taxonID	type	description	language	source
B74A87D8C517987FFF71FB25FC2CF837.taxon	distribution	Distribution. China, India, Indonesia, Japan, Korea, Malaysia, Myanmar, Nepal, Russia, Sri Lanka, Thailand, The Philippines, Vietnam (Betrem 1928; Gupta & Jonathan 2003; Liu et al. 2021 a; Pham & van Achterberg 2023).	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C514987DFF71F8DCFB9DFEB2.taxon	materials_examined	Type material examined. Holotype (CCEC), ♀, ‘ Philippines, Mindanao, Sarangani, Kiamba, I. 2016, Ismaël Lumawig leg. ’. Paratypes: 1 ♂, idem holotype (CECC); 1 ♂, Bukidnon, Cabanglasan, II. 2017; 1 ♂, Davao oriental, Mati, V. 2017; 1 ♂, Lanao du Nord, Kapatagan, III. 2016; 1 ♂, idem, VII. 2017; 1 ♀, Davao oriental, Governor Generoso (Sigaboy), VIII. 2017; 1 ♀, Cotabato du Sud, mont Parker, IV. 2014; 1 ♀, Agusan del Sur, San Francisco, X. 2017; 1 ♀, Mindoro oriental, Baco, III. 2014; 1 ♀, Iloilo, Leon, V. 2016 (all Ismaël Lumawig leg., CJBC). Other material examined. The Philippines. Mindanao. Agusan del Sur province. 1 ♀ 1 ♂, San Luis, XII. 2022; 1 ♀ 2 ♂, San Felomina, XII. 2022. (all Ismaël Lumawig leg.) [all CJBC].	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C514987DFF71F8DCFB9DFEB2.taxon	distribution	Distribution. The Philippines (Mindanao, Luzon, and Panay) (Castagnet 2021 a).	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C5159876FF71FE45FF15FC56.taxon	description	urn: lsid: zoobank. org: act: 8 C 23 E 594 - 6 BA 5 - 46 EA-BAB 2 - E 32501 B 44 BA 2 (Figs 1 – 3)	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C5159876FF71FE45FF15FC56.taxon	materials_examined	Type material. Holotype (CCEC), ♂, [red label] ‘ PHILIPPINES / Mindanao Isl. / Surigao del Sur pr. / Tandag, IX. 2017 / Local collector leg. ’, [red label] ‘ HOLOTYPE ♂ / Megacampsomeris funesta / Castagnet & Cabon, 2025 ’. Paratypes: 1 ♀ (CCEC), ‘ PHILIPPINES, Mindanao Isl., Davao Oriental pr., Governor Generoso, X. 2017 / Local collector leg. ’, ‘ PARATYPE ♀ / Megacampsomeris funesta / Castagnet & Cabon, 2025 ’; 1 ♀ (CJBC) ‘ PHILIPPINES / Mindanao Isl. / Lanao del Sur pr. / Amai Manabilang, II. 2024 / Local collector leg. ’, second label same as the precedent; 2 ♂ (CJBC), ‘ PHILIPPINES / Mindanao Isl. / Surigao del Sur pr. / San Agustin, III. 2023 / Local collector leg. ’, ‘ PARATYPE / ♂ / Megacampsomeris funesta / Castagnet & Cabon, 2025 ’; 1 ♂ (CFC), ‘ PHILIPPINES / Mindanao Isl. / Davao del Sur pr. / Tamayong, II. 2021 / Local collector leg. ’, second label same as the precedent.	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C5159876FF71FE45FF15FC56.taxon	diagnosis	Diagnosis. M. funesta sp. nov. is distinguished in females from all other species of Megacampsomeris by having the punctation of the frons and vertex dense, very fine, and largely coalescent; and the metanotum with extremely fine, dense, and coalescent punctation. In contrast, in other Megacampsomeris species, the punctation of the frons and / or vertex is always coarser, partly more widely spaced, or with more or less extensive impunctate areas; and the metanotum always has a coarser punctation than in M. funesta. Moreover, most of Megacampsomeris female (M. asiatica (Saussure, 1858), M. schultessi Betrem, 1928, M. grossiformis Betrem, 1928, M. farrenwithei Betrem, 1928, M. grossa (Fabricius, 1804), M. mojiensis (Uchida, 1933), M. uchidai Betrem, 1928, M. cochinensis Betrem, 1928, M. binghami Betrem, 1928, M. formosensis Betrem, 1928, M. limbata (Saussure & Sichel, 1864), M. lindenii (Lepeletier, 1845), M. prismatica (Smith, 1855), M. ceylonica (Kirby, 1889), M. pulchrivestita (Cameron, 1902), M. acutinerva (Cameron, 1902), M. shillongensis Betrem, 1928, M. dehraensis Betrem, 1928, M. limbangensis Betrem, 1928, M. montanum Gupta & Jonathan, 2003, M. nilgirensis Gupta & Jonathan, 2003, M. westermanni Betrem, 1928) have the first tergites covered with colored pubescence (whitish, yellowish, orange, or brownish); whereas in M. funesta sp. nov. the metasomal pubescence is entirely black. Only M. luzonensis (Rohwer, 1921), comb. nov., M. reticulata (Cameron, 1892), M. vanoordti Betrem, 1928, M. habrocoma, and M. bitschi Castagnet, 2021 also have the metasoma entirely covered with black pubescence. The males of Megacampsomeris species are very similar, but M. funesta sp. nov. can be distinguished from those of other Megacampsomeris species by the following combination of characters: the mesosoma with brown pilosity; the metasoma entirely melanistic covered with black pilosity; and distinctly yellowish wings. In contrast, the other species never exhibit this exact combination: the mesosoma has whitish, yellowish, orangish, brownish or black pilosity; the metasoma often has yellowish markings; the basal tergites usually have colored pilosity (whitish, yellowish, orange, or brownish); and the wings may be either dark or yellowish. M. habrocoma may present a somewhat similar habitus; for further details, see the identification key and remarks.	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C5159876FF71FE45FF15FC56.taxon	description	Description of the male (Fig. 1). Length 25 – 29 mm. Head (Figs 1 B, C). Inner margin of mandibles with three median teeth. Clypeus impunctate medially, with coarse to fine punctures laterally and basally. Frontal spatium with dense, fine, tight and coalescent punctures. Fissura frontalis weakly impressed, ending in frontal pit. Transversal groove delimiting frontal space weakly visible. Frons with more or less coarse and dense punctures, with a fine impunctate area behind median ocellus. Vertex almost impunctate between eye and posterior ocellus, with very dense and fine punctures behind posterior ocelli. Mesosoma (Fig. 1 D). Pronotum with very dense, fine and tight punctures. Scutum with a medially elevated area; with coarse, dense and tight punctures, sparser punctures medially and with a V-shaped impunctate area. Scutellum with very dense and fine punctures, largely impunctate apically. Metanotum with very fine, tight and almost coalescent punctures. Mesopleuron with dense, fine and tight punctures, with a fine area impunctate or almost so along metapleuron. Upper plate of metapleuron with scattered, fine and shallow punctures medially. Lower plate of metapleuron largely punctate; with dense, coarse and deep punctures. Lateral area of propodeum with dense, coarse and deep punctures; at level of transition between lateral and dorso-lateral areas of propodeum with very dense and coalescent punctures. Lateral carina of propodeum weakly visible. Dorso-median area of propodeum with fine, dense, tight and almost coalescent punctures. Dorso-lateral area of propodeum with fine, dense, tight and almost coalescent punctures, without distinct impunctate area basally (mirror). Postero-median and lateral areas of propodeum with fine, dense, tight and coalescent punctures. Metasoma (Fig. 1 E). T 1 with dense, coarse and deep punctures. T 2 - T 3 with dense, coarse and deep punctures; medially and apically in middle with sparser and shallower punctures. T 4 - T 6 with similar punctation. S 1 with a transverse groove crossing entire sternite; with dense, coarse and deep punctures, weakly impunctate apically. S 2 with distinct tubercle; with dense and coarse punctures, sparser in middle. S 3 - S 5 similarly punctate, with sparser punctures medially. Genitalia. See Fig. 3 A. Coloration (Fig. 1). Head entirely black with brown / black pilosity. Mesosoma black with brown-black pilosity and tawny tomentosity mainly on pronotum, scutum and scutellum. Legs black with black pilosity. Metatibial spurs dark orangish. Wings yellowish subhyaline. Metasoma black with green / violet iridescence, with black pilosity. Female (Fig. 2). Length 35 – 37 mm. Head (Figs 2 B, C). Clypeus wrinkled apically, impunctate medially, basally and laterally with very dense and fine punctures. Frontal spatium with very dense, tight, fine and almost coalescent punctures. Fissura frontalis visible, reaching median ocelli. Frons with fine, dense, and tight punctures in eye notch and laterally; with dense but more widely spaced punctures in front of median ocellus; and with a group of dense, tight, and fine punctures in front of posterior ocelli. Along eye and eye notch, with a sharp raised and impunctate area. Vertex with a fine impunctate band behind posterior ocelli, and with dense, fine punctures on remaining surface. Declivous part of vertex bears very dense, tight, and fine punctures; area behind eyes largely impunctate. Temporal groove weakly visible. Mesosoma (Fig. 2 D). Pronotum with very dense, tight, fine and coalescent punctures. Scutum with very dense, deep and coalescent punctures, a fine impunctate band laterally along tegula and medially with a large V-shaped impunctate area; with micropunctation over almost entire surface, except on impunctate area. Scutellum with very dense, fine, and coalescent punctures, except for a narrow impunctate band medially and apically. Metanotum very dense, tight, fine and coalescent punctures, except for a narrow impunctate band medially. Upper plate of mesopleuron without tubercle; with dense and fine punctures medially. Lower plate of mesopleuron with a weakly visible epicnemial carina; with dense and fine punctures along epicnemial carina. Upper plate of metapleuron with fine, dense, and tight punctures on upper quarter. Lateral area of propodeum largely impunctate along lower plate of metapleuron, with dense, deep and coarse punctures along dorso-lateral and postero-lateral areas of propodeum. Lateral carina of propodeum slightly exceeding spiracle. Dorso-lateral area of propodeum with an impunctate area (mirror); with dense, tight and fine punctures on remaining surface. Dorso-median area of propodeum with dense, tight deep, coarse and coalescent punctures; transition zone between dorso-median and postero-median areas of propodeum with a raised crest projecting backward. Postero-median area of propodeum with dense; deep and coarse punctures. Postero-lateral area of propodeum with dense, shallow, tight and fine punctures. Metatibial spur dark orangish. Forewing with two recurrent veins. Metasoma (Fig. 2 E). T 1 almost impunctate medially; with dense and fine punctures laterally and apically. T 2 - T 3 with dense, shallow and fine punctures basally; largely impunctate medially; with pre-apical band impunctate or nearly so. T 4 - T 5 similarly punctate with denser and coarser punctures. S 1 with fine and dense punctures laterally, impunctate on remaining surface. S 2 with coarse punctures basally; impunctate medially, with few scattered dots; a row of coarse and deep punctures largely interrupted apically in middle. S 3 - S 4 similarly punctate. Coloration (Fig. 2). Head black with brown / black pilosity, except declivous part of vertex with tawny tomentosity. Mesosoma black with brown-black pilosity and tawny tomentosity mainly on scutum. Legs black with black pilosity. Metatibial spurs orange-brownish. Wings yellowish subhyaline. Metasoma black and mat (mainly on T 2 - T 3), with black pilosity.	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C5159876FF71FE45FF15FC56.taxon	etymology	Etymology. The name funesta refers to the dark color of the species.	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C5159876FF71FE45FF15FC56.taxon	biology_ecology	Biology. Unknown.	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C5159876FF71FE45FF15FC56.taxon	distribution	Distribution. The Philippines (Mindanao) (Fig. 4).	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C5159876FF71FE45FF15FC56.taxon	diagnosis	Comparative diagnosis. This taxon is assigned to the genus Megacampsomeris because it shares the following generic characteristics with Megacampsomeris: in females, posterior tibial spurs are whitish or orangish; frons without deep punctate cluster in front of the median ocellus; excavation of the pronotum short and weakly defined, sometimes partially limited by a sharp margin; first submarginal cell glabrous, except near the margin of the stigma and costal area, where the pilosity may extend medially. In males, the posterior tibial spurs are whitish or orangish, and the genitalia are characteristic of the genus Megacampsomeris: lamina volsellaris densely punctate, bearing long, erect setae; basal part of the volsellae with such dense and closely spaced punctures that the bases of the setae are almost invisible, with long and erect pilosity, forming a brush. Three species are particularly close to M. funesta and can be distinguished from M. funesta as follows: M. funesta sp. nov. is easily distinguished from M. luzonensis in females, by the dense punctation of the head and metasoma, whereas in M. luzonensis, both the head and metasoma exhibit large impunctate areas. M. funesta sp. nov. can be distinguished from M. bitschi by the frons and vertex are covered with very dense, fine, tight, and almost coalescent punctures; the metanotum bears very dense, fine, and coalescent punctures; the dorso-median area of propodeum has a distinct and prominent apical ridge, with very dense, coarse, and coalescent punctures; and the postero-lateral and median areas of propodeum show dense, coarse, and deep punctures. In contrast, in M. bitschi, the frons and vertex are densely but more sparsely and coarsely punctate; the metanotum has dense, coarse punctures that are clearly not coalescent; the dorso-median area of propodeum lacks a distinct apical ridge, and is punctate with dense, coarse, but clearly non-coalescent punctures; the postero-lateral and median areas of propodeum are covered with scattered, minute, and shallow punctures. M. habrocoma (Smith, 1855), recorded from Indonesia (Betrem, 1928) (material examined: Indonesia: West Java province. 3 ♀ 3 ♂, Mount Halimun, IX. 2017 (local collector leg.) [CJBC]), exhibits a habitus very similar to that of M. funesta sp. nov. in females: entirely melanistic, with black to brownish pilosity and clear wings that are not entirely dark. However, females of M. funesta sp. nov. can be distinguished from M. habrocoma by the following characters: the frons and vertex are covered with very dense, fine, tight, and almost coalescent punctures; the metanotum bears very dense, fine, and coalescent punctures; the dorso-median area of propodeum has a distinct and prominent apical ridge, with very dense, coarse, and coalescent punctures; and the postero-lateral and median areas of propodeum show dense, coarse, and deep punctures. In contrast, in M. habrocoma, the frons and vertex are densely but more sparsely and coarsely punctate; the metanotum has dense, coarse punctures that are clearly not coalescent; the dorso-median area of propodeum shows only a weakly visible apical ridge, and is punctate with dense, coarse, but clearly non-coalescent punctures; the postero-lateral and median areas of propodeum are covered with scattered, minute, and shallow punctures. In males, M. funesta sp. nov. can be distinguished from M. luzonensis, M. habrocoma and M. bitschi by the dorso-median area of propodeum, which bears fine, dense, tight, and almost coalescent punctures; the lower plate similarly covered with dense, coarse, and deep punctures, and at the transition between the lateral and dorso-lateral areas, the punctures become very dense and coalescent. In contrast, in M. luzonensis, M. habrocoma and M. bitschi, the dorso-median area of propodeum has coarser and sparser punctures; the lower plate of the metapleuron is weakly punctate, with scattered, fine, and shallow punctures; the lateral area of propodeum shows sparse, fine, and shallow punctures, and at the transition between the lateral and dorso-lateral areas, the punctures are dense and almost coalescent. Two other species known only by females, M. malaccensis and M. laevivertex from Indonesia (in the vicinity of Malacca), were reported to have a similar habitus to that of the new species. M. laevivertex differs from M. funesta sp. nov., according to Betrem’s (1928) description, by the punctation of the vertex, which is entirely impunctate, whereas in M. funesta sp. nov. the vertex is densely and finely punctate. M. malaccensis differs from the new species, according to Betrem’s (1928) description, by having the vertex with very coarse and deep punctures, the scutum being entirely densely punctate, the dorso-median area of propodeum punctate like the metanotum, and by a uniformly charcoal-black pilosity, almost lacking any tomentosity. In contrast, in M. funesta sp. nov., the vertex is densely and finely punctate, the scutum shows a large V-shaped impunctate area medially, the punctation of the dorso-median area of propodeum is much coarser than that of the metanotum, and the pilosity is brownish to black, with a dense tawny tomentosity. The holotype specimens of M. malaccensis and M. laevivertex were originally deposited in the Federated Malay States Museums (formerly the Selangor Museum and now Muzium Negara). Unfortunately, during World War II, part of the Federated Malay States Museums was destroyed by bombing carried out by the Allied Air Force on 10 March 1945. As a result, most of the collections were lost, and these two holotypes remain untraceable in the collections of Muzium Negara to this day (I. M. Abdullah & M. K. Jemangin com. pers.).	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C51E9874FF71F8B8FBCCFC8A.taxon	materials_examined	Type material examined. Holotype of Scolia luzonensis (USNM), ♀, [white label] ‘ Scolia luzonensis Type ♀ Roh. ’, [red label] ‘ Type No. 23587 U. S. N. M. ’. Holotype of Scolia panayensis (USNM), ♂, [white label] ‘ Scolia panayensis type ♂ ’, [red label] ‘ Type No. 41539 U. S. N. M. ’, [white labels] ‘ USNMENT 01546675 ’, ‘ Culasi 6. 20.18 ’, ‘ Scolia I ♂ ’, ‘ Culasi Panay PI June 1918 McGregor’. Holotype of Campsomeris pseudopropodealis (NMKB), ♂, [white labels] ‘ C. pseudopropodealis Holotypus Betr det. Betrem’, ‘ Zamboanga Mindanao Schulze’, [red label] ‘ holotype’. Other material examined. Philippines. Mindanao. Agusan del Sur province. 1 ♀, Esperanza, VIII. 2014; 2 ♀, Prosperidad, III. 2016; 1 ♀ 1 ♂, San Francisco, IV. 2019. Bukidnon province. 1 ♂, Impasug-ong, IV. 2014; 1 ♀ 1 ♂, Intravas, IV. 2014; 1 ♂, Panamokan, V. 2014; 1 ♀, Dominorog, XII. 2018; 1 ♀, same place, I. 2019; 1 ♂, Kabanglasan, III. 2018. Catobato province. 1 ♂, Kidapawan, I. 2016; 1 ♀, Alamada, IV. 2019; 2 ♀, same place, X. 2019. Catobato del Sur province. 1 ♂, mont Parker, IV. 2014. Davao Oriental province. 1 ♂, Mati, V. 2017. Davao del Sur province. 3 ♀, Kapatagan, IV. 2018; 5 ♀, same place, IV. 2019. Davao de Oro province. 5 ♀, Maragusan, IV. 2018. Lanao del Norte province. 1 ♂, Kapatagan, VIII. 2015; 1 ♂, same place, XII. 2015; 1 ♂, same place, III. 2016; 3 ♂, same place, VII. 2016; 1 ♂, same place, I. 2017; 2 ♂, same place, VII. 2017; 2 ♀, same place, IV. 2018. Lanao del Sur province. 1 ♂, Wao, V. 2018; 2 ♀ 1 ♂, same place, VI. 2018; 1 ♂, same place, X. 2018; 1 ♀, Maguing, X. 2018. Misamis Oriental province. 2 ♀ 1 ♂, Balingasag, IV. 2014; 1 ♂, same place, VI. 2014. Sarangani province. 2 ♂, Kiamba, I. 2016; 1 ♀, same place, II. 2017. Surigao del Sur province. 1 ♀, Barobo, II. 2019; 1 ♂, Luagonj, III. 2019; 1 ♀, Lingig, III. 2017. Luzon. Aurora province. 1 ♀ 1 ♂, Dingalan, VIII. 2017; 1 ♀, same place, III. 2018. Mindoro Oriental province. 1 ♂, Baco, VIII. 2017. Negros. Negros occidental province. 1 ♀, Salvador Benedicto, VIII. 2015; 1 ♂, same place, I. 2019. Negros oriental province. 1 ♂, Dumaguete, V. 2015. Samar. Samar del Norte province. 1 ♀, Lope de Vega, II. 2017. (all local coll. leg.) [all CJBC].	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C51E9874FF71F8B8FBCCFC8A.taxon	distribution	Distribution. Philippines (Mindanao, Luzon, Visayas) (Rohwer 1921; Betrem 1928; Castagnet 2021 a).	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
B74A87D8C51E9874FF71F8B8FBCCFC8A.taxon	discussion	Remarks. Based on morphological criteria, the taxa luzonensis and panayensis were synonymized and transferred to the genus Laevicampsomeris from the genus Megacampsomeris (Castagnet 2021 b). Indeed, females of luzonensis share the following characteristics with species of the genus Laevicampsomeris: the upper part of the frons, vertex, center of scutum, scutellum, and metanotum are largely impunctate; the pronotum, in lateral view, forms a prominent crest; and the dorso-median area of propodeum is clearly wider than long. In both sexes, one recurrent vein is also present. However, the female of luzonensis clearly differs from other Laevicampsomeris species by its entirely punctate propodeum, whereas in Laevicampsomeris species the propodeum is almost completely impunctate. In males, luzonensis is distinguished by an elongated T 1 (much longer than wide), while in Laevicampsomeris it is shorter (slightly longer than wide). Examination of a larger number of L. luzonensis specimens, as well as of L. solomonis Krombein, 1963, L. bonguensis Betrem, 1933, and particularly of the male genitalia, reveals my (JBC) earlier mistake in assigning luzonensis to the genus Laevicampsomeris. The taxon luzonensis, although atypical and sharing several traits with Laevicampsomeris, should instead be transferred to the genus Megacampsomeris. Indeed, in both sexes of the taxon luzonensis, the tibial spur is orange in the panayensis form, but slightly darker in the typical form (luzonensis), as is the case in Megacampsomeris, which consistently exhibit pale metatibial spurs. This contrasts with Laevicampsomeris, in which the metatibial spurs are always black. The presence of one recurrent vein is sometimes observed in Megacampsomeris, which usually have two recurrent veins (notably M. bitschi and M. acutinerva), whereas Laevicampsomeris always has only one recurrent vein (Betrem 1928; Castagnet 2021 a). Therefore, this character alone cannot be used to distinguish between the two genera. My error (JBC) stemmed from the fact that, in females, luzonensis exhibits an atypical reduction of mesosoma punctation, and that the pronotum bears a broad oblique groove at the boundary between the dorsal and lateral surfaces, forming a prominent crest in lateral view, generally less pronounced in Megacampsomeris, though exceptions exist, such as M. farrenwhitei Betrem, 1928 (Liu et al. 2021 b), which shows similarly reduction of mesosoma punctation; and the dorso-median area of propodeum is clearly wider than long. However, females of luzonensis remain more densely punctate, particularly across the entire propodeum, than those of Laevicampsomeris, in which the propodeum is largely impunctate. Furthermore, the dorso-median area of propodeum in Laevicampsomeris is nearly four times wider than long, whereas in luzonensis it is approximately three times longer than wide. The examination of the genitalia of L. luzonensis (see Castagnet 2021 b), M. bitschi (see Castagnet 2021 a), M. funesta sp. nov. (Fig. 3 A), and representatives of the genus Laevicampsomeris (Fig. 3 B) clearly confirm that luzonensis shares the same characteristics as species of the genus Megacampsomeris. To avoid further confusion resulting from my earlier misplacement of the taxon luzonensis within the genus Laevicampsomeris (Castagnet 2021 b), the following characters can be used to distinguish the genera Megacampsomeris and Laevicampsomeris. The genus Megacampsomeris is distinguished in both sexes by having pale metatibial spurs (ranging from whitish to dark orangish), whereas those of Laevicampsomeris are always black. In Megacampsomeris, the forewing generally has two recurrent veins, sometimes only one, whereas Laevicampsomeris always has a single recurrent vein. In females of Megacampsomeris, the punctation is variable, sometimes largely impunctate on the vertex, scutum, and scutellum, whereas in Laevicampsomeris, the head (mostly) and mesosoma (except for the pronotum and basal part of the scutum) are almost entirely impunctate. In males, the genitalia offer a clear distinction between the two genera. In Megacampsomeris, the paramere is long, slender, and apically pointed. The lamina volsellaris is densely punctate and bears long, erect setae. The basal part of the volsellae is so densely and finely punctate that the bases of the setae are almost indistinguishable; the pilosity is long and erect, forming a brush-like structure. In contrast, in Laevicampsomeris, the paramere is short and thick, broadly rounded apically. The lamina volsellaris is less densely punctate and bears short setae. The basal part of the volsellae, more sparsely punctate, has visible seta bases; the pilosity is short, sparse, and does not form a brush. The genus Ilkamilka Argaman, 1996, previously treated incorrectly as a synonym of Laevicampsomeris (Castagnet 2021 b), must now be considered a synonym of Megacampsomeris.	en	Castagnet, Jean-Baptiste, Cabon, Flavien (2025): A new melanistic species of Megacampsomeris Betrem, 1928 from the Philippines and taxonomic reassessment of Laevicampsomeris luzonensis (Rohwer, 1921) (Hymenoptera, Scoliidae, Campsomerini). Zootaxa 5701 (5): 573-585, DOI: 10.11646/zootaxa.5701.5.6, URL: https://doi.org/10.11646/zootaxa.5701.5.6
