taxonID	type	description	language	source
B448EA62FFE9B276FE97F9093B4C4EF6.taxon	description	(Fig. 20 A – E)	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE9B276FE97F9093B4C4EF6.taxon	materials_examined	Material examined: 1 ♀, Myanmar: Myita, Doherty ‘ Drabescus laetisigna Walk’. (NHM). Remarks: Tis species was described from an unknown number of female specimens (syntypic) with the following data: ‘ Birmah. (Mus. Holm.) ’. Te depository refers to the Museum Holmen in Copenhagen, Denmark. Distant (1908: 307) recorded the specimen examined above from ‘ Tennasserim; Myita (Doherty) ’ under the name Drabescus laetisigna Walker (see below). Te specimen he subsequently recorded from ‘ Lower Burma: Mergui (W. Doherty) ’ is missing from its mount in NHM and which erroneously bears a red holotype disc. As both Zhang and Webb (1996) and Viraktamath (1998) referred to seeing the Stål holotype, this specimen should be regarded as the lectotype, with the former credited with the designation, as Stål made no mention of specimen number.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE8B276FDADF94B39754EFE.taxon	description	(Fig. 21 A – K)	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE8B276FDADF94B39754EFE.taxon	materials_examined	Material examined. Lectotype ♂ (designated by Zhang and Webb 1996: 20), Malaysia: Sarawak, NHMUK 013589075 (NHM); 1 ♂, Malaysia: Sarawak, Semongoh Forest Reserve, 01 º 25 ' N, 110 º 17 ' E, 15 – 19. xi. 1976, P. Cranston (NHM); 1 ♂, Malaysia, Sabah, Sandakan, (USNM); 1 ♂, Malaysia: Sabah; Danum Valley (05 º 01 ′ N, 117 º 47 ′ E), 30 September 1987, 200 m, light trap sample, understory forest, lowland mixed dipterocarp forest; Grid No. 52, coll. A. H. Kirk-Spriggs (NMW). Distribution: Malaysia (Sabah, Sarawak). Remarks: Tis species was described from an unspecified number of male and female specimens (syntypic) from Borneo, collected by Wallace. Te lectotype bears a circular label with ‘ SAR’ (hand-writen) and a rectangular label with ‘ Sarawak’ (printed) and another with ‘ 68 4 ’ (printed). Te later number refers to an entry in the NHM register for 1868 4 stating that the collection lot originated from various localities donated by ‘ Saunders’. Labels with this registration number are frequently found on Wallace collected specimens from Borneo in the NHM.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE9B277FC4CFDB939C94EB5.taxon	materials_examined	Type species: Bythoscopus laetisigna Walker, 1857. Diagnosis: Similar in coloration and structure to Malaysiapona and Halimunella but differs in having the aedeagus symmetrical with an unpaired process arising near the base of the shaf and extending distad along the shaf, and the style apex digitiform without a pre-apical angulate projection. Description: Body olive green. Crown with anterior half bright orange, without black spots. Eye orange with round black medial spot. Face unmarked. Pronotum with paired antero-medial, medial, and posterolateral symmetrical orange markings. Mesonotum and scutellum each with pair of submedial orange markings, apex of scutellum orange. Forewing smoky hyaline with costal and commissural margins orange and orange spot at base of corium, apex of corium and first two R branches broadly black. Front and middle tibia orange. Body oblong, relatively broad, slightly depressed. Head slightly broader than pronotum; anterior margin relatively thick in profile, weakly bicarinate with few additional fine transverse striations. Ocelli small, situated in sulcus of foremargin, separated from adjacent eye by distance equal to own diameter. Crown flat, slightly depressed behind anterior margin, anterior margin broadly rounded in dorsal view, only slightly longer medially than next to eye. Face with lateral frontal sutures strongly divergent dorsad of antennal pits, extended to ocelli; frontoclypeus relatively narrow, distinctly swollen dorsally, depressed ventrally; clypeal suture straight; anteclypeus with lateral margins concave, apex expanded; lora broader than basal width of anteclypeus; gena emarginate below eyes; antenna longer than half-body length; antennal pits situated near upper corner of eyes, encroaching onto clypeus; antennal ledges moderately developed and oblique but not carinate. Pronotum well produced between eyes, hind margin slightly concave; lateral margin short, carinate; posterior two-thirds with irregular transverse striations; mesonotum and scutellum together slightly broader than long; scutellar suture arcuate. Forewing with four apical cells and three subapical cells, inner subapical cell open; two anterior branches of R reflexed; appendix broad. Fore femur with AM 1 near midheight of femur, AV with few stout setae in basal half; IC with series of several fine, close-set setae; fore tibia with dorsal setal arrangement of rows AD and PD: 1 (small stout seta) + 5. Hind femur macrosetae 2 + 2 + 1. Male genitalia: Pygofer dorsal bridge moderately long; lobe triangularly produced, tapered to narrowly rounded apex, with numerous macrosetae, posterior margin denticulate, processes absent. Valve broadly triangular. Xth segment moderately long, well sclerotized dorsally and laterally, anterodorsal margin V-shaped in dorsal view. Subgenital plate with base broad and rounded laterally, apex atenuate, macrosetae absent, scatered long fine setae present. Style apophysis stout, digitiform, without angulate lateral process, finely denticulate, pre-apical lobe poorly developed. Connective small, Y-shaped with stem broadened apically and slightly longer than arms, arms thickened and weakly divergent. Aedeagus with dorsal apodeme moderately well developed, column-like; shaf moderately long, tubular, curved dorsad with unpaired process arising near base and extended distad along shaf; gonopore large, apical; preatrium absent. Small sclerite articulated between apex of connective and base of aedeagus. Etymology: Te genus name was formed by combing the Latin prefix ‘ aurantico - ’ (orange), referring to the bright orange coloration, with ‘ - pona ’ the suffix of the name of the type genus (Parabolopona) of the subtribe to which the new genus belongs. Gender: feminine. Distribution: Malaysia (Sabah, Sarawak), Myanmar. Remarks: Auranticopona comprises two previously described species, both of which are here transferred into the genus from Roxasella: A. laetisigna (Walker, 1857) comb. nov. and A. egregia (Stål, 1864) comb. nov .. Te later species is known only from the female syntype but is placed here based on its close morphological similarity to the type species, the male genitalia of which were illustrated by Zhang and Webb (1996). In our phylogenetic analysis based on combined morphological and molecular data, the two included species of this genus formed a polytomy with the clade comprising Halimunella and Malaysiapona. Tese three genera are very similar in overall structure and coloration but have distinctive male genitalia. Te phylogenetic position of A. egregia was poorly resolved because only female characters could be scored for the analysis and molecular data could not be obtained. Checklist of Auranticopona species	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD4B24FFECEF8F23B7E4A52.taxon	description	(Fig. 24 A – I)	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD4B24FFECEF8F23B7E4A52.taxon	materials_examined	Material examined: Lectotype ♀ (designated by Zhang and Webb 1996: 18); Myanmar, Mergui, W. Doherty (NHM); 1 ♂ China, Yunnan Prov., Mengla Wangtianshu landscape, 21 April 2018, coll. Lu Lin. Remarks: Tis species was described from an unspecified number of specimens (syntypic) with the following data: ‘ Lower Burma; Mergui (W. Doherty) ’. It can be distinguished by its distinctive orange and dark brown markings including a dark spot against each ocellus. Roxasellana Zhang and Zhang	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD4B24FFECEF8F23B7E4A52.taxon	distribution	Distribution: China and Vietnam. Checklist of Roxasellana species	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD4B24FFECEF8F23B7E4A52.taxon	discussion	Remarks: Roxasellana is similar to Roxasella and Tenompoella based on the characters of bright orange markings on the head, thorax, and forewings and shape of the anterior margin of the vertex. It can be distinguished from the former by the aedeagus situated dorsad of the connective and linked to it by membrane, and from the later by the absence of any processes of the connective, forewing with large inner apical cell, and dorsal margin of face markings lacking a dark band.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD6B248FEBDFC0F3E294EC8.taxon	description	(Fig. 22 A – Q) (URN: LSID: ZOOBANK. ORG: ACT: D 6 B 08017 - C 0 D 8 - 4 C 49 - B 9 C 5 - 9 DF 9 D 9 EE 505 C) Description: Length (including tegmen): male 7.5 mm. Fore femur IC with about 10 setae, AV with six stout setae. Male genitalia: Style apex bilobed, pre-apical lobe not developed. Aedeagus with shaf smooth and relatively short, in lateral view strongly curved, slightly broader near the middle and tapered to apex. Paraphysis larger than aedeagus, expanded at base with dorsal surface rough in lateral view and curved strongly dorsad, apex forked in ventral view with branches strongly divergent and tapered distally. Material examined: Holotype: ♂, Philippines, Bukidnon, Brgy Impalutao, CEDAR [Center for Ecological Development and Recreation], (08 ° 2509 ' N, 125 ° 034 ' E), 780 m, 7 July 2010, coll. J. Jones (PNM). Etymology: Tis species name is a Latin adjective meaning ‘ forked’, referring to the bifurcate paraphysis between the aedeagus and connective.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE8B248FEF5F943399B4BB3.taxon	materials_examined	Type species: Paraphysipona furculapona sp. nov. Diagnosis: Tis genus resembles Roxasella and Malaysiapona in coloration, head structure, wing venation, and leg chaetotaxy but differs in having three prominent black spots along the anterior margin of the crown, an elongate connective with stem longer than arms, a simple aedeagus without processes, and a paraphysis articulated between the aedeagus and connective. Description: Coloration: Body olive green. Crown with three prominent black spots on apical margin between eyes and two larger bright orange spots between black spots. Face with thin black transverse band just below dorsal margin. Pronotum with paired anteromedial, medial and posterolateral pairs of symmetrical orange markings. Mesonotum each with pair of orange spots, apex of scutellum orange. Forewing smoky hyaline with costal and commissural margins orange and orange spot at base of clavus, first two branches of R and apices of claval veins black. Legs unmarked. Structure: Body oblong, relatively broad, somewhat depressed. Head slightly broader than pronotum; anterior margin relatively thin in profile, weakly bicarinate with few additional fine transverse striations. Ocelli small, situated in sulcus of foremargin, separated from adjacent eye by distance equal to own diameter. Crown flat, strongly depressed behind anterior margin, anterior margin broadly rounded in dorsal view, approximately as long medially than the length next to eye. Face with lateral frontal sutures strongly divergent dorsad of antennal pits, extended to ocelli; frontoclypeus relatively narrow, lightly swollen dorsally, depressed ventrally; clypeal suture straight; anteclypeus with lateral margins concave, apex expanded; lora broader than basal width of anteclypeus; gena strongly emarginate below eyes; antenna longer than half-body length; antennal pits situated near upper corner of eyes, encroaching on to frontoclypeus; antennal ledges moderately developed and oblique but not carinate. Pronotum well produced between eyes, hind margin concave; lateral margin short, carinate; posterior two-thirds with irregular transverse striations. Mesonotum and scutellum together distinctly broader than long; scutellar suture arcuate. Forewing with four apical cells and three subapical cells, inner subapical cell open; two anterior branches of R reflexed; appendix broad. Fore femur with AM 1 near midheight of femur, AV with few stout setae in basal half; IC with series of several fine, close-set setae; fore tibia with dorsal setal arrangement of rows AD and PD: 6 + 4. Hind femur macrosetae 2 + 2 + 1. Male genitalia: Pygofer dorsal bridge moderately long; lobe attenuate, tapered to narrowly rounded apex, posterior margin with dense row of short, stout setae, macrosetae numerous and scattered over distal half, processes absent. Valve broadly triangular. Xth segment moderately long and well sclerotized dorsally and laterally, anterodorsal margin V-shaped in dorsal view. Subgenital plate with base broad and rounded laterally, apex atenuate and weakly sclerotized, macrosetae absent, scatered long fine setae present. Style apophysis stout with lateroposterior angle acutely produced (avicephaliform), finely denticulate, pre-apical lobe poorly developed. Connective small, Y-shaped with stem broadened and truncate apically, much longer than arms, arms slender and bowed laterad. Single paraphysis present, well sclerotized and articulated between connective apex and base of aedeagus, branched in distal half. Aedeagus with dorsal apodeme short, rounded in lateral and posterior views; preatrium absent; shaf simple, U-shaped in lateral view, without processes. Gonopore pre-apical on posterior surface. Female unknown. Etymology: Te genus name combines the prefix ‘ paraphysi - ’ referring to the structure articulated between the connective and aedeagus, with ‘ - pona ’, the suffix of the type genus (Parabolopona) of the subtribe to which this new genus belongs, Paraboloponina. Gender: feminine. Distribution: Philippines. Remarks: Phylogenetic analysis of combined morphological and DNA sequence data placed the type species of this monotypic genus on an independent branch within a larger clade comprising Malaysiapona, Halimunella, Tenompoella, Roxasellana, and Nakula. It resembles these other genera in having bright orange coloration on the dorsum of the head and thorax but differs in having three prominent black spots along the anterior margin of the crown (as in Philipona) and in having a paraphysis articulated between the connective and aedeagus. Some other members of this clade (Fig. 3, Clade 2) have one or more processes fused to the base of the aedeagus (Malaysiapona and Roxasellana) or have the posterior stem of the connective bearing a process (Tenompoella); the condition in Paraphysipona seems to be intermediate between these two conditions.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD6B24AF90BF9353F794B7D.taxon	materials_examined	Type species: Omanella philippina Merino, 1936. Diagnosis: Tis genus closely resembles Halimunella and Paraphysipona in coloration and overall structure but differs in lacking wide black maculae along the apices of the anterior branches of forewing vein R and in having the aedeagus symmetrical with a pair of distal processes and paraphysis absent. Description: Coloration: Body olive green to dull yellow. Crown and dorsum of compound eyes bright orange with three small black anterior submarginal spots between eyes. Face with thin transverse orange band dorsally continuous with orange area of crown. Pronotum blue-green with antero-medial, medial, and posterolateral pairs of symmetrical orange markings. Mesonotum with black macula at each corner of basolateral triangle and pair of submedial orange stripes extended on to scutellum, scutellum apex orange. Forewing smoky hyaline with costal and commissural margins orange, orange spot at base of clavus, and apices of claval veins marked with black. Front and middle tibia orange. Body oblong, relatively broad, slightly depressed. Head slightly broader than pronotum; anterior margin relatively thick in profile, weakly bicarinate with few additional fine transverse striations. Ocelli small, situated in sulcus of foremargin, separated from adjacent eye by distance equal to own diameter. Crown flat, short, slightly depressed behind anterior margin, anterior margin broadly rounded in dorsal view, parallel to posterior margin. Face with lateral frontal sutures strongly divergent dorsad of antennal pits, extended to ocelli; clypeus relatively narrow, distinctly swollen dorsally, depressed ventrally; clypeal suture straight; anteclypeus with lateral margins concave, apex expanded; lora broader than basal width of anteclypeus; gena strongly emarginate below eyes; antenna longer than half-body length; antennal pits situated near upper corner of eyes, encroaching on to clypeus; antennal ledges moderately developed and oblique but not carinate. Pronotum well produced between eyes, hind margin slightly concave; lateral margin short, carinate; posterior two-thirds with irregular transverse striations. Mesonotum and scutellum together slightly broader than long; scutellar suture arcuate. Forewing with four apical cells and three subapical cells, inner subapical cell open; two anterior branches of R weakly reflexed; appendix broad. Fore femur with AM 1 near midheight of femur, AV with few stout setae in basal half; IC with series of several fine, close-set setae; fore tibia with dorsal setal arrangement of rows AD and PD: 1 + 4. Hind femur macrosetae 2 + 2 + 1. Male genitalia: Pygofer dorsal bridge moderately long; lobe broadly rounded with numerous macrosetae, posterior margin smooth, processes absent. Valve broadly triangular. Xth segment moderately long, well sclerotized dorsally and laterally, anterodorsal margin V-shaped in dorsal view. Subgenital plate with base broad and rounded laterally, apex atenuate, macrosetae absent, scatered long, fine setae present. Style apophysis stout with lateroposterior angle acutely produced (avicephaliform), finely denticulate, pre-apical lobe poorly developed. Connective small, Y-shaped with stem shorter than arms and broadened apically, arms thickened and weakly divergent. Paraphysis absent. Aedeagus with dorsal apodeme well developed, column-like, shallowly bifurcate apically; preatrium absent; shaf moderately long, curved dorsad with pair of slender retrorse processes apically; gonopore large, pre-apical on posterior surface. Etymology: Te genus name was formed by combining a prefix from the name of its home country (Philippines) with ‘ - pona ’, a suffix from the name of the type genus (Parabolopona) of the subtribe to which the new genus belongs. Gender: feminine. Distribution: Philippines. Remarks: Te type species of this monotypic genus grouped with Nakula in a sister-clade to the remaining members of Clade 2 in our phylogenetic analysis of combined morphological and molecular data (Fig. 3). It also did not group with the other included species of Roxasella in the phylogenomic analysis of Cao et al. (2022). Philipona resembles Paraphysipona in coloration and in having three black anterior spots on the vertex but differs in having paraphysis between aedeagus and connective absent. It resembles Roxasella in the structure of the male genitalia but differs from the later in having head with anterior margin moderately thick and aedeagus without paired apical processes.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD4B24AFE7BFCC13F1C4E8D.taxon	description	(Fig. 23 A – O) (URN: LSID: ZOOBANK. ORG: ACT: 83 A 1272 A-A 793 - 45 FD-A 23 C- 3887499 E 66 AF)	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD4B24AFE7BFCC13F1C4E8D.taxon	distribution	Distribution: Philippines. Material examined: 1 ♂, Philippines, Bukidnon, Brgy Impalutao, CEDAR (Center for Ecological Development and Recreation), (8 ° 25.09 ' N, 125 ° 034 ' E), 780 m, 7 July 2010, coll. J. Jones (INHS); holotype, Philippines (NMNH). Remarks: Tis species was described from a holotype (‘ type’) and paratypes with the following data: ‘ Mindanao, Zamboanga (type, Baker collection, U. S. N. M. ’. Te holotype (male) and allotype in USNM were examined and figured by Zhang and Webb (1996).	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD4B24AFE7BFCC13F1C4E8D.taxon	distribution	Distribution: China (Guangxi, Yunnan); Myanmar, Tailand. Remarks: Tis monotypic genus can be distinguished from all other Roxasella group genera externally by its distinctive orange and dark brown markings, vertex short and in the male genitalia by the caudal spine on the pygofer and shape of the subgenital plate with a few short spine-like setae (Fig. 24 E).	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFFDB263FC3BF90C39F24DBC.taxon	description	(Fig. 7 A – P) (URN: LSID: ZOOBANK. ORG: ACT: 654 AD 5 A 2 - 6 D 36 - 4489 - 8 BB 7 - DC 5 F 225 A 9 BA 8) Diagnosis: Head and thorax with orange markings. Crown with narrow black band at anterior margin. Male pygofer with caudoventral margin smooth. Style apex not bilobed, lateroapical margin truncate. Aedeagus with one pair of apical processes and extended ventrolateral beyond midlength of shaf in lateral view, shaf apex emarginate in ventral view. Small sclerite present between connective and aedeagus. Fore tibia with dorsal setal arrangement of rows AD and PD 4 + 7; fore femur with IC with about 14 setae, AV with four stout setae. Description: Length (including tegmen): male 8.0 mm. Coloration: Body yellowish. Crown with narrow black band at anterior margin and broader orange band between anterior eye corners. Face stramineous dorsally, white ventrally, dorsal margin with narrow black band. Pronotum with anterior margin white, irregularly arcuate orange band in middle, area behind orange band greenish brown. Crown parabolic, anterior and posterior margins subparallel. Fore tibia with dorsal setal arrangement of rows AD and PD 4 + 7; fore femur with IC with about 14 setae, AV with four stout setae. Male genitalia: Style (Fig. 7 O) without short apical extension beyond lateral beak-like lateral lobe. Connective (Fig. 7 K) with stem approximately as long as arms. Aedeagus with shaft relatively long and slender cylindrical, in lateral view (Fig. 7 J) sinuate in basal half and thereafter slightly bent dorsad; with one pair of long apical processes extended ventrolateral to midlength of shaft in posterior view. Small rectangular sclerite (paraphysis) articulated between connective and aedeagus. Material examined: Holotype: ♂, Malaysia: Sarawak (Borneo), Lambir Hills Nat’l Park, S of Miri (04 º 11.53.9 ′ N, 114 º 02.31.4 ′ E), 12 – 23 October 2006, coll. Cryan and Urban (INHS). Etymology: Te specific epithet is a Latinized adjective derived from the Greek word acontos (spear) referring to the spear-like aedeagal shaf. Remarks: Tis species is similar to R. abrupta in form and coloration but differs in the shape of the style apex and aedeagal shaf and in having a small rectangular sclerite (paraphysis) articulated between connective and aedeagus. Both species were collected together.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFFDB264FED4FA0138194DB4.taxon	description	(Fig. 8 A – Q) (URN: LSID: ZOOBANK. ORG: ACT: 846 E 9 D 39 - D 1 C 9 - 4 C 7 F-B 872 - 8 B 04 D 2642698) Diagnosis: Head and thorax with orange markings, crown with narrow black band at anterior margin. Male pygofer with caudoventral margin smooth. Style apex not bilobed, lateroapical margin convex. Aedeagus with one pair of subapical processes not extending beyond midlength of shaf, shaf apex entire in ventral view. Small sclerite present between connective and aedeagus. Fore tibia with dorsal setal arrangement of rows AD and PD 4 + 7; fore femur IC with 13 setae, AV with six stout setae. Description: Length (including tegmen): male 8.0 mm, female 9.0 mm. Coloration: Body yellowish. Crown with narrow black band at anterior margin and broader orange band between anterior eye corners. Face stramineous with narrow black transverse band at dorsal margin. Pronotum with anterior margin white, irregularly arcuate orange band in middle, area behind orange band greenish brown. Crown parabolic, anterior and posterior margins subparallel. Fore tibia with dorsal setal arrangement of rows AD and PD 4 + 7; fore femur IC with 13 setae, AV with six stout setae. Male genitalia: Style (Fig. 8 N) without short apical extension beyond lateral beak-like lateral lobe. Connective (Fig. 8 P) with stem slightly shorter than arms. Aedeagus with shaf slightly laterally compressed, in lateral view (Fig. 8 M) evenly curved and gradually widened from base to midlength, then parallel sided, in posterior view (Fig. 8 O) parallel sided throughout length; with one pair of short slightly sinuate apical processes extended ventrolateral. Small sclerite (paraphysis) articulated between connective stem and base of aedeagus, narrow and shallowly V-shaped in ventral aspect. Female (non-type) genitalia: Abdominal sternite VII (Fig. 8 D) with posterior margin with small lobe on each side of mid-line and concave in middle. Material examined: Holotype: ♂, Malaysia, Sabah, FMU 10, Apin apin, Keningau (05 ° 44.31 ′ 07 N, 116 ° 45.15 ′ 72 E), 1217 m, 24 April – 3 May 2016, coll. Z. S. Song. Paratypes: 1 ♂, same data as holotype (NWAFU). Additional (non-type) material: 1 ♀ (tentatively associated to males, see generic Remarks), same data as types (NWAFU). Etymology: Te specific epithet, a Latinized adjective, refers to the atached (appenda) sclerite between the aedeagus and connective. Remarks: Tis species can be distinguished by its relatively robust aedeagal shaf in lateral view with processes relatively short and in having a sclerite (paraphysis) articulated between the connective stem and base of the aedeagus.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFFAB264FC35FA0B3ED34D5E.taxon	description	(Fig. 9 A – Q) (URN: LSID: ZOOBANK. ORG: ACT: 0 A 87781 E- 4 FE 9 - 45 A 6 - BDD 6 - 3300 D 778 E 0 CC) Diagnosis: Head with two pairs of dark spots on anterior margin of crown. Male pygofer with caudoventral margin smooth. Style apex not bilobed, lateroapical margin convex. Aedeagus with two pairs of apical processes, one pair extended ventrolateral to shaf midlength and curved toward shaf, second pair very long and thin, extended slightly distad then sharply curved basad and extending nearly to shaf base. Without sclerite between connective and aedeagus. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about 14 setae. Description: Length (including tegmen): male 7.5 mm, female 8.0 mm. Coloration: Body tan to yellow. Crown with two pairs of black markings at anterior margin, one submedial and one sublateral adjacent to ocelli, remainder of crown tan anteriorly fading to yellow posteriorly. Face stramineous with narrow black band at dorsal margin. Pronotum tan anteriorly and laterally, darker greenish posteromedially. Crown produced, parabolically rounded, longer medially than next to eyes. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about 14 setae. Male genitalia: Style (Fig. 9 O) without short apical extension beyond lateral beak-like lateral lobe. Connective (Fig. 9 O) with stem similar in length to arms. Aedeagus with shaf relatively slender cylindrical, in lateral view (Fig. 9 N); shaf arcuate with two pairs of elongate apical processes, one pair extended cephalad and curved toward shaf, second pair longer and thinner, directed caudad and extended to near base of shaf. Female (non-type) genitalia: Abdominal sternite VII (Fig. 9 E) medially produced. Material examined: Holotype: ♂, Malaysia, Sabah, FMU 10, Apin apin, Keningau (05 ° 44.31 ′ 07 N, 116 ° 45.15 ′ 72 E), 1217 m, 24 April – 3 May 2016, coll. Z. S. Song. Additional (non-type) material: 1 ♀ (tentatively associated to males, see generic Remarks), same data as holotype (NWAFU). Etymology: Te specific epithet, a Latinized adjective, refers to the curved (arcuate) aedeagal shaf. Remarks: Te female tentatively identified as this species has the abdominal sternite VII similar to R. dolichospina but can be distinguished from the later by the greater projection of the posterolateral corner of sternite VII and the absence of intensely concave arcuated sculpture of sternite VII.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFFAB279FEBEFAA33AD24FCD.taxon	description	(Fig. 10 A – J)	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFFAB279FEBEFAA33AD24FCD.taxon	diagnosis	Diagnosis: Crown with narrow black band at anterior margin with orange band between anterior eye corners. Style apex bilobed, lateroapical margin distinctly concave. Aedeagus with two pairs of apical processes, shaf relatively slender and evenly curved in lateral view, apex slightly bifid, gonopore apical. Description: Body yellowish; crown with narrow black band at anterior margin and broader orange band between anterior eye corners. Male genitalia: Style with short apical extension beyond lateral beak-like lateral lobe (bilobed). Connective with stem similar in length to arms. Aedeagus with shaf relatively long and slender cylindrical, in lateral view evenly curved dorsad with apex expanded, in posterior view gradually broadened distally with one pair of long apical processes extended ventrocaudally beyond midlength of shaf and second pair of more slender and shorter processes curved lateroventrad. Material examined: Lectotype male (designated by Zhang and Webb 1996): Sarawak, NHMUK 013589074 (NHM). Distribution: Malaysia (Sarawak). Remarks: Tis species was described from an unknown number of female specimens (syntypic) from Borneo collected by Wallace. Te lectotype bears a circular data label with ‘ SAR’ for Sarawak, on one side and 57 36 on the reverse. Te later numbers refer to an entry in the NHM register for 1857 36 stating ‘ Borneo’ from ‘ Stevens’, the later acquiring specimens from Wallace.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE7B278FEDBFA41388D4B5D.taxon	description	(Fig. 20 F)	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE7B278FEDBFA41388D4B5D.taxon	description	Description (from Merino 1936): Length (including tegmen): male 7.0 mm, female 7.25 mm. Vertex, with a narrow ochraceous band at extreme anterior margin, preceded by a fuscous band two and two-thirds times as long as distance between the eyes; ocelli at apex of vertex just above and close to eyes; eyes fuscous. Face ochraceous; base of frons rounded with fuscous band, a deep excavated part above cheek, between base of frons and eyes, where antennae are located, clypeus narrower at base. Pronotum brown, roughly and transversely striate at posterior two-thirds, scutellum triangular, slightly wider than long, anterior portion brown, posterior angle ochraceous. Tegmina semitransparent pale brown, long veins brown, crimson bands on commissural region, on claval suture region, and on costal and first apical cells; pectoral and abdominal regions ochraceous with a light brown tinge. Posterior legs regularly marked with brown. Material examined: No material examined. Distribution: Philippines. Remarks: Tis species was described from the holotype and allotype from ‘ Luzon, Laguna Province, Los Baños (holotype S. Gonzales; allotype, Baker collection, USNM’. Zhang and Webb (1996) examined and illustrated the female allotype but stated that the male holotype was lost. Its absence in the USNM collection is confirmed by James Zahniser (USNM) who provided the habitus image of the allotype given here. However, from the information provided with the original description, the holotype was presumably retained in the private collection of S. Gonzales, the whereabouts of which is unknown. Te species is similar to R. losbanosa in external appearance, but its identity will remain uncertain until the genitalia of a male can be studied.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE6B278FC27FCA23F464B16.taxon	description	(Fig. 11 A – R) (URN: LSID: ZOOBANK. ORG: ACT: FBDF 4 F 77 - EA 9 B- 4770 - 90 B 0 - E 577 B 90 C 4 FA 7) Diagnosis: Head and thorax with orange markings, crown with narrow black band at anterior margin. Male pygofer with row of small teeth on caudoventral margin and short distal spine. Style apex bilobed, pre-apical lobe indistinct. Aedeagus with two pairs of distal processes. One apical pair extends beyond a midlength shaf, second pair subapical close to shaf ventrally, gonopore apical. Fore tibia with dorsal setal arrangement of rows AD and PD 4 + 7. Fore femur IC with about 16 setae, AV with four stout setae. Description: Length (including tegmen): male 8.5 mm, female 9.0 – 9.5 mm. Coloration: Body yellowish. Crown with narrow black band at anterior margin, area posterad of band ochraceous except narrowly yellow posterior margin. Face tan dorsally and yellowish more ventrad, with narrow black band at dorsal margin. Pronotum with anterior margin broadly yellow, irregularly arcuate orange medial band, and greenish brown posteromedial area. Forewing smoky hyaline with fuscous markings at apex of clavus and along anterior branches of R. Crown parabolic, anterior and posterior margins subparallel. Fore tibia with dorsal setal arrangement of rows AD and PD 4 + 7. Fore femur IC with about 16 setae, AV with four stout setae. Male genitalia: Pygofer with caudodorsal margin narrowly rounded, with short apical spine extended posteromesad, posteroventral margin serrate. Style apophysis expanded, apex with large posteromedial lobe and beaklike lateral pre-apical projection, pre-apical lobe indistinct. Connective short, stem broad and slightly longer than arms. Aedeagus with dorsal apodeme long, T-shaped in posterior view, bent near midlength in lateral view; shaf in lateral view nearly straight, slightly widened distally, in posterior view parallel sided through most of length with slight pre-apical bulge, with two pairs of distal processes, one apical pair extended ventrolaterally beyond a midlength shaf, curved posterad in lateral view, second pair subapical close to shaf ventrally, gonopore apical. Female (non-type) genitalia: Female sternite VII posterior margin with two short medial processes and pair of lateral concavities. Material examined: Holotype: ♂, Malaysia, Sabah, FMU 10, Apin apin, Keningau (05 ° 44.31 ′ 07 N, 116 ° 45.15 ′ 72 E), 1217 m, 24 April – 3 May 2016, coll. Z. S. Song. Additional (non-type) material: 2 ♀ (tentatively associated to males, see generic Remarks), same data as holotype (NWAFU). Etymology: Tis specific epithet, a Latinized adjective, refers to the spine at the apex of the pygofer lobe. Remarks: Tis species closely resembles R. appendasclerita but differs in having the aedeagal processes longer, the style with a distinctive large posteromedial lobe, and the male pygofer lobe with a distal spine and serrate posteroventral margin.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE6B27DFED5FC6A3AF34BF2.taxon	description	(Fig. 12 A – S) (URN: LSID: ZOOBANK. ORG: ACT: BF 840 CBE- 3 D 96 - 44 BB-B 35 F- CB 9 FB 9 DDAF 14) Diagnosis: Crown with a narrow black band at anterior margin interrupted at middle. Connective slender and elongate. Pygofer with caudodorsal margin bluntly rounded. Style apophysis weakly expanded apically, apex rounded medially. Aedeagus with two pairs of distal processes and extended nearly to atrium. Fore tibia with dorsal setal arrangement of rows AD and PD 4 + 4; fore femur IC with about 12 setae, AV with six setae basally. Description: Length (including tegmen): male 7.2 mm, female 8.2 mm. Coloration: Body yellowish; crown with a narrow black band at anterior margin interrupted at middle, remainder of crown ochraceous anteriorly fading to pale yellow posteriorly. Face stramineous with narrow black band dorsally. Pronotum tan medially, yellowish laterally. Forewing pale hyaline with three small fuscous spots along commissural margin of clavus, one just distad of claval apex, two in middle of corium and two at apices of anterior branches of R. Crown produced, parabolically rounded, distinctly longer medially than next to eyes. Fore tibia with dorsal setal arrangement of rows AD and PD 4 + 4; fore femur IC with about 12 setae, AV with six setae basally. Malegenitalia: Pygoferwithcaudodorsalmarginbluntlyrounded. Style apophysis weakly expanded apically, apex rounded medially and acute laterally, pre-apical lobe absent. Connective relatively slender and elongate with stem much longer than arms. Aedeagus with dorsal apodeme constricted near base then evenly widened dorsad with slightly convex dorsal end in posterior view; shaf relatively short, moderately slender, evenly curved and slightly widened distad in lateral view, very slightly widened from base to apex in posterior view, with two pairs of long apical processes extended ventrolateral, one pair evenly divergent from shaf, other pair curved toward shaf in posterior view, one pair extended more anterad than other in lateral view; apex very slightly emarginate in posterior view, gonopore apical. Female genitalia: Female sternite VII posterior margin trilobed with medial process angulate. Material examined: Holotype: ♂, Malaysia, Sabah, FMU 10, Apin apin, Keningau (05 ° 44.31 ′ 07 N, 116 ° 45.15 ′ 72 E), 1217 m, 24 April – 3 May 2016, coll. Z. S. Song. Paratypes: 1 ♂, same data as holotype (NWAFU). Additional (non-type) material: 1 ♀ (tentatively associated to males, see generic Remarks), same data as holotype (NWAFU). Etymology: Te specific epithet, a Latinized adjective, refers to the elongate processes of the aedeagus. Remarks: Te tentatively associated female of this species has the abdominal sternite VII similar to R. arcuata (see Remarks under R. arcuata).	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE3B27DFC10FC4F3BF84FC1.taxon	description	(Fig. 13 A – M)	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE3B27DFC10FC4F3BF84FC1.taxon	diagnosis	Diagnosis: Head thin, with fuscous markings on vertex, pronotum, exposed mesonotum. Pygofer lobe with caudal and ventral margins smooth. Style apophysis irregularly rounded. Aedeagus with shaf relatively short and broad, with two pairs of short apical processes. Male genitalia. Pygofer lobe with many macrosetae, caudal and ventral margins smooth. Valve broad, triangular. Subgenital plate long, triangular. Style apophysis irregularly rounded posteromedially with acute lateral projection. Connective short, Y-shaped with stem equal to lateral arms. Aedeagus with shaf relatively short and broad, with two pairs of short apical processes, one curved anterodorsal, other extended ventrolateral; dorsal atrium developed caudally, gonopore large, apical on posterior surface. Material examined: 1 ♂, 1 ♀, Philippines, Luzon, Mt. Makiling, Baker (NHM, donated by USNM). Distribution: Philippines. Remarks. Tis species was described from the holotype and paratypes from the Philippines with the following data, holotype: ' Luzon, Laguna Prov., Los Banos, (type, Baker collection, U. S. N. M.) ', paratypes: ' Mindanao, Surigao, Lanao, Zamboanga’. It was noted that the paratypes were slightly darker. Te holotype was examined by Zhang and Webb (1996) and images and figures of the same are provided here.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE3B27DFEF7FF643E504E14.taxon	description	(Fig. 14 A – O) (URN: LSID: ZOOBANK. ORG: ACT: F 69 D 6768 - B 475 - 4631 - ACE 5 - 3 EC 6 E 1118 C 19) Diagnosis: Crown with narrow black band at anterior margin. Style apex not bilobed, with lateroapical margin convex. Pygofer with caudodorsal margin narrowly rounded. Aedeagus with two pairs of apical processes extended ventrolateral, one pair curved anterad and other curved posterad in lateral view. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about 14 setae; AV with four setae. Description: Length (including tegmen): male 7.0 mm, female 7.7 mm. Coloration: Body yellowish. Crown with narrow black band at anterior margin, area posterad of band ochraceous except narrow pale area along posterior margin. Pronotum yellow anteriorly, greenish brown posteriorly. Crown parabolic, anterior margin slightly more produced than posterior margin. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about 14 setae; AV with four setae. Male genitalia: Pygofer with caudodorsal margin narrowly rounded. Style apophysis weakly expanded apically, apex with inner margin obtusely angulate and outer angle acute, pre-apical lobe absent. Connective stem broad and slightly longer than arms. Aedeagus with dorsal apodeme abruptly constricted then broadened near apex in posterior view; shaf moderately long and narrow, evenly curved dorsad in lateral view, narrow and parallel sided in posterior view, with two pairs of apical processes extended ventrolateral, one pair curved anterad and other curved posterad in lateral view, both extended to near midlength of shaf, gonopore apical. Material examined: Holotype: ♂, Malaysia: Sarawak; Gunung Mulu National Park, near headquarters (04 º 04. 25 ′ N, 114 º 81.38 ′ E), 30 June – 28 July 2009, coll. S. Krishnankuty, mercury vapour lights (INHS). Etymology: Tis species takes its name from the Mulu National Park. Remarks: Tis species resembles R. biarcuata in external form, coloration, and male genitalia but lacks an orange arcuate marking on the pronotum and has the processes of the aedeagus similar to each other in size.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFE3B271FEE1F96838354BB1.taxon	description	(Fig. 15 A – N) (URN: LSID: ZOOBANK. ORG: ACT: 1 B 8 CA 5 C 4 - 74 E 5 - 4 C 90 - 9113 - C 9 E 829 BDBF 8 D) Diagnosis: Crown with two pairs of black spots at anterior margin. Style apex not bilobed, with lateroapical margin convex. Pygofer with caudodorsal margin acutely angulate. Aedeagus with two pairs of apical processes extended basolaterad, approximately equal in length and extended to midlength of shaf. Small sclerite articulated between connective and aedeagus. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about 14 setae, AV with six stout setae. Description: Length (including tegmen): male 7.3 mm. Coloration: Body greenish yellow. Crown with two pairs of black spots at anterior margin, one submedial and one sublateral next to ocelli, with pair of transverse submedial orange bars more posterad. Face stramineous with narrow black band at dorsal margin. Pronotum yellow anteriorly, greenish brown posteriorly. Forewing with three fuscous spots along commissural margin of clavus and crossveins of corium marked with fuscous. Crown parabolic, longer medially than next to eyes. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about 14 setae, AV with six stout setae. Male genitalia: Pygofer with caudodorsal margin acutely angulate. Style apophysis weakly expanded, apex with inner margin obtusely angulate and lateral margin acute, pre-apical lobe very weakly developed. Connective moderately short, stem shorter than arms. Aedeagus with dorsal apodeme widened dorsad and truncate in posterior view, shaf moderately wide and gradually curved dorsad in lateral view, narrow basally and gradually expanded distally in posterior view, with two pairs of apical processes extended basolaterad, approximately equal in length and extended to midlength of shaf, one pair curved strongly anterad in lateral view, second pair only slightly curved and posterad of first pair, with apex slightly emarginate; gonopore apical. Small sclerite (paraphysis) articulated between connective and base of aedeagus. Material examined: Holotype: ♂, Malaysia, Sarawak, Gunung Mulu National Park, near headquarters (04 º 04. 25 ′ N, 114 º 81.38 ′ E), 30 June – 28 July 2009, coll. S. Krishnankuty, mercury vapour lights (INHS). Etymology: Te specific epithet, a Latinized adjective, refers to the four black spots on the anterior margin of the crown. Remarks: Tis species is similar to R. arcuata in form and coloration but differs in having the two pairs of aedeagal processes similar in size, the connective much shorter and broader, and the crown broader and less strongly produced.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFEFB271FC32FC0D3E5D4BAC.taxon	description	(Fig. 16 A – Q) (URN: LSID: ZOOBANK. ORG: ACT: 69 DE 7 BF 9 - 901 A- 4169 - AA 40 - 5 B 0191 BE 1 F 80) Diagnosis: Head and thorax with orange band. Crown with thin black band. Style apex bilobed with lateral projection beaklike. Aedeagal shaf relatively long and slender with one pair of stout apical processes having some strigae and basal spiculate or scale-like microsculpture, gonopore apical. Description: Length (including tegmen): male 8.0 – 8.5 mm, female 9.0 mm. Coloration: Body yellowish. Head with thin black band either side of anterior margin and crown with broader ochraceous band between anterior margins of eyes. Face stramineous with narrow black band at dorsal margin. Pronotum yellow anteriorly with irregularly arcuate ochraceous medial band, greenish brown posteromedially. Exposed mesonotum without orange markings. Crown parabolic, anterior and posterior margins subparallel. Fore tibia with dorsal setal arrangement of rows AD and PD 4 + 7. Fore femur IC with about 14 setae, AV with five stout setae. Male genitalia: Pygofer with caudodorsal margin narrowly rounded. Style apophysis avicephaliform with posteromedial lobe subquadrate and lateral projection beaklike, pre-apical lobe absent. Connective short, stem as long as arms. Aedeagus with dorsal apodeme slightly broadened dorsally in posterior view, dorsal margin truncate; shaf relatively long and slender, in lateral view somewhat irregularly curved dorsad and nearly parallel sided, in posterior view somewhat narrower and slightly narrowed near midlength, with one pair of stout apical processes, extended ventrolateral to near shaf midlength, curved slightly posterad in lateral view, with strigae and basal spiculate or scale-like microsculpture; gonopore apical. Female genitalia: Sternite VII posterior margin with short medial brown process slightly concave in middle. Material examined: Holotype: ♂, Malaysia, Sabah, FMU 10, Apin apin, Keningau, (05 ° 44.31 ′ 07 N, 116 ° 45.15 ′ 72 E), 1217 m, 24 April – 3 May 2016, coll. Z. S. Song. Paratypes: 1 ♂, same data as holotype (NWAFU); 1 ♂, Malaysia, Sarawak, Gunung Mulu National Park, Mulu Camp 2.5, 1000 m, February, RGS Mulu Expedition, Site 14, J. D. Holloway (NHM). Additional (non-type) material: 1 ♀ (tentatively associated to males, see generic Remarks), same data as holotype (NWAFU). Etymology: Te species takes its name from the type locality (Sabah). Remarks: Tis species is very similar to R. abrupta in form and coloration but differs in having the style apex with the posteromedial lobe more produced and quadrate, and the aedeagal processes much shorter with strigae and basal spiculate or scale-like microsculpture.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFEFB272FEE4FC1138CF4B19.taxon	description	(Fig. 17 A – N) (URN: LSID: ZOOBANK. ORG: ACT: 1951 C 16 C- 29 E 2 - 473 F- 94 C 4 - 859 BE 3 A 6 FF 4 B) Diagnosis: Head with two narrow transverse black bands at anterior margin. Style apex not bilobed, with lateroapical margin convex. Aedeagal shaf relatively short and slightly broader at base, thereafer parallel sided to apex in lateral view, with one pair of short sinuate apical processes. Without additional sclerite between connective and aedeagus. Fore tibia with dorsal setal arrangement of rows AD and PD 1 + 5. Description: Length (including tegmen): male 7.8 mm. Coloration: Body sordid yellow. Head with two narrow transverse black bands at anterior margin. Vertex with orange band between anterior eye corners; eyes crimson either side of white transverse band in line with anterior margin of head. Head slightly wider than pronotum. Crown produced medially by approximately one and a half times length next to eyes. Fore tibia with dorsal setal arrangement of rows AD and PD 1 + 5. Male genitalia: Pygofer lobe with caudal margin acutely rounded. Style apophysis very short, robust with posterior lateral angle acutely produced, pre-apical lobe moderately well developed. Connective with stem similar in length to arms. Aedeagus with shaf relatively short, in lateral view evenly curved, very slightly broader at base, thereafer parallel sided to apex, in ventral view slightly tapered from base to apex, digitate, with pair of short sinuate apical processes narrow at base, thereafer broadened to midlength and tapered to acute apex, extended ventrolaterally, gonopore apical, small; basal apodeme slightly broadened from base to apex in ventral view, apical margin slightly curved. Without additional sclerite (paraphysis) between connective and aedeagus. Material examined: Holotype: ♂, Indonesia, Sulawesi Utara, Dumoga-Bone National Park, 230 m, 11 July 1985, Fog 13, Plot A (NHM). Paratypes: Indonesia, Sulawesi Utara, Dumoga-Bone National Park, 1 ♂, near Edward’s camp, 664 m, 27. iv. 1985, M. Horak; 1 ♂, ii. 1985; 1 ♂, Edwards camp, 664 m, J. H. Martin; 1 ♂, Toraut, near base camp, c. 200 m, vi. 1985, M. R. Wilson. All types and non-types (see below) collected on Royal Entomological Society of London Project Wallace, B. M. 1985: 10 (NHM). Additional (non-type) material: Indonesia, Sulawesi Utara, Dumoga-Bone National Park, 1 ♀, ii. 1985; 2 ♀♀, Edwards camp, 664 m, J. H. Martin, 12. iv. 1985; 2 ♀♀, Toraut, near base camp, ca 200 m, vi. 1985, M. R. Wilson. Etymology: Tis species takes its name from the type locality (Sulawesi). Remarks: Tis species is similar to R. acontata and R. appendasclerita in shape and coloration but differs from the former in having the aedeagal shaf shorter and broader and from the later in having the aedeagal processes situated apically on the shaf rather than subapically and lacking a small accessory sclerite (paraphysis) between the connective and aedeagus.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFECB272FC34FC603E9A4ADB.taxon	description	(Fig. 18 A – Q) (URN: LSID: ZOOBANK. ORG: ACT: F 312 D 0 B 1 - CABB- 4397 - 83 D 6 - 902881 CDA 95 F) Diagnosis: Head with orange markings. Crown with narrow black band at anterior margin. Male pygofer with caudodorsal margin narrowly rounded. Style apex broad and falcate. Connective short. Aedeagus with two pairs of long apical processes nearly straight distally. Without additional sclerite between connective and aedeagus. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about 12 setae, AV with five stout setae. Description: Length (including tegmen): male 8.0 mm. Coloration: Body dull stramineous. Crown with narrow black band at anterior margin and broader ochraceous band between anterior corners of eyes. Face dull stramineous dorsally, becoming paler ventrally, with narrow black band at dorsal margin. Pronotum mostly greenish brown. Exposed mesonotum stramineous except basolateral triangles greenish brown. Crown parabolic with anterior and posterior margins subparallel. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about 12 setae, AV with five stout setae. Male genitalia: Pygofer with caudodorsal margin narrowly rounded. Style apophysis slightly expanded, apex broad and falcate, without distinct posteromedial lobe, pre-apical lobe very weakly developed and obtuse. Connective short, arms slightly longer than stem; aedeagus with dorsal apodeme slightly broadened near apex in posterior view; shaf moderately long and narrow, in lateral view curved dorsad at base then relatively straight, broader at base and apex than near middle, in posterior view narrow and nearly parallel sided, with two pairs of long apical processes extended basolaterad to midlength of shaf, one pair extended slightly posterad, other pair extended slightly anterad of shaf; gonopore apical. Material examined: Holotype: ♂, Malaysia, Sabah, FMU 10, Apin apin, Keningau (05 ° 44.31 ′ 07 N, 116 ° 45.15 ′ 72 E), 1217 m, 24 April – 03 May 2016, coll. Z. S. Song (NWAFU). Etymology: Te specific epithet refers to the umbel-like apex of the aedeagus in ventral view. Remarks: Tis species closely resembles R. muluensis in form and coloration but differs in having the aedeagal shaf relatively straight in lateral view with the processes more slender, the stem of the connective shorter and narrower, and the crown with anterior and posterior margins parallel.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFECB277FEE1FD273AEC4A43.taxon	description	(Fig. 19 A – Q) (URN: LSID: ZOOBANK. ORG: ACT: 14155 EB 9 - 83 E 6 - 4 FDA- 8906 - 5 A 91133417 E 0) Diagnosis: Crown with narrow black band at anterior margin. Style apophysis falcate apically. Aedeagus with shaf moderately short, with two pair of short processes, distal processes uncinate laterally and second pair laterally on shaf subapically. Without additional sclerite between connective and aedeagus. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Connective moderately long. Fore femur IC with about nine setae, AV with five stout setae. Description: Length (including tegmen): male 7.0 mm, female 7.5 mm. Coloration: Body stramineous. Crown with narrow black band at anterior margin and broader ochraceous band between anterior corners of eyes. Pronotum stramineous anteriorly and laterally, greenish brown posteromedially. Crown parabolic, anterior and posterior margins subparallel. Fore tibia with dorsal setal arrangement of rows AD and PD 2 + 4. Fore femur IC with about nine setae, AV with five stout setae. Male genitalia: Pygofer with caudodorsal margin narrowly rounded. Style apophysis falcate apically, pre-apical lobe indistinct. Connective moderately long and slender, stem longer than arms. Aedeagus with dorsal apodeme nearly parallel and apically truncate in posterior view; shaf relatively short, evenly curved dorsad, in lateral view broad at base, narrowed distally and slightly expanded apically, in posterior view unevenly narrowed from base to apex, with two pair of distal and subapical moderately short processes, one pair extended posteroventrad, not diverging from shaf in posterior view, second pair laterally on shaf subapically, slightly diverging from shaf in posterior view, uncinate laterally; gonopore apical. Female genitalia: Sternite VII posterior margin with short medial brown process. Material examined: Holotype: ♂, Malaysia, Sabah, FMU 10, Apin apin, Keningau (05 ° 44.31 ′ 07 N, 116 ° 45.15 ′ 72 E), 1217 m, 24 April – 03 May 2016, coll. Z. S. Song (NWAFU). Paratypes: 7 ♂♂, same data as holotype (NWAFU, NHM). Additional material: 4 ♀♀ (tentatively associated to males, see Remarks), same data as types. Etymology: Te specific epithet, a Latinized adjective, refers to the uncinate (hook-like) aedeagal apex. Remarks: Tis species resembles R. umbella in form and coloration but differs in having two pairs of relatively short aedeagal processes.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD1B24FFDF2FDA93B474D2C.taxon	description	(Fig. 25 A – I)	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD1B24FFDF2FDA93B474D2C.taxon	materials_examined	Material examined: China: 1 ♂ (holotype, BMSYS), Kwangtung Prov., Lung-tall Shan, Alt. 6 m, Yiu Vill., above Tsokokwan, 7 June 1947, Gressit et Lam. 1 ♂ (paratype, BMSYS) Kwangtung Prov., between Tsokokwan and Fung-Wan, Kukong District, 200 – 400 m, 11 June 1947, coll. Gressit; 1 ♀ (paratype, NHM), Guangxi Prov., Lingchuan, Lingtian, 5 June 1984, coll. Wu Zhengliang and Lu Xiaolin; 1 ♂ (paratype, CAU), Fujian Prov., Dehua, Shuikou, 7 Nov. 1974, coll. Li Fasheng; 1 ♂, Hainan Prov., Jianfengling Mountain, May- 3 - 2018 (NWAFU). Distribution: China (Hunan, Fu jian, Guangdong, Hainan, Guangxi, Guizhou, Yunnan). Remarks: Tis species is very similar to R. vietnamensis in its external structure and coloration, but can be distinguished from the later by its aedeagus with dorsal apodeme moderately slender and straight laterally, and aedeagus with apex extended nearly half of the length of the dorsal apodeme (equal in length in later) and the base of shaf curved arcuately rather than angularly in lateral view.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD1B24FFDB2FA933B664ED6.taxon	description	(Fig. 26 A – I)	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD1B24FFDB2FA933B664ED6.taxon	materials_examined	Material examined: 2 ♂♂, Vietnam, Ninh Binh, Cuc Phuong National Park, 125 m, 4 July 2019, coll. Lu Lin, light trap (NWAFU). Distribution: Vietnam.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD1B24FFC5BF9553E004E43.taxon	materials_examined	Type species: Sabahanus ornamentus sp. nov. Description: Body stramineous or ochraceous with conspicuous orange markings on head and thorax. Forewing with longitudinal orange bands basally, apical cells marked with brown, veins with a few scatered dark brown spots. Fore and hind tibia with setal bases dark brown. Head shorter than pronotum with two anterior carinae. Crown angularly produced with middle length longer than next to eyes, slightly transversely sulcate subapically; coronal suture distinct; faintly longitudinally striate. Ocelli slightly larger than twice their own diameter from eyes. Face slightly shorter than long; antennae situated near upper corner of eye in facial view, longer than half-body length; antennal pit deep and slightly encroaching on to clypeus; lateral frontal sutures well developed, evenly curved from above antennal pits to ocelli; gena slightly incurved below eye. Pronotum shorter than mesonotum, with carina on lateral margin. Forewing slightly tapered distally to apex, with appendix well developed; four apical cells and three subapical cells, inner subapical cell open, claval veins indistinct. Hindwing with four apical cells and two transverse veins. Fore femur setal formulation: IC = 12; AM 1 close to middle. Fore tibia with PD: AD = 2: 4; hind femur setal formula: 2 + 2 + 1. Male genitalia: Anal tube long. Pygofer produced acutely to apex with some macrosetae on lateral lobe, posterior margin sinuated in profile. Valve pentagonal. Subgenital plate with lateral margin convex in basal half with three to four lateral macrosetae in close oblique row, apex long and digital. Style with pre-apical lobe rectangular, apical process moderately robust, beak-like. Connective short, articulated with aedeagus, stem approximately as long as arms. Aedeagus with dorsal apodeme well developed, apex bifurcate in dorsal view; shaf elongate cylindrical, gonopore subapical on dorsal surface; base of aedeagus with a pair of ventral processes. Distribution: Malaysia (Borneo). Remarks: Tis genus is externally similar to Roxasella - group and previously treated as an undescribed group list in ‘ unplaced’ (Supporting Information, Table S 1), but can be distinguished externally by its bright orange markings on the head, pronotum, and forewings. It differs from the other paraboloponine genera in the male genitalia by the short oblique row of three to four macrosetae on the subgenital plate (with their bases darkly pigmented) similar to Tenompoella Zhang and Webb but in this genus the aedeagus is disassociated from the connective. Etymology: Te genus is named for the collecting locality of its type species ‘ Sabah’.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
B448EA62FFD1B241FED9F9B839814EC7.taxon	description	(Fig. 27 A – N) (URN: LSID: ZOOBANK. ORG: ACT: 25864 CA 8 - 7437 - 4 DBA-BE 60 - E 05 EDD 7963 CB) Description: Body length: Male, 6.5 mm. Body stramineous or ochraceous. Crown with patch distally and a pair of rounded spots near hind margin, orange; coronal suture orange. Pronotum with a pair of longitudinal sinuate bands near lateral margin, oblique paired sinuate bands from anterior margin and an inverted T-shape mark medially to hind margin, orange. Mesonotum with basal triangles and medial area orange. Forewing with orange band on claval suture, base and commissural region, and with smoky markings on apical cells and some scatered dark brown spots on veins. Fore and hind tibia with setal pits dark brown. Male genitalia: Aedeagal shaf slightly sinuate with apex curved ventrad hook-like in profile; with a pair of divergent sinuate elongate processes arising from base of aedeagus, tapered to acute apex and slightly longer than shaf. Material examined: Holotype: ♂, Malaysia, Sabah, FMU 10, Apin apin, Keningau, (05 ° 44.3107 ' N, 116 ° 45.1572 ' E), 1217 m, 2016.4.24 – 5.3, Z. S. Song (NWAFU). Paratype: 1 male, Malaysia, Borneo, Sabah, Bukit Monkobo, 51 ′ 48 ′′ N, 116 ′ 58 ′′, 1200 m, 21 - viii- 1987, A. H. Kirk-Spriggs, Light trap sample, camp 1, stunted hill forest, Bukit Monkobo (NMW). Other material: 1 female, same data as holotype (see Remarks). Revision of the Oriental Roxasella group • 41 Distribution: Malaysia (Borneo). Etymology: Te scientific name ‘ ornamentus ’ refers to the ornamental colour patern of this species. Remarks: Although the coloration of this species is very distinctive, two other new species from Sabah and Brunei (NHM) are similar. Terefore, we cannot be sure the female studied here is conspecific. If it is the same species then sexual dimorphism occurs, with the male crown having an anterior orange patch and orange coronal suture which are absent in the female.	en	Lu, Lin, Dietrich, Christopher H., Webb, M. D., Zhang, Yalin (2025): Revision of the Oriental Roxasella leafopper group: phylogeny, morphological evolution, biogeography, and description of 14 new species. Zoological Journal of the Linnean Society 204 (3), DOI: 10.1093/zoolinnean/zlaf040, URL: https://doi.org/10.1093/zoolinnean/zlaf040
