taxonID	type	description	language	source
BFFADEC3AF1954FB9AAE340AE6A60DFD.taxon	description	Figs 3, 4 English name: Yangchun Horned Toad Chinese name: yáng chūn jiǎo chán (阳春角蟾)	en	Zhao, Yi-Yang, Lin, Shi-Shi, Li, Yuan-Hang, Lyu, Zhi-Tong, Zheng, Ke, Zeng, Zhao-Chi, Borzée, Amaël, Wang, Jian (2025): A new species of Chinese horned toad (Anura, Megophryidae, Boulenophrys) from western Guangdong Province, China, with recommendations for extinction risk assessments for several related congeners. Herpetozoa 38: 345-357, DOI: 10.3897/herpetozoa.38.e171400
BFFADEC3AF1954FB9AAE340AE6A60DFD.taxon	etymology	Etymology. The specific epithet yangchunensis is in reference to the type locality, Yangchun City.	en	Zhao, Yi-Yang, Lin, Shi-Shi, Li, Yuan-Hang, Lyu, Zhi-Tong, Zheng, Ke, Zeng, Zhao-Chi, Borzée, Amaël, Wang, Jian (2025): A new species of Chinese horned toad (Anura, Megophryidae, Boulenophrys) from western Guangdong Province, China, with recommendations for extinction risk assessments for several related congeners. Herpetozoa 38: 345-357, DOI: 10.3897/herpetozoa.38.e171400
BFFADEC3AF1954FB9AAE340AE6A60DFD.taxon	diagnosis	Diagnosis. (1) small size (SVL 27.6 – 28.7 mm in five adult males); (2) snout sharpened in dorsal view, canthus rostralis well developed, tongue not notched distally; (3) tympanum distinct; (4) vomerine ridges and vomerine teeth absent; (5) dorsal skin rough and highly granular, discontinuous X-shaped ridge on center of dorsum, discontinuous dorsolateral ridges present, sparse large tubercles on flanks, dorsal limbs with discontinuous transverse ridges and tubercles; (6) outer margin of upper eyelid with a small horn-like prominent tubercle, supratympanic fold distinct and narrow, curving posteroventrally to above arm; (7) two metacarpal tubercles distinct, inner one observably enlarged, relative finger lengths I <II <IV <III, distinct subarticular tubercle at base of each finger; (8) heels just meeting when hindlimbs folded, tibio-tarsal articulation reaching middle to anterior corner of eye; (9) toes without webbing and lateral fringes, inner metatarsal tubercle long ovoid, outer one absent, relative toe length I <II <V <III <IV; (10) dorsal surface yellowish-brown to dark brown, with indistinct irregular dark-brown patches and small orange dots; dark-brown triangular marking between eyes; dorsal limbs and digits light brown with dark-brown transverse bands; and (11) a single nuptial pad bearing dense nuptial spines on dorsal bases of fingers I and II in breeding adult males, subgular vocal sac present in males.	en	Zhao, Yi-Yang, Lin, Shi-Shi, Li, Yuan-Hang, Lyu, Zhi-Tong, Zheng, Ke, Zeng, Zhao-Chi, Borzée, Amaël, Wang, Jian (2025): A new species of Chinese horned toad (Anura, Megophryidae, Boulenophrys) from western Guangdong Province, China, with recommendations for extinction risk assessments for several related congeners. Herpetozoa 38: 345-357, DOI: 10.3897/herpetozoa.38.e171400
BFFADEC3AF1954FB9AAE340AE6A60DFD.taxon	description	Description of holotype. Adult male. Body size small, SVL 27.9 mm. Head length almost equal to head width, HDW / HDL 1.00; snout sharpened in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond margin of lower jaw; top of head flat; eyes moderate in size, ED 0.40 of HDL, pupil vertical, near diamond-shaped; nostril obliquely ovoid; canthus rostralis well developed; loreal region slightly oblique; internasal distance slightly larger than interorbital distance; tympanic region oblique, tympanum distinct and visible in dorsal view; tympanum moderate in size, margin clear, upper margin in contact with supratympanic fold, lower margin in contact with upper lip, TD / ED 0.50; large ovoid choanae at base of maxilla; vomerine ridge and vomerine teeth absent, maxillary teeth present; margin of tongue rounded, not notched distally; presence of single subgular vocal sac. Forearm length 0.20 of SVL, hand 0.30 of SVL; webbing absent between fingers, lateral fringes absent, relative finger length I <II <IV <III; tips of fingers slightly dilated, round; subarticular tubercles on base of fingers present, distinct; inner metacarpal tubercle observably enlarged, outer one slightly smaller; single nuptial pad bearing dense nuptial spines present on dorsal surface of first and second fingers, respectively. Hindlimbs moderate in length, tibio-tarsal articulation reaching forward to anterior corner of eye when hindlimb stretched along body; heels just meeting when flexed hindlimbs held at right angles to body axis; crus length 0.50 of SVL and foot length 0.70 of SVL; relative toe length I <II <V <III <IV; tips of toes round and slightly dilated; toes without lateral fringes and webbing; subarticular tubercles on base of toes present and distinct; inner metatarsal tubercle long ovoid and lacking outer metatarsal tubercle. Dorsal skin rough and highly granular; dense large tubercles on flanks; single horn-like prominent tubercle on edge of upper eyelid; obvious supratympanic fold curving posteroventrally from posterior corner of eye to level above insertion of arm; discontinuous X-shaped ridge on center of dorsum, discontinuous dorsolateral ridges present; dorsal limbs with discontinuous transverse ridges and tubercles; ventral surface smooth; small and distinct pectoral gland closer to axilla; single femoral gland positioned on posterior surface of thigh at midpoint between knee and cloaca.	en	Zhao, Yi-Yang, Lin, Shi-Shi, Li, Yuan-Hang, Lyu, Zhi-Tong, Zheng, Ke, Zeng, Zhao-Chi, Borzée, Amaël, Wang, Jian (2025): A new species of Chinese horned toad (Anura, Megophryidae, Boulenophrys) from western Guangdong Province, China, with recommendations for extinction risk assessments for several related congeners. Herpetozoa 38: 345-357, DOI: 10.3897/herpetozoa.38.e171400
BFFADEC3AF1954FB9AAE340AE6A60DFD.taxon	distribution	Distribution and natural history. Currently, Boulenophrys yangchunensis sp. nov. is only known from the type locality, Guangdong Yangchun Ehuangzhang Provincial Nature Reserve, western Guangdong, China. It inhabits flowing montane streams and the nearby forest floor and leaf litter. The habitat is surrounded by evergreen broad-leaved forest mixed with bamboo groves at elevations between 900 and 1,100 m (Fig. 5). Advertisement calls of males were heard from February to May. Males were found calling while perching on leaves that are 10 – 30 cm above the ground.	en	Zhao, Yi-Yang, Lin, Shi-Shi, Li, Yuan-Hang, Lyu, Zhi-Tong, Zheng, Ke, Zeng, Zhao-Chi, Borzée, Amaël, Wang, Jian (2025): A new species of Chinese horned toad (Anura, Megophryidae, Boulenophrys) from western Guangdong Province, China, with recommendations for extinction risk assessments for several related congeners. Herpetozoa 38: 345-357, DOI: 10.3897/herpetozoa.38.e171400
