identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
AC2487EDFFFCB854A9EF68B9504D20AB.text	AC2487EDFFFCB854A9EF68B9504D20AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sonerila cornuta K. S. Nguyen, Aver. & C. W. Lin 2025	<div><p>Sonerila cornuta K.S.Nguyen, Aver. &amp; C.W.Lin, sp. nov. (Figs. 1 &amp; 2)</p><p>Type: — VIETNAM. Nghe An Province: Ky Son District, Na Ngoi Municipality, eastern slopes of Phu Xai Lai Leng Mountain, primary very humid broad-leaved forest on steep mountain slopes composed of sandstone and gray shale at elevation of 2400–2700 m a.s.l., around point 19°12’17.7’’N 104°11’06.6’’E, lithophytic or terrestrial creeping herb, flowers purple, very common from 2,400 – 2,700 m a.s.l., 25 October 2013, L. Averyanov, N. T. Hiep, N. S. Khang, L. M. Tuan, N. A. Trang, L. H. Dan, CPC 6245 (holotype: LE LE 01253136 http://re.herbariumle.ru/01253136, photos LE LE 01124785 http://re.herbariumle.ru/01124785).</p><p>Diagnosis: — Sonerila cornuta resembles S. reptans J.Wai &amp; J.M.Hu (2023: 131) with creeping to ascending, many branched stems, hairy leaves, and terminal scorpioid, few-flowered cymes. However, it is distinguished from S. reptans in ovate or broadly lanceolate leaves (vs. elliptic or elliptic-ovate) acute or acuminate at apex (vs. acute or obtuse), and 3 or 4 (vs. 2 or 3) pairs of secondary veins, obclavate hypanthium (vs. cylindrical-campanulate or obconical) covered by purple hispid, partially glandular hairs (vs. hairs white, simple, woolly), and purple subulate anthers (vs. yellow linear-lanceolate) with two prominent, S-curved basal lobes (vs. basal lobes rather straight, inconspicuous).</p><p>Etymology: —The species name refers two prominent, horn-shaped basal lobes of anther.</p><p>Description:— Lithophytic or terrestrial, perennial herbs. Stems creeping with suberect upper stem, many-branched, rooting at nodes when procumbent; stem terete, 8–14 cm tall, internodes 1–3.5 cm long, 1.5–2.5 mm thick, shorter on the distal branches, pale yellow, pale pinkish, pink, or light greenish, villous, hirsute or pilose. Leaves opposite, decussate, isomorphic or slightly dimorphic, subequal or unequal in sizes in each pair; petioles yellowishgreen, 0.7–2.8 cm long, densely pink to magenta, villous, hirsute or pilose; leaf blade chartaceous, symmetrical or sometimes slightly oblique, ovate or broadly lanceolate, 0.7–4.2 × 0.5–3 cm; adaxial surface bright green, densely white to pinkish hirsute, abaxial surface sparsely to densely white to pinkish hirsute; apex acute or attenuate, base cordate or rounded, margins denticulate with rows of magenta hairs; venation pinnate, primary vein shallowly grooved above, and prominent on the abaxial surface; secondary veins 3 or 4 pairs, arising from both sides of primary vein in an alternate or occasionally sub-opposite manner. Inflorescence terminal, simple scorpioid cyme, erect or ascending, 2–6-flowered; peduncles terete, 0.8–3.5 cm long, yellowish-pink, pale orange or light green, sparsely to densely hirsute with glandular hispid hairs; pedicels, ovary, hypanthium, sepals and petals pubescent with scattered to dense, purple, long, hispid, multicellular, simple and glandular (capitate) hairs to 1 mm long. Flowers 3-merous; pedicels 1.5–2 mm long, yellowish-pink. Hypanthium obclavate, pink to yellowish, 5–6 mm long, 2.2–3 mm in diameter. Sepals 3, broadly triangular, 1–1.5 × ca. 1.5 mm. Petals 3, pink-purple, oblong to obovate-oblong, slightly oblique, 13–15 × 4–5 mm. Stamens 3, isomorphic, entirely purplish-pink and glabrous; filaments slightly flattened, 7–10 mm long; anther narrowly cylindric, incurved, subulate and falcate, 8–10 mm long, with two pores at apex; anther base forked into 2 finger-like, S-curved lobes, 2–2.5 mm long. Ovary ca. 3–4 mm long, 3-locular; ovary crown shortly cylindrical, 3-winged, ca. 1 mm high; ovules numerous. Style filiform, purple, 15–17 mm long; stigma obscurely capitate. Fruit cupuliform capsule, 6–7 × 3.5–4.5 mm.</p><p>Distribution: — Vietnam, Nghe An Province (Ky Son District). Probably stenoendemic to Phu Xai Lai Leng Mountains.</p><p>Ecology and phenology: —Lithophytic and terrestrial creeping herb. Primary humid broad-leaved evergreen highland tropical forest on sandstone and shale at elevations of 2100–2700 m a.s.l. Occasional. Flowers in September– October.</p><p>Proposed IUCN conservation status: —This species is presently known from two close locations distant for ca. 10 km from each other in the highlands of the Phu Xai Lai Leng Mountains. The current population status of the new species is unknown after the accomplishment of road network construction in the past. Therefore, its conservation status should be considered as Data Deficient (DD).</p><p>Note: —Among the known Sonerila species in Vietnam, S. cornuta bears morphological resemblances to S. cantonensis Stapf (1892: 302) . Both species share characteristics such as shorter plants, hairy stems and inflorescence, and ovate leaves. However, S. cornuta is distinct from the latter due to its lamina densely hirsute above (vs. sparsely appressed setose or strigose) with a cordate or rounded base (vs. cuneate to obtuse), and the anthers uniformly purple (vs. yellow) with two S-curved base lobes (vs. base deeply cordate). A detailed comparison of S. cornuta and two other morphologically similar species, S. reptans and S. cantonensis are presented in Table 1.</p><p>Additionally studied specimen (paratype): — VIETNAM, Nghe An Province, Ky Son District, Na Ngoi Municipality, north-eastern slopes of Phu Xai Lai Leng mountain system, primary very humid broad-leaved forest with Cunninghamia konishii Hayata as emergent on steep mountain slopes composed of gray shale at elevation 2100–2700 m a.s.l. around point 19°13’52.9’’N 104°05’30.5’’E, terrestrial creeping herb on very steep shady humid slope of northern exposition, flowers purple, locally common, 25 October 2013, L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dan, CPC 6261 (LE LE01253092 http://re.herbariumle.ru/01253092).</p></div>	https://treatment.plazi.org/id/AC2487EDFFFCB854A9EF68B9504D20AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nguyen, Khang Sinh;Averyanov, Leonid V.;Lin, Che Wei	Nguyen, Khang Sinh, Averyanov, Leonid V., Lin, Che Wei (2025): New species, Sonerila cornuta and S. phaluongensis (Melastomataceae), from northern Vietnam. Phytotaxa 696 (2): 158-168, DOI: 10.11646/phytotaxa.696.2.5, URL: https://doi.org/10.11646/phytotaxa.696.2.5
AC2487EDFFF9B857A9EF6C1651BF2A65.text	AC2487EDFFF9B857A9EF6C1651BF2A65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sonerila phaluongensis K. S. Nguyen, Aver. & C. W. Lin 2025	<div><p>Sonerila phaluongensis K.S.Nguyen, Aver. &amp; C.W.Lin, sp. nov. (Figs. 3 &amp; 4)</p><p>Type: — VIETNAM. Son La Province: Van Ho District, Chieng Xuan Municipality, Co Hong Village, Xuan Nha Nature Reserve, Pha Luong Mountain, primary coniferous and mixed forest with Pinus cernua on very steep slopes along ridge edge composed by brown sandstone at elevation of 1200–1400 m a.s.l., around point 20°41’40.5’’N 104°39’24.7’’E, terrestrial undershrub, flowers pink, anthers yellow, locally common, 13 November 2013, L. Averyanov, N. T. Hiep, N. S. Khang, N.D. Thang, L.D. Qui, CPC 7044 (holotype: LE LE 01253138 http://re.herbariumle.ru/01253138, photos LE LE 01124786 http://re.herbariumle.ru/01124786).</p><p>Diagnosis: — Sonerila phaluongensis is a distinctive species among Vietnamese Sonerila, easily distinguishable from others by its dimorphic leaves in each pair. It superficially resembles S. betongensis J.Wai &amp; J.M.Hu (2023: 114), but differs in having a 4-winged stem (vs. terete), glabrous leaves (vs. densely hispid), a cylindric hypanthium (vs. cupuliform to campanulate) that is sparsely covered with gland-tipped hispid hairs (vs. relatively densely hispid), and the anthers with two prominent basal lobes (vs. basal lobes insignificant).</p><p>Etymology:— The specific epithet refers to the name of mountain (Pha Luong), where the new species was discovered.</p><p>Description:— Terrestrial perennial herb or small undershrub. Stems suberect, few- to many-branched, quadrangular in cross section, 4-winged, up to 30 cm tall, internodes 0.8–3 cm long, 3–6 mm thick (wings 1–2 mm wide), shorter on the distal branches, glabrous; green, turning yellowish-brown and slightly lignified at age. Leaves glabrous, strongly dimorphic in each pair and held nearly horizontally. Larger leaves: petiole 0.5–1.2 cm long; leaf blade thick, obliquely oblanceolate, 4.5–11 × 1–2.5 cm, at base asymmetric, subcordate, margin denticulate on the upper two-thirds and entire on the lower one-third, apex acuminate to caudate; adaxial surface emerald green; abaxial surface pale green; venation acrodromous, with 3 or 5 veins, median nerve grooved and lateral two or four veins slightly impressed to flat on the adaxial surface, and prominent on the abaxial surface, secondary and tertiary veins inconspicuous. Smaller leaves: petiole 1–2.5 mm long; leaf blade ovate to widely ovate, 0.5–2.5(3.3) × 0.4–1.1 cm; adaxially emerald, abaxially pale green, apex obtuse, acute to shortly acuminate, base cordate, margin entire or sparsely denticulate; venation acrodromous, with 3 or 5 main veins. Inflorescence terminal or axillary, ascending, simple scorpioid cyme with 2–4 flowers; peduncles 4-winged, 0.8–1.5 cm long, green, glabrous or with few scattered purple hispid, glandular hairs. Flowers 3-merous; pedicels 3–5(10) mm long, pinkish to pale green, glabrous. Hypanthium cylindrical, pale pinkish, yellowish or greenish, sometimes with reddish tint; 5–7(8) mm long, 2–2.3 mm in diameter, with scattered minute yellowish-white or purple, hispid, glandular hairs. Sepals 3, broadly triangular, acute, 1.4–1.5 mm long and wide, with entire margins, abaxially with scattered hispid, yellowish-white to pink, glandular hairs. Petals 3, pale pink-purple, obovate-oblong to elliptic, slightly oblique, 12–14 × 6–7 mm, glabrous. Stamens 3, isomorphic, glabrous, filamentose; filaments almost terete, 12–15 mm long, white to light pinkish; anthers subulate, incurved, 10–12 mm long (appendages included), white, with green tint at the base and with light purplish-pink tint at apex, apically with two pores, at base forked into two pale yellow, short, falcate, lobes, ca. 2 mm long. Ovary ca. 2.5 mm long, 3-locular; ovary crown ca. 1 mm high; ovules numerous. Style filiform, pink to purple, 1.7–1.9 cm long; stigma obscurely capitate. Fruit cylindric to obconoid capsule, often slightly constricted at middle, 8–10 × 4–5 mm.</p><p>Distribution: — Vietnam, Son La Province (Van Ho District, Xuan Nha Nature Reserve). Probably stenoendemic to Pha Luong Mountain.</p><p>Ecology and phenology: —Terrestrial herb or small undershrub. Primary coniferous and mixed submontane forests with Pinus cernua on brown eroded sandstone at elevations of 1200–1400 m a.s.l., commonly found in shady mossy places on steep slopes, sandstone cliffs, and along ridge edges. Locally common. Flowers and fruits observed in November.</p><p>Proposed IUCN conservation status: —The new species is currently known only from a single location (Co Hong Village Area) in Pha Luong Mountain near the Vietnam-Laotian border (Son La Province, Van Ho District, and Houaphanh Province, Sop Bao District). Our two field surveys in November 2013 and September 2016 confirmed the existence of only a single population of about 30–50 mature plants. This fact gives sure evidence that Sonerila phaluongensis, like Pinus cernua Aver., K.S.Nguyen &amp; T.H.Nguyên, represents the strict endemism of Pha Luong Mountain. Though the habitat of the newly discovered species is formally protected by regional authorities from any anthropogenic disturbance, it is provisionally accessed as Critically Endangered according to criteria B1ab(ii,iii,v) + B2ab(ii,iii,v); D.</p><p>Note: — Sonerila phaluongensis somewhat resembles S. plagiocardia Diels (1932: 117) in its cylindrical hypanthium, covered with scattered hispid glandular hairs and colour scheme of anthers forked at the base into two prominent lobes. However, it differs from the latter in having dimorphic leavess, shorter petioles, oblanceolate leaf blade, and isomorphic stamens. A detailed comparison of S. phaluongensis and closely related species, S. betongensis and S. plagiocardia is presented in Table 2.</p><p>vestiture glabrous densely hispid subglabrous wings 4-winged wing absent 4-winged</p><p>Leaf dimorphic dimorphic isomorphic petiole length (cm) 0.5–1.2 (in larger leaf) 1–3.5 (in larger leaf) 2–10 ...continued on the next page</p></div>	https://treatment.plazi.org/id/AC2487EDFFF9B857A9EF6C1651BF2A65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nguyen, Khang Sinh;Averyanov, Leonid V.;Lin, Che Wei	Nguyen, Khang Sinh, Averyanov, Leonid V., Lin, Che Wei (2025): New species, Sonerila cornuta and S. phaluongensis (Melastomataceae), from northern Vietnam. Phytotaxa 696 (2): 158-168, DOI: 10.11646/phytotaxa.696.2.5, URL: https://doi.org/10.11646/phytotaxa.696.2.5
