identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
99B3C3437DA25C69AA3DF5C5ED1B6E8D.text	99B3C3437DA25C69AA3DF5C5ED1B6E8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liphistius buyphradi Sivayyapram & Warrit 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Liphistius buyphradi Sivayyapram &amp; Warrit sp. nov.</p>
            <p>Figs 1, 10, 11, 14</p>
            <p>Type material.</p>
            <p>Holotype: Thailand • 1♂; Saraburi, Mueang Saraburi District, Nong Pla Lai; alt. 90 m; 17 August 2017; N. Warrit, V. Sivayyapram, C. Kunsete, N. Chatthanabun, P. Traiyasut leg.; ARA-2017-139. Allotype: Thailand • 1♀; same data as for the holotype; ARA-2017-140. Paratype: Thailand • 1♀; same data as for the holotype; ARA-2017-138.</p>
            <p>Diagnosis.</p>
            <p> Liphistius buyphradi sp. nov. is similar to  L. kaengkhoi sp. nov.,  L. champakpheaw sp. nov.,  L. hintung sp. nov. and  L. suwat in its general appearance. The male of  L. buyphradi sp. nov. can be distinguished from those species, except for  L. champakpheaw sp. nov., by the male palp with flat paracymbium (Fig. 14; Schwendinger 1996: fig. 43A); and from  L. champakpheaw sp. nov. by the male palp with proximal edge of the tegulum moderately long, arched, and finely dentate (short, oblique in  L. champakpheaw sp. nov.; Fig. 13C). The female  L. buyphradi sp. nov. is difficult to distinguish from those of the  L. kaengkhoi sp. nov.,  L. champakpheaw sp. nov.,  L. hintung sp. nov. and  L. suwat Schwendinger, 1996. Molecular phylogeny and species delimitation using COI and multi-locus data support monophyly and species status of all new species described here (Sivayyapram et al. 2023). </p>
            <p>Description.</p>
            <p>Male (Holotype: ARA-2017-139; Fig. 10A, B). Coloration (in alcohol): carapace brown with black stripe along the margins; abdominal tergite olive green, except on the white posterior margins; membranous parts of the opisthosoma cream colored with black mottled spots; chelicerae olive green, paler on the proximal part; palp and legs olive green, without distinct annulation.</p>
            <p>Palp (Fig. 11A-D): tibial apophysis round, carrying one long slender and three tapering megaspines; paracymbium round but flat, dark patch with spicules isolated by a pale band; cumulus plane, bearing long black bristles; subtegulum without apophysis; contrategulum without apophysis, distal edge of contrategulum narrow with a depression leading to the oblique conical dorsal apex; tegulum large, axe-blade shaped, possessing a long ridge on the surface, indistinctly separated from the contrategulum, proximal margin with arched, moderately dentate edge, distal margin oblique with moderately apophysis; pigmental bridge between tegulum and contrategulum distinct; paraembolic plate round, as long as wide, project into scale like-plate; embolus proper: sclerotized part with two longitudinal ridges reaching to the tip.</p>
            <p>Measurements: Total length 15.60; carapace 7.80 long, 7.28 wide; opisthosoma 8.19 long, 7.02 wide; ocular tubercle 1.04 long, 1.04 wide; palpal coxa 2.10 long, 1.17 wide; labium 0.72 long, 1.44 wide; sternum 3.92 long, 2.16 wide (1.02 on ventral surface); palp 12.48 long (4.14 + 2.34 + 3.90 + - + 2.10); leg I 20.88 long (6.06 + 3.00 + 4.26 + 5.22 + 2.34); leg II 22.62 long (6.24 + 3.12 + 5.16 + 7.14 + 2.64); leg III 24.66 long (6.60 + 3.12 + 5.16 + 7.14 + 2.64); leg IV 31.44 long (8.22 + 3.30 + 6.54 + 9.66 + 3.72).</p>
            <p>Female (Allotype: ARA-2017-140; Fig. 10C, D). Coloration (in alcohol): carapace orange with black marking behind the ocular tubercle and on the peripheral area of the thoracic region; abdominal tergites black, except on the white posterior margins; membranous part of the opisthosoma cream colored with black mottled spots; chelicerae orange, paler at the proximal part; palp and leg femora to metatarsi orange, distal part of metatarsi I-III with black mottled marking, tarsi I-III black, tarsi IV orange with black annulations on the proximal and distal parts.</p>
            <p>Vulva (Fig. 12E, F): vulva plate almost round; genital atrium with lateral hairs; posterior stalk narrow, V-shaped; pore plate quadrangular slightly wider than long; lateral margins thickened, projected into a lip, bearing moderate anterolateral lobes; anterior margin arched, thickened, not projected into a lip; receptacular cluster racemose; central dorsal opening wide.</p>
            <p>Measurements: Total length 24.70; carapace 10.66 long, 8.97 wide; opisthosoma 12.35 long, 8.84 wide; ocular tubercle 1.04 long, 1.04 wide; palpal coxa 3.24 long, 2.88 wide; labium 1.16 long, 2.48 wide; sternum 5.36 long, 2.80 wide (1.76 on ventral surface); palp 16.90 long (5.80 + 3.40 + 3.90 + - + 3.80); leg I 20.20 long (6.90 + 3.90 + 4.00 + 3.50 + 1.90); leg II 20.40 long (6.60 + 3.80 + 4.10 + 4.10 + 1.80); leg III 22.30 long (6.80 + 4.00 + 4.10 + 5.00 + 2.40); leg IV 30.80 long (8.40 + 4.20 + 6.10 + 8.20 + 3.90).</p>
            <p>Etymology.</p>
            <p>The specific epithet buyphradi is dedicated to Mr. Phuri Buyphrad for providing information on the type locality of the new species.</p>
            <p>Distribution.</p>
            <p>Known only from the type locality.</p>
            <p>Comment.</p>
            <p> This new species name was mentioned as  Liphistius sp. SL in Sivayyapram et al. (2023). </p>
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	https://treatment.plazi.org/id/99B3C3437DA25C69AA3DF5C5ED1B6E8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sivayyapram, Varat;Kunsete, Chawakorn;Xu, Xin;Smith, Deborah R.;Traiyasut, Prapun;Deowanish, Sureerat;Aung, Mu Mu;Ono, Hirotsugu;Li, Daiqin;Warrit, Natapot	Sivayyapram, Varat, Kunsete, Chawakorn, Xu, Xin, Smith, Deborah R., Traiyasut, Prapun, Deowanish, Sureerat, Aung, Mu Mu, Ono, Hirotsugu, Li, Daiqin, Warrit, Natapot (2024): Seven new species of the segmented spider genus Liphistius (Mesothelae, Liphistiidae) in Thailand and Myanmar. ZooKeys 1189: 201-229, DOI: http://dx.doi.org/10.3897/zookeys.1189.115850, URL: http://dx.doi.org/10.3897/zookeys.1189.115850
EF63018D21715F2781E25C40AF10B29D.text	EF63018D21715F2781E25C40AF10B29D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liphistius champakpheaw Sivayyapram & Warrit 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Liphistius champakpheaw Sivayyapram &amp; Warrit sp. nov.</p>
            <p>Figs 1, 12, 13, 14</p>
            <p>Type material.</p>
            <p>Holotype: Thailand • 1♂; Saraburi, Kaeng Khoi District, Cham Phak Phaeo; alt. 82 m; 19 August 2017; N. Warrit, V. Sivayyapram, C. Kunsete, N. Chatthanabun, P. Traiyasut leg.; ARA-2017-146. Allotype: Thailand • 1♀; same data as for the holotype; ARA-2017-146a.</p>
            <p>Diagnosis.</p>
            <p> Liphistius champakpheaw sp. nov. is similar to  L. kaengkhoi sp. nov.,  L. hintung sp. nov.,  L. buyphradi sp. nov., and  L. suwat in its general appearance.  Liphistius champakpheaw sp. nov. is closely similar to  L. buyphradi sp. nov. but can be distinguished by the male palp with tegulum that is short, obliqued, with dentate proximal edge (moderately long, arched in  L. buyphradi sp. nov., Fig. 11C). The female  L. champakpheaw sp. nov. is difficult to distinguish from those of the  L. kaengkhoi sp. nov.,  L. hintung sp. nov.,  L. buyphradi sp. nov., and  L. suwat Schwendinger, 1996. Molecular phylogeny and species delimitation using COI and multi-locus data support monophyly and species status of all new species described here (Sivayyapram et al. 2023). </p>
            <p>Description.</p>
            <p>Male (Holotype: ARA-2017-146; Fig. 12A, B). Coloration (in alcohol): carapace brown, slightly paler in the central area, without any distinct markings; abdominal tergites black, except for cream colored spots on the posterior margin; membranous part of the opisthosoma cream colored with black mottled spots; chelicerae olive green, paler at the proximal part; palp and legs uniformly brown.</p>
            <p>Palp (Fig. 13A-D): tibial apophysis wide distally, carrying four tapering megaspines; paracymbium round but flat, dark patch with spicules isolated by a pale band; cumulus plain, bearing several long bristles; subtegulum without apophysis; contrategulum without apophysis, distal edge of the contrategulum long with slight concavity leading to the truncate apex; tegulum moderate, proximal margins with short, tooth-like coarsely dentate edge; distal margin oblique, with large apophysis; paraembolic plate projected into scale-like plate, large, basally wide, running to blunt distal margin; embolus proper: sclerotized part with two longitudinal ridges reaching to the tip.</p>
            <p>Measurements: Total length 14.82; carapace 7.54 long, 7.02 wide; opisthosoma 7.41 long, 5.46 wide; ocular tubercle 1.17 long, 1.43 wide; palpal coxa 2.10 long, 1.17 wide; labium 0.78 long, 1.30 wide; sternum 3.90 long, 3.64 wide (1.17 on ventral surface); palp 12.22 long (4.03 + 1.56 + 4.42 + - + 2.21); leg I 23.53 long (6.89 + 2.86 + 5.07 + 6.24 + 2.47); leg II 24.83 long (7.02 + 2.99 + 5.07 + 6.76 + 2.99); leg III 28.34 long (7.28 + 3.38 + 5.72 + 8.32 + 3.64); leg IV 34.84 long (8.84 + 3.51 + 7.15 + 10.92 + 4.42).</p>
            <p>Female (Allotype: ARA-2017-146a; Fig. 12C, D). Coloration (in alcohol): carapace orange with black markings on the coxal elevations and the carapace margins; abdominal tergites with black markings except for the cream color on the posterior margin; membranous part of the opisthosoma cream colored with mottled black spots; chelicerae orange; palp and legs orange with black annulation on the proximal and distal part of tarsi.</p>
            <p>Vulva (Fig. 13E, F): vulva plate triangular, as long as wide; genital atrium with a few hairs and wrinkle posterior margin; posterior stalk U-shape, short but wide; pore plate quadrangular, slightly longer than wide; lateral margins thickened and projected into lips, bearing moderate anterolateral lobe; anterior margin arched, less thickened but not projected into a lip; receptacular cluster racemose longer than wide; central dorsal opening longer than wide.</p>
            <p>Measurements: Total length 19.37; carapace 9.23 long, 7.41 wide; opisthosoma 9.88 long, 8.06 wide; ocular tubercle 1.17 long, 1.56 wide; palpal coxa 3.25 long, 2.08 wide; labium 1.17 long, 2.47 wide; sternum 5.07 long, 2.86 wide (1.69 on ventral surface); palp 15.73 long (5.59 + 3.12 + 3.77 + - + 3.25); leg I 18.72 long (6.24 + 3.51 + 3.77 + 3.38 + 1.82) leg II 18.59 long (5.72 + 3.38 + 3.77 + 3.90 + 1.82); leg III 20.15 long (6.11 + 3.38 + 3.90 + 4.42 + 2.34); leg IV 28.99 long (8.32 + 3.90 + 5.59 + 7.80 + 3.38).</p>
            <p>Etymology.</p>
            <p>The specific epithet champakpheaw refers to Cham Pak Pheaw subdistrict, the type locality of the new species in Saraburi, Thailand.</p>
            <p>Distribution.</p>
            <p>Known only from the type locality.</p>
            <p>Comment.</p>
            <p> This new species name was mentioned as  Liphistius sp. CPP in Sivayyapram et al. (2023). </p>
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	https://treatment.plazi.org/id/EF63018D21715F2781E25C40AF10B29D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sivayyapram, Varat;Kunsete, Chawakorn;Xu, Xin;Smith, Deborah R.;Traiyasut, Prapun;Deowanish, Sureerat;Aung, Mu Mu;Ono, Hirotsugu;Li, Daiqin;Warrit, Natapot	Sivayyapram, Varat, Kunsete, Chawakorn, Xu, Xin, Smith, Deborah R., Traiyasut, Prapun, Deowanish, Sureerat, Aung, Mu Mu, Ono, Hirotsugu, Li, Daiqin, Warrit, Natapot (2024): Seven new species of the segmented spider genus Liphistius (Mesothelae, Liphistiidae) in Thailand and Myanmar. ZooKeys 1189: 201-229, DOI: http://dx.doi.org/10.3897/zookeys.1189.115850, URL: http://dx.doi.org/10.3897/zookeys.1189.115850
3E9526E89913578AA05870431C379398.text	3E9526E89913578AA05870431C379398.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liphistius choosaki Sivayyapram & Warrit 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Liphistius choosaki Sivayyapram &amp; Warrit sp. nov.</p>
            <p>Figs 1, 4</p>
            <p>Type material.</p>
            <p>Holotype: Thailand • 1♀; Phrae, Wang Chin District, Mae Koeng; alt. 265 m; 5 October 2019; N. Warrit, V. Sivayyapram, C. Kunsete, N. Chatthanabun, P. Traiyasut leg.; ARA-2019-057. Paratype: Thailand • 1♀; same data as for the holotype; ARA-2019-056.</p>
            <p>Other materials.</p>
            <p> Thailand • 3 juveniles; same data as for the holotype; ARA-2019-059, ARA-2019-061, ARA-2019-062 . </p>
            <p>Diagnosis.</p>
            <p> Liphistius choosaki sp. nov. is similar to  L. dawei sp. nov.,  L. inthanon , and  L. yamasakii in its uniformly dark coloration. The female of  L. choosaki sp. nov. can be distinguished from those of  L. dawei ,  L. inthanon , and  L. yamasakii by the characters of vulva: pore plate distinctly wider than long with almost straight anterior margin; and by its larger body size. </p>
            <p>Description.</p>
            <p>Male. Unknown.</p>
            <p>Female (Holotype: ARA-2019-057; Fig. 4A). Coloration (in alcohol): carapace uniformly brown; abdominal tergites dark brown; membranous part of the opisthosoma cream color with thin mottled black marking; chelicerae bicolor, orange on proximal part and dark brown on distal part; palp and legs brown, without distinct annulations.</p>
            <p>Vulva (Fig. 4B, C): vulva plate hexagonal, genital atrium with folded lateral margins, carrying lateral hairs; posterior stalk very wide, M-shaped posterior margin; pore plate rectangular, wider than long, lateral margin convex, anterior margin almost straight, all margins thickened and projected into a lip; receptacular cluster racemose, longer than wide; central dorsal opening wide longer than wide.</p>
            <p>Measurements: Total length 36.40; carapace 18.33 long, 16.38 wide; opisthosoma 17.94 long, 15.34 wide; ocular tubercle 1.82 long, 1.95 wide; palpal coxa 5.46 long, 3.25 wide; labium 2.08 long, 4.29 wide; sternum 9.75 long, 3.90 wide (1.95 on ventral surface); palp 29.12 long (10.66 + 5.59 + 7.02 + - + 5.85); leg I 34.71 long (12.48 + 6.50 + 7.15 + 6.24 + 2.34); leg II 36.40 long (11.96 + 5.85 + 7.41 +7.54 +3.64); leg III 40.82 long (12.74 + 6.89 + 7.67 + 9.49 + 4.03); leg IV 53.69 long (15.21 + 7.41 + 10.79 + 14.95 + 5.33).</p>
            <p>Etymology.</p>
            <p>The specific epithet choosaki honors the late Mr. Choosak Pungrusmee, father to Mr. Sarawut Pungrusmee and dedicated philanthropist to the study of biodiversity in our research laboratory.</p>
            <p>Distribution.</p>
            <p>Known only from the type locality.</p>
            <p>Comment.</p>
            <p> This new species name was mentioned as  Liphistius sp. WKS in Sivayyapram et al. (2023). </p>
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	https://treatment.plazi.org/id/3E9526E89913578AA05870431C379398	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sivayyapram, Varat;Kunsete, Chawakorn;Xu, Xin;Smith, Deborah R.;Traiyasut, Prapun;Deowanish, Sureerat;Aung, Mu Mu;Ono, Hirotsugu;Li, Daiqin;Warrit, Natapot	Sivayyapram, Varat, Kunsete, Chawakorn, Xu, Xin, Smith, Deborah R., Traiyasut, Prapun, Deowanish, Sureerat, Aung, Mu Mu, Ono, Hirotsugu, Li, Daiqin, Warrit, Natapot (2024): Seven new species of the segmented spider genus Liphistius (Mesothelae, Liphistiidae) in Thailand and Myanmar. ZooKeys 1189: 201-229, DOI: http://dx.doi.org/10.3897/zookeys.1189.115850, URL: http://dx.doi.org/10.3897/zookeys.1189.115850
CD688CF05F6A539DA44F37C5CE6D9EAF.text	CD688CF05F6A539DA44F37C5CE6D9EAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liphistius dawei Sivayyapram & Warrit 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Liphistius dawei Sivayyapram &amp; Warrit sp. nov.</p>
            <p>Figs 1, 2, 3</p>
            <p>Type material.</p>
            <p>Holotype: Myanmar • 1♂; Dawei, Pa Kar Ri; alt. 20 m; 4 May 2018; N. Warrit, V. Sivayyapram, C. Kunsete, N. Chatthanabun, P. Traiyasut leg.; ARA-2018-143. Allotype: Myanmar • 1♀; same data as for the holotype; ARA-2018-138. Paratypes: Myanmar • 1♂ 5♀; same data as for the holotype; ARA-2018-136, ARA-2018-137, ARA-2018-139, ARA-2018-140, ARA-2018-144, ARA-2018-147.</p>
            <p>Other material.</p>
            <p> Myanmar • 3 juveniles; same data as for the holotype; ARA-2018-146, ARA-2018-148, ARA-2018-149 . </p>
            <p>Diagnosis.</p>
            <p> Liphistius dawei sp. nov. is similar to  L. inthanon Zhan &amp; Xu, 2022 and  L. yamasakii Ono, 1988 in large body size with uniformly dark color. The male of  L. dawei sp. nov. can be distinguished from those of  L. inthanon and  L. yamasakii by the palp: subtegulum with moderate apophysis, not enlarged at the tip; tegulum with finer dentate edge of proximal margin and more pronounced marginal apophysis (Fig. 3A-D; Ono 1988: fig. 8; Schwendinger 1990: fig. 18; Zhan et al. 2022: fig. 4A-G). The female of  L. dawei sp. nov. can be distinguished from those of  L. inthanon and  L. yamasakii by the vulva: posterior stalk axe-blade shaped, constricted at the base; pore plate with less projecting posterior corners of the lateral lips (Fig. 3E, F; Ono 1988: figs 6, 7; Schwendinger 1990: fig. 19; Zhan et al. 2022: fig. 4H-M). </p>
            <p>Description.</p>
            <p>Male (Holotype: ARA-2018-143; Fig. 2A). Coloration (in alcohol): carapace uniformly brown, with black stripe along the margins, bearing short black setae on cephalic region and coxal elevations; abdominal tergite brown, bearing short black setae, paler on the glabrous area; membranous part of the opisthosoma cream color with a smear of black pigment; chelicerae pale brown; palp and legs pale brown, without distinct annulations.</p>
            <p>Palp (Fig. 3A-D): tibial apophysis short, truncate, carrying four black tapering megaspines; cumulus distinctly elevated, bearing long black bristles; paracymbium long, narrow, dark patch with spicules partially isolated by a pale band; subtegulum with moderated apophysis, not enlarged at the tip; contrategulum without apophysis, distal edge of contrategulum arched, leading to conical apex; tegulum kidney-shaped, wider than long, proximal margin convex with finely dentate edge, distal margin slightly concave with round and pronounced apophysis; pigmental bridge between contrategulum and tegulum indistinct; paraembolic plate indistinct, not projecting into a scale-like plate, sclerotized part of the embolus with two longitudinal ridges reaching to the tip.</p>
            <p>Measurements: Total length 20.15; carapace 9.75 long, 9.10 wide; opisthosoma 9.36 long, 7.28 wide; ocular tubercle 1.30 long, 1.56 wide; palpal coxa 3.12 long, 1.82 wide; labium 1.04 long, 1.82 wide; sternum 4.55 long, 2.73 wide (1.43 on ventral surface); palp 16.90 long (5.59 + 3.38 + 5.33 + - + 2.60); leg I 28.21 long (8.58 + 4.16 + 5.85 + 7.02 + 2.60); leg II 30.42 long (8.97 + 4.03 + 6.50 + 7.93 + 2.93); leg III 32.04 long (8.58 + 4.16 + 6.50 + 7.93 + 2.93); leg IV 39.91 long (10.7 + 4.29 + 8.19 + 12.48 + 4.68).</p>
            <p>Female (Allotype: ARA-2018-138; Fig. 2B). Coloration (in alcohol): carapace uniformly brown, bearing short black setae on the cephalic region and coxal elevations; abdominal tergites brown, darker in the area with short black setae; membranous part of the opisthosoma cream colored with thin layer of smear black marking; chelicerae bicolor, orange on proximal part and brown on distal part; palp and legs brown, without distinct annulations.</p>
            <p>Vulva (Fig. 3E, F): vulva plate hexagonal, genital atrium with folded lateral margins, carrying lateral hairs; posterior stalk axe-blade shaped, constricted at the based, posterior margin convex; pore plate rectangular and wider than long, lateral margin thickened and projecting into a lips, more distinct on anterior portion, posterior corner slightly projecting, anterior margin thickened and projecting into a lip, slightly arched; receptacular cluster racemose, longer than wide; central dorsal opening wide longer than wide.</p>
            <p>Measurements: Total length 27.43; carapace 12.87 long, 12.09 wide; opisthosoma 14.04 long, 11.44 wide; ocular tubercle 1.69 long, 1.95 wide; palpal coxa 4.42 long, 2.47 wide; labium 1.82 long, 3.12 wide; sternum 6.24 long, 3.51 wide (2.08 on ventral surface); palp 22.23 long (8.06 + 4.16 + 5.07 + - + 4.94); leg I 28.47 long (9.75 + 4.94 + 5.98 + 5.46 + 2.34); leg II 29.51 long (9.62 + 5.07 + 5.98 + 5.72 + 3.12); leg III 30.42 long (8.97 + 5.33 + 5.85 + 7.28 + 2.99); leg IV 41.34 long (11.44 + 5.46 + 7.80 + 12.09 + 4.55).</p>
            <p>Etymology.</p>
            <p>The specific epithet dawei refers to the type locality of the new species in Dawei State, Myanmar.</p>
            <p>Distribution.</p>
            <p>Known only from the type locality.</p>
            <p>Comment.</p>
            <p> The new species was mentioned as  Liphistius sp. DW in Sivayyapram et al. (2023). </p>
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	https://treatment.plazi.org/id/CD688CF05F6A539DA44F37C5CE6D9EAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sivayyapram, Varat;Kunsete, Chawakorn;Xu, Xin;Smith, Deborah R.;Traiyasut, Prapun;Deowanish, Sureerat;Aung, Mu Mu;Ono, Hirotsugu;Li, Daiqin;Warrit, Natapot	Sivayyapram, Varat, Kunsete, Chawakorn, Xu, Xin, Smith, Deborah R., Traiyasut, Prapun, Deowanish, Sureerat, Aung, Mu Mu, Ono, Hirotsugu, Li, Daiqin, Warrit, Natapot (2024): Seven new species of the segmented spider genus Liphistius (Mesothelae, Liphistiidae) in Thailand and Myanmar. ZooKeys 1189: 201-229, DOI: http://dx.doi.org/10.3897/zookeys.1189.115850, URL: http://dx.doi.org/10.3897/zookeys.1189.115850
EA7D1033369F52DBBB3751E083F2535B.text	EA7D1033369F52DBBB3751E083F2535B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liphistius hintung Sivayyapram & Warrit 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Liphistius hintung Sivayyapram &amp; Warrit sp. nov.</p>
            <p>Figs 1, 8, 9, 14</p>
            <p>Type material.</p>
            <p>Holotype: Thailand • 1♂; Nakhon Nayok, Mueang Nakhon Nayok District, Hin Tung; alt. 90 m; 27 November 2018; D. Li, L. Yu V. Sivayyapram leg.; ARA-2018-299. Allotype: Thailand • 1♀; same data as for the holotype; ARA-2018-296. Paratype: Thailand • 1 juvenile; same data as for the holotype; ARA-2018-297.</p>
            <p>Diagnosis.</p>
            <p> Liphistius hintung sp. nov. is similar to  L. kaengkhoi sp. nov.,  L. buyphradi sp. nov.,  L. champakpheaw sp. nov., and  L. suwat in its general appearance.  Liphistius hintung sp. nov. is closely similar to  L. kaengkhoi sp. nov. but can be distinguished by the male palp: tegulum with coarser proximal dental edge and longer paraembolic plate (Figs 7A-D, 9A-D). The female  L. hintung sp. nov. is difficult to distinguish from those of the  L. kaengkhoi sp. nov.,  L. buyphradi sp. nov.,  L. champakpheaw sp. nov., and  L. suwat Schwendinger, 1996. Molecular phylogeny and species delimitation using COI and multi-locus data support monophyly and species status of all new species described here (Sivayyapram et al. 2023). </p>
            <p>Description.</p>
            <p>Male (Holotype: ARA-2018-299; Fig. 8A, B). Coloration (in alcohol): carapace pale brown, with black stripe along the lateral and posterior margins; abdominal tergites almost black, paler on the posterior ones; membranous part of the opisthosoma cream in color with black mottled spots; chelicerae olive green, paler at the proximal part; palp and legs uniformly brown.</p>
            <p>Palp (Fig. 9A-D): tibial apophysis pronounce, carrying one long slender and three tapering megaspines; paracymbium conical, dark patch with spicules isolated by a pale band; cumulus plain, bearing some bristles; subtegulum without apophysis; contrategulum without apophysis, distal edge of contrategulum long and thick, slightly concave leading to the blunt apex; tegulum large, indistinctly separated from the contrategulum, axe-blade shaped, proximal margin with moderate long, coarsely dentate edge, distal margin oblique with large apophysis; pigmental bridge between the tegulum and contrategulum distinct; paraembolic plate projected into scale-like plate, basally wide with long and pointed distal edge; embolus proper: sclerotized part with two longitudinal ridges running to the truncate apex.</p>
            <p>Measurements: Total length 12.09; carapace 6.76 long, 6.37 wide; opisthosoma 5.33 long, 3.64 wide; ocular tubercle 1.17 long, 1.17 wide; palpal coxa 2.21 long, 1.56 wide; labium 0.78 long, 1.82 wide; sternum 3.64 long, 1.95 wide (1.04 on ventral surface); palp 12.35 long (3.90 + 2.34 + 4.03 + - + 2.08); leg I 19.37 long (5.85 + 2.99 + 3.90 + 4.42 + 2.21); leg II 19.89 long (5.72 + 2.73 + 4.03 + 5.20 + 2.21); leg III 22.88 long (6.11 +3.12 + 4.55 + 6.37 + 2.73); leg IV 27.81 long (7.14 + 3.25 + 5.72 + 8.45 + 3.25).</p>
            <p>Female (Allotype: ARA-2018-296; Fig. 8C, D). Coloration (in alcohol): carapace orange, with black marking behind the ocular tubercle running to the fovea and coxal elevations and black stripe along the carapace margins, thicker on the anterior margin; abdominal tergites pale brown with large black marking; membranous part of the opisthosoma cream colored with mottled black spots; palp and legs orange with black annulations on the metatarsus and tarsus.</p>
            <p>Vulva (Fig. 9E, F): vulva plate hexagonal; genital atrium with a few hairs; posterior stalk trapezoidal, wider anteriorly; pore plate almost square, lateral margins thickened and projected into a lip, bearing indistinct anterolateral lobes; anterior margin convex, less thicken and not project into a lip; receptacular clusters racemose, longer than wide; central dorsal opening wide, longer than wide.</p>
            <p>Measurements: Total length 16.90; carapace 8.06 long, 7.02 wide; opisthosoma 8.32 long, 6.63 wide; ocular tubercle 1.17 long, 1.17 wide; palpal coxa 2.60 long, 1.43 wide; labium 1.04 long, 2.08 wide; sternum 3.77 long, 2.60 wide (1.56 on ventral surface); palp 13.91 long (4.94 + 2.47 + 3.38 + - + 3.12); leg I 16.90 long (5.59 + 3.12 + 3.38 + 3.12 + 1.69); leg II 17.81 long (5.72 + 2.99 + 3.77 + 3.51 + 1.82); leg III 18.72 long (5.59 + 3.12 + 3.77 + 4.16 + 2.08); leg IV 26.26 long (7.54 + 3.51 + 5.20 + 6.63 + 3.38).</p>
            <p>Etymology.</p>
            <p>The specific epithet hintung refers to Hintung District, the type locality of the new species in Nakhon Nayok, Thailand.</p>
            <p>Distribution.</p>
            <p>Known only from the type locality.</p>
            <p>Comment.</p>
            <p> This new species name was mentioned as  Liphistius sp. WTK in Sivayyapram et al. (2023) </p>
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	https://treatment.plazi.org/id/EA7D1033369F52DBBB3751E083F2535B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sivayyapram, Varat;Kunsete, Chawakorn;Xu, Xin;Smith, Deborah R.;Traiyasut, Prapun;Deowanish, Sureerat;Aung, Mu Mu;Ono, Hirotsugu;Li, Daiqin;Warrit, Natapot	Sivayyapram, Varat, Kunsete, Chawakorn, Xu, Xin, Smith, Deborah R., Traiyasut, Prapun, Deowanish, Sureerat, Aung, Mu Mu, Ono, Hirotsugu, Li, Daiqin, Warrit, Natapot (2024): Seven new species of the segmented spider genus Liphistius (Mesothelae, Liphistiidae) in Thailand and Myanmar. ZooKeys 1189: 201-229, DOI: http://dx.doi.org/10.3897/zookeys.1189.115850, URL: http://dx.doi.org/10.3897/zookeys.1189.115850
74A1866DA8BC5195A75412C5A658644C.text	74A1866DA8BC5195A75412C5A658644C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liphistius kaengkhoi Sivayyapram & Warrit 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Liphistius kaengkhoi Sivayyapram &amp; Warrit sp. nov.</p>
            <p>Figs 1, 6, 7, 14</p>
            <p>Type material.</p>
            <p>Holotype: Thailand • 1♂; Saraburi, Kaeng Khoi District, Cha Om; alt. 127 m; 14 October 2018; N. Warrit, V. Sivayyapram, C. Kunsete, N. Chatthanabun, P. Traiyasut leg.; ARA-2018-284. Allotype: Thailand • 1♀, same data as for the holotype; ARA-2018-286. Paratypes: Thailand • 2♂ 4♀; same data as for the holotype; ARA-2018-281, ARA-2018-282, ARA-2018-283, ARA-2018-285, ARA-2018-289, ARA-2018-291.</p>
            <p>Diagnosis.</p>
            <p> Liphistius kaengkhoi sp. nov. is similar to  L. buyphradi sp. nov.,  L. champakpheaw sp. nov.,  L. hintung sp. nov., and  L. suwat Schwendinger, 1996 in general appearance. The male of  L. kaengkhoi sp. nov. can be distinguished from that of  L. suwat by the paracymbium not bent outward; and the contrategulum without short blunt cone (Fig. 7; Schwendinger 1996: figs 43, 43A); from those of  L. buyphradi sp. nov. and  L. champakpheaw sp. nov. by the male palp with swollen paracymbium (flat in  L. buyphradi sp. nov.,  L. champakpheaw sp. nov.; Figs 11, 13);  L. kaengkhoi sp. nov. is very similar to  L. hintung sp. nov. but can be distinguished by the shorter and finer dentate edge on the proximal margin of the tegulum and shorter paraembolic plate (Figs 7A-D, 9A-D). The female  L. kaengkhoi sp. nov. is difficult to distinguish from those of the  L. buyphradi sp. nov.,  L. champakpheaw sp. nov.,  L. hintung sp. nov., and  L. suwat Schwendinger, 1996. Molecular phylogeny and species delimitation using COI and multi-locus data support monophyly and species status of all new species described here (Sivayyapram et al. 2023). </p>
            <p>Description.</p>
            <p>Male (Holotype: ARA-2018-284; Fig. 6A, B). Coloration (in alcohol): carapace brown with indistinct black mottling on cephalic region and coxal elevations; abdominal tergites black; membranous part of the opisthosoma cream colored with black mottling on antero-dorsal portion; chelicerae brown, paler on proximal portion; palp and legs brown without distinct annulation.</p>
            <p>Palp (Fig. 7A-D): tibial apophysis large, carrying one long slender and three tapering megaspines; paracymbium short, almost round, dark patch with spicules isolated by a pale band; cumulus plain, bearing thin black bristles; subtegulum without apophysis; contrategulum without apophysis, distal edge of contrategulum long, slightly invaginate leading to the conical apex; tegulum large, axe-blade shaped, indistinctly separated from the contrategulum by corrugated surface, proximal margin with a short moderate dentate edge, distal margin almost straight, with round apophysis; pigmental bridge between tegulum and contrategulum sigmoid in shape; paraembolic plate projecting to a scale-liked plate, basally wide, leading into short triangular distal margin; embolus proper: sclerotized part with two longitudinal ridges reaching to the truncated apex.</p>
            <p>Measurements: Total length 11.44; carapace 5.85 long, 5.72 wide; opisthosoma 5.82 long, 5.07 wide; ocular tubercle 0.97 long, 1.17 wide; palpal coxa 2.21 long, 1.30 wide; labium 0.65 long, 1.3 wide; sternum 3.99 long, 1.95 wide (0.91 on ventral surface); palp 12.09 long (3.90 + 2.21 + 3.90 + - + 2.08); leg I 18.72 long (5.72 + 2.73 + 3.77 + 4.42 + 2.08); leg II 19.50 long (5.72 + 2.60 + 3.90 + 4.81 + 2.47); leg III 20.80 long (5.46 + 2.60 + 4.29 + 5.85 + 2.60); leg IV 26.25 long (6.89 + 2.99 + 5.20 + 7.67 + 3.51).</p>
            <p>Female (Allotype: ARA-2018-286; Fig. 6C, D). Coloration (in alcohol): carapace orange with thick black band on the anterior margin and black mottled marking in the posterior portion of the cephalic region and the thoracic region; abdominal tergites cream colored with large mottled black markings; membranous part of the opisthosoma cream with black mottled marking; chelicerae orange; palp and legs: femur to tibia orange, tarsi and metatarsi of legs I-III black, metatarsi of leg IV orange with black annulation on the proximal and distal area.</p>
            <p>Vulva (Fig. 7E, F): vulva plate almost round, genital atrium without lateral hair; posterior stalk short, V-shaped; pore plate rectangular, lateral margins thickened and project into lips, anterior margin convex, less thicken, not project into a lip; receptacular clusters racemose, grape-like in shape; central dorsal opening wide.</p>
            <p>Measurements: Total length 16.2; carapace 6.89 long, 6.50 wide; opisthosoma 9.49 long, 9.67 wide; ocular tubercle 1.04 long, 1.04 wide; palpal coxa 2.60 long, 1.56 wide; labium 0.78 long, 1.95 wide; sternum 3.64 long, 2.21 wide (1.30 on ventral surface); palp 12.87 long (4.68 + 2.47 + 2.86 + - + 2.86); leg I 15.47 long (5.20+ 2.86 + 2.99 + 2.86 + 1.56); leg II 15.99 long (5.20 + 2.73 + 2.99 + 3.25 + 1.82); leg III 16.64 long (5.07 + 2.60 + 3.12 + 3.77 + 2.08); leg IV 23.81 long (7.02 + 3.38 + 4.18 + 6.24 + 2.99).</p>
            <p>Etymology.</p>
            <p>The specific epithet kaengkhoi refers to Kaeng Khoi District, the type locality of the new species in Saraburi, Thailand.</p>
            <p>Distribution.</p>
            <p>Known only from the type locality.</p>
            <p>Comment.</p>
            <p> This new species name was mentioned as  Liphistius sp. CK in Sivayyapram et al. (2023). </p>
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	https://treatment.plazi.org/id/74A1866DA8BC5195A75412C5A658644C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sivayyapram, Varat;Kunsete, Chawakorn;Xu, Xin;Smith, Deborah R.;Traiyasut, Prapun;Deowanish, Sureerat;Aung, Mu Mu;Ono, Hirotsugu;Li, Daiqin;Warrit, Natapot	Sivayyapram, Varat, Kunsete, Chawakorn, Xu, Xin, Smith, Deborah R., Traiyasut, Prapun, Deowanish, Sureerat, Aung, Mu Mu, Ono, Hirotsugu, Li, Daiqin, Warrit, Natapot (2024): Seven new species of the segmented spider genus Liphistius (Mesothelae, Liphistiidae) in Thailand and Myanmar. ZooKeys 1189: 201-229, DOI: http://dx.doi.org/10.3897/zookeys.1189.115850, URL: http://dx.doi.org/10.3897/zookeys.1189.115850
2238C5393A3651219CEBCA60115E69AC.text	2238C5393A3651219CEBCA60115E69AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liphistius lansak Sivayyapram & Warrit 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Liphistius lansak Sivayyapram &amp; Warrit sp. nov.</p>
            <p>Figs 1, 5</p>
            <p>Type material.</p>
            <p>Holotype: Thailand • 1♀; Uthai Thani, Lan Sak District, Rabam; alt. 200 m; 29 October 2020; V. Sivayyapram leg.; ARA-2021-067. Paratype: Thailand • 1♀, same data as for the holotype; ARA-2021-068.</p>
            <p>Diagnosis.</p>
            <p> Liphistius lansak sp. nov. is a small  Liphistius species recognized by the unique vulva: pore plate with receptacular cluster flanked by a pair of large vesicles. </p>
            <p>Description.</p>
            <p>Male. Unknown.</p>
            <p>Female (Holotype: ARA-2021-067; Fig. 5A, B). Coloration (in alcohol): carapace pale brown with black marking on the cephalic region and the margin of the thoracic region; abdominal tergites pale brown, with black marking on the anterior and lateral margins of each plate; membranous part of the opisthosoma cream colored with thin mottled black marking; chelicerae dark brown, with black marking, except on the proximal part; palp and legs pale brown, with black annulations on the proximal and distal part of each joint (Fig. 5B).</p>
            <p>Vulva (Fig. 5C, D): vulva plate hexagonal, wider than long, genital atrium without lateral hair; posterior stalk short but wide, posterior margin W-shaped; pore plate rectangular, wider than long, lateral margin not projected into a lip, anterior margin invaginated, thickened and projected into a lip; receptacular cluster racemose, as long as wide, flanked by a pair of large vesicles; central dorsal opening round and wide.</p>
            <p>Measurements: Total length 14.17; carapace 5.98 long, 5.46 wide; opisthosoma 8.19 long, 6.89 wide; ocular tubercle 0.91 long, 1.04 wide; palpal coxa 2.21 long, 1.17 wide; labium 0.78 long, 1.82 wide; sternum 3.12 long, 1.95 wide (1.30 on ventral surface); palp 10.79 long (3.77 + 1.95 + 2.47 + - + 2.60); leg I 14.04 long (4.55 + 2.08 + 2.86 + 2.99 + 1.56); leg II 15.08 long (4.81 + 2.21 + 2.99 + 3.38 + 1.56); leg III 17.16 long (4.68 + 2.47 + 3.25 + 4.03 + 2.73); leg IV 20.94 long (5.33 + 2.47 + 4.03 + 5.98 + 3.13).</p>
            <p>Etymology.</p>
            <p>The specific epithet lansak refers to Lan Sak District, the type locality of the new species in Uthai Thani, Thailand.</p>
            <p>Distribution.</p>
            <p>Known only from the type locality.</p>
            <p>Comment.</p>
            <p> This new species name was mentioned as  Liphistius sp. HKK in Sivayyapram et al. (2023). </p>
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	https://treatment.plazi.org/id/2238C5393A3651219CEBCA60115E69AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sivayyapram, Varat;Kunsete, Chawakorn;Xu, Xin;Smith, Deborah R.;Traiyasut, Prapun;Deowanish, Sureerat;Aung, Mu Mu;Ono, Hirotsugu;Li, Daiqin;Warrit, Natapot	Sivayyapram, Varat, Kunsete, Chawakorn, Xu, Xin, Smith, Deborah R., Traiyasut, Prapun, Deowanish, Sureerat, Aung, Mu Mu, Ono, Hirotsugu, Li, Daiqin, Warrit, Natapot (2024): Seven new species of the segmented spider genus Liphistius (Mesothelae, Liphistiidae) in Thailand and Myanmar. ZooKeys 1189: 201-229, DOI: http://dx.doi.org/10.3897/zookeys.1189.115850, URL: http://dx.doi.org/10.3897/zookeys.1189.115850
D6EF8D7312E45D2E954111419C077FE8.text	D6EF8D7312E45D2E954111419C077FE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liphistius Schiodte 1849	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  
Liphistius 
Schiodte
, 1849
</p>
            <p>Type species.</p>
            <p> Liphistius desultor Schiødte , 1849. </p>
            <p>Diagnosis.</p>
            <p> Liphistius can be distinguished from the heptathelid genera by the male palp possessing a tibial apophysis; the vulva modified into a pore plate or plate-like spermatheca; and the nest structure equipped with signal lines, unique silk lines radiating from the burrow entrance (Platnick and Sedgwick 1984; Xu et al. 2015a). </p>
            <p>Distribution.</p>
            <p>China (Yunnan Province), Indonesia (Sumatra), Laos, Peninsular Malaysia, Myanmar, and Thailand.</p>
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	https://treatment.plazi.org/id/D6EF8D7312E45D2E954111419C077FE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sivayyapram, Varat;Kunsete, Chawakorn;Xu, Xin;Smith, Deborah R.;Traiyasut, Prapun;Deowanish, Sureerat;Aung, Mu Mu;Ono, Hirotsugu;Li, Daiqin;Warrit, Natapot	Sivayyapram, Varat, Kunsete, Chawakorn, Xu, Xin, Smith, Deborah R., Traiyasut, Prapun, Deowanish, Sureerat, Aung, Mu Mu, Ono, Hirotsugu, Li, Daiqin, Warrit, Natapot (2024): Seven new species of the segmented spider genus Liphistius (Mesothelae, Liphistiidae) in Thailand and Myanmar. ZooKeys 1189: 201-229, DOI: http://dx.doi.org/10.3897/zookeys.1189.115850, URL: http://dx.doi.org/10.3897/zookeys.1189.115850
