 April 2014; 1 male, same data except 20 April 2014; 1 female, 3 males, Rajhana, Toba Tek Singh, Punjab, Paksitan, 30 ° 50 ' N, 72 ° 33 ' E, 27 October 2014; 1 male Dera Ghazi Khan, Basti Budhan, 29 ° 49 ' N, 70 ° 36 ' E, 18 August 2014, undisturbed soil; 1 male, same data except 28 February 2014; 1 male, Layyah, 30 ° 58 ' N, 70 ° 56 ' E, 27 April 2014, undisturbed soil. Notes Joharchi et al. (2016) referred to this species as Laelaspisella kabitae, using a broad concept of Laelaspisella Marais & Loots, 1969 that included six species. In a simultaneous paper, Nemati and Gwiazdowicz (2016 b) used a narrow concept of Laelaspisella that includes only the two original species L. epigynialis Marais & Loots 1969 and L. macrodorsalis Marais & Loots, 1969. It is difficult to include G. kabitae in Laelaspisella without also including Pogonolaelaps Nemati & Gwiazdowicz, 2016 b. Nemati et al. (2019) discussed the morphology of L. kabitae in detail, but could not decide on its generic placement. Moraes et al. (2022) also considered it as incertae sedis. We provisionally place G. kabitae in Gymnolaelaps until all these genera are thoroughly revised.	en	Khan, Ahmad Kamran, Halliday, Bruce, Bashir, Muhammad Hamid (2024): New records of soil mites from Pakistan, with description of a new species (Acari: Laelapidae). Persian Journal of Acarology 13: 719-732, DOI: 10.22073/pja.v13i4.84483
A91987BEFFE9CA2EFD952522FEAFD9E9.taxon	description	One female, Pakistan, Rajhana Toba Tek Singh, 30 ° 50 ' N, 72 ° 33 ' E, 1 August 2014, soil core, undisturbed soil. Notes	en	Khan, Ahmad Kamran, Halliday, Bruce, Bashir, Muhammad Hamid (2024): New records of soil mites from Pakistan, with description of a new species (Acari: Laelapidae). Persian Journal of Acarology 13: 719-732, DOI: 10.22073/pja.v13i4.84483
A91987BEFFE8CA2FFD8C2792FA6CDFEC.taxon	materials_examined	Material examined Ten females, University of Agriculture Faisalabad, Pakistan, 20 August 2014, soil core, agricultural soil; 3 females, same data except 21 October 2014; 1 female, same data except 2 June 2014. Notes Allred (1969) recorded specimens identified as Hypoaspis miles (Berlese, 1892) associated with small mammals in Pakistan. Walter and Campbell (2003) discussed the difficulty of identifying species in this group, and we have not been able to examine Allred's specimens to confirm their identification. In view of the fact that S. scimitus is a widespread inhabitant of nests and burrows, we provisionally suggest that Allred's specimens from mammals in Pakistan were actually S. scimitus. These two species are distinguished by the shape of the opisthonotal shield (tapering gradually behind S 2 in S. miles; tapering sharply behind S 2 in S. scimitus), the length of the post-anal seta (much shorter than para-anal setae in S. miles; subequal to para-anal setae in S. scimitus), and the form of the palp tarsal claw (two subequal tines in S. miles, two subequal tines and a small basal tine in S. scimitus).	en	Khan, Ahmad Kamran, Halliday, Bruce, Bashir, Muhammad Hamid (2024): New records of soil mites from Pakistan, with description of a new species (Acari: Laelapidae). Persian Journal of Acarology 13: 719-732, DOI: 10.22073/pja.v13i4.84483
