identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
BC161708FFB9FFD4FF28F90A2498FCD1.text	BC161708FFB9FFD4FF28F90A2498FCD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vemanidae Lowry & Myers 2017	<div><p>Family VEMANIDAE Lowry &amp; Myers, 2017</p><p>Included genera. Vemana Barnard, 1964</p><p>Type species. Vemana compressa Barnard, 1964 (type by original designation)</p><p>Included species. Vemana contains seven species: Vemana compressa Barnard, 1964; V. cuspidata sp. nov.; V. geyserensis Ledoyer, 1986; V. hortonae sp. nov.; V. lemuresa Barnard, 1967; V. lizata Barnard, 1964; V. touzeti Ortiz &amp; Winfield, 2022 .</p><p>Diagnosis. (modified after Lowry &amp; Myers 2017)</p><p>[Antennae subequal] or antenna 1 shorter than antenna 2; callynophore well developed in both sexes; accessory flagellum short or minute. Antennae 1–2 calceoli absent (but present in terminal male). [Gland cone prominent] except in V. lizata . Mandibular incisor dentate; molar large, triturative. Maxilla 1 inner plate setose along medial margin; palps asymmetric. Maxilla 2 inner plate with strong, oblique setal row. Maxilliped palps well-developed. [Gnathopod 1 simple]. Gnathopod 2 subchelate [except in V. touzeti Ortiz &amp; Winfield, 2022 which lacks a palm], although described as possessing a transverse palm. Pereopods 1–4 coxae not acute ventrally. Coxa 4 subequal to or smaller than coxa 3. [Uropod 1 without apical spines]. Telson laminar, moderately cleft.</p><p>[divergencies are shown in square brackets]</p><p>Remarks. Maxilla 1 inner plate bears 13–22 setae on the medial margin in four of the five described species (unknown in V. lizata) but 42 in one of the new species described herein. Gnathopod 1 is here regarded as simple. Although the dactyl is shown as reflected to varying degrees in some illustrations, e.g. V. geyserensis, in no species is there any evidence for the existence of defining palmar spines. Gnathopod 2 has a strongly oblique palm in all species except V. touzeti and V. geyserensis . In V. touzeti, the palm is absent according to the illustration of Ortiz &amp; Winfield (2022, fig. 4, also p. 587), although described as being present and transverse. The palm of V. geyserensis is short and oblique, but almost transverse (Ledoyer, 1986, fig. 404, also p. 1022). Coxa 3 is deeper than coxa 4 to a greater or lesser degree in all species of Vemana except V. lizata . Based on the shape of the coxae of pereopods 3 and 4 as shown by Barnard (1964, fig. 31J, K), it is probable that the labelling has been transposed and that coxa 3 is deeper than coxa 4 in that species also. Coxa 4 lacks a posterior excavation in all species except V. touzeti and the new species, V. hortonae . Vemana compressa is the only species with a rounded posteroventral corner of epimeron 3.</p></div>	https://treatment.plazi.org/id/BC161708FFB9FFD4FF28F90A2498FCD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hendrycks, Ed A.;Thurston, Michael H.	Hendrycks, Ed A., Thurston, Michael H. (2025): Two new species of Vemana Barnard, 1964 (Crustacea, Amphipoda, Vemanidae) from abyssal depths in the eastern tropical Atlantic Ocean. Zootaxa 5653 (1): 97-115, DOI: 10.11646/zootaxa.5653.1.5, URL: https://doi.org/10.11646/zootaxa.5653.1.5
BC161708FFBAFFD4FF28FC36207DFB0C.text	BC161708FFBAFFD4FF28FC36207DFB0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vemana Barnard 1964	<div><p>Vemana Barnard, 1964</p><p>Diagnosis. As for family. Palps of maxillae 1 asymmetric, left maxilla 1 palp article 2 broad and spatulate, with stouter apical spines. Uropod 2, lateral margin of peduncle with stout spines (except in V. lemuresa Barnard, 1967).</p><p>Remarks. The outer ramus or both rami of uropod 2 bear terminal spines in most species. In V. geyserensis and V. touzeti the spines appear to be inset into the apices of the rami as shown for Tryphosella in Lowry &amp; Myers (2013, fig. 1F) and thus distinct from the apical spines which characterize the Senticaudata .</p></div>	https://treatment.plazi.org/id/BC161708FFBAFFD4FF28FC36207DFB0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hendrycks, Ed A.;Thurston, Michael H.	Hendrycks, Ed A., Thurston, Michael H. (2025): Two new species of Vemana Barnard, 1964 (Crustacea, Amphipoda, Vemanidae) from abyssal depths in the eastern tropical Atlantic Ocean. Zootaxa 5653 (1): 97-115, DOI: 10.11646/zootaxa.5653.1.5, URL: https://doi.org/10.11646/zootaxa.5653.1.5
BC161708FFBAFFDFFF28FB7825F1FD2F.text	BC161708FFBAFFDFFF28FB7825F1FD2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vemana cuspidata Hendrycks & Thurston 2025	<div><p>Vemana cuspidata sp. nov.</p><p>(Figs. 1–5)</p><p>Type material. Holotype: Mature female, length 10.0 mm, carcass plus 2 microscope slides (CMNC 2025-0858); DIVA-2 M63/2 Expedition, RV Meteor, Station #69 (Guinea Basin), 0.25m ² USNEL box corer, 17 March 2005, 0°26.7’ S 2°20.5’ W, 5060 m . Paratype: Male, posterior fragment (pereonite 5 to telson), total length of complete specimen ca 9.5 mm (NHMUK 2025.1102), Discovery Cruise 63, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-22.7&amp;materialsCitation.latitude=20.776667" title="Search Plazi for locations around (long -22.7/lat 20.776667)">Station 8524#1</a>, 28 June 1974, 20°45.5’ N 22°42.5’ W to 20°46.6’ N 22°42.0’ W, BN1.5/5C, 4412 m .</p><p>Etymology. The specific epithet, cuspidata, refers to the strong distoventral tooth on peduncle 2 of antenna 1.</p><p>Diagnosis. Lateral cephalic lobe narrowly rounded, about as long as rostrum; antenna 1 peduncle article 2 with distoventral tooth; coxa 1 produced anteroventrally, apically rounded, depth greater than length, anterior margin straight; coxa 4 lacking posterodistal lobe; gnathopod 1 carpus and propodus subequal; gnathopod 2 palm oblique; pereopod 7 posterior margin of basis convex; epimeron 3 strongly produced posteriorly with strong acute tooth at posteroventral corner; telson cleft about 20%, lobes not tapering.</p><p>Description. Holotype, mature female, 10 mm, CMNC 2025-0858.</p><p>Body (Fig.1) vaulted, pereonites broader than deep. Urosome (Figs.1, 4) urosomite 1 gently rounded posterodorsally, with small boss and lateral ridge; urosomite 2 very short; urosomite 3 shorter than urosomite 1, slightly convex, with lateral ridge.</p><p>Head (Fig. 1) eyes absent; rostrum short, apically rounded, weakly downcurved; lateral cephalic lobe, triangular, subacute. Antenna 1 (Fig. 1) longer than length of head and pereonites 1–3 combined, peduncle article 2 with distoventral tooth, peduncle article 3 very short; accessory flagellum slender, biarticulate, about half as long as callynophore; flagellum 9-articulate. Antenna 2 (Fig. 1) 1.3x length of antenna 1, peduncle article 5 slightly shorter than articles 3 and 4 combined; flagellum 13-articulate.</p><p>Upper lip (Fig. 2) rounded distally, smooth with fine marginal setules. Mandible (Fig. 2), incisor processes strongly toothed, 4-dentate; left lacinia mobilis 5-dentate, right stout, transversally truncate, multidentate; accessory spine row, left of about 8 pectinate or setose stout setae, right with 4 spines, the three adjacent to the lacinia broadened into blade-like structures; molar process broad, triturative; palp attached level with molar, article 1 short, article 2 length 4.6x width with 15 A2 setae, article 3 length 0.6x article 2 with 1 A3 seta, 15 D3 setae and 3 E3 setae. Lower lip (Fig. 2) distal mesial margin with dense fine setules overlying simple setae, mandibular lobes oval. Maxilla 1 (Fig. 2) inner plate subtriangular, tapering distally with 22 plumose setae along inner margin; outer plate with 9 bifurcate or multidentate stout spine-teeth in a 5/4 arrangement, palp article 2 broad with 7 stout nodular spines on mesiodistal margin of left palp; right palp slender with 5–6 slender spines on mesiodistal margin. Maxilla 2 (Fig. 2) inner plate slightly shorter than outer with parallel margins, distally narrowly rounded, with strong mediofacial row of 24 plumose setae; outer plate lateral margin convex, apically setose. Maxilliped (Fig. 2) inner plate broad, slightly exceeding distal end of palp article 1, with 3 nodular spines apically and 7 long plumose setae on medial margin; outer plate broad, not reaching end of palp article 2, setose apically, with 11 elongated spines distally on medial margin; palp 4-articulate, article 2 longest, stout, inner margins setose, article 3 broad, 0.6x length of article 2, setose distally, dactyl short, unguis prominent.</p><p>Gnathopod 1 (Fig. 3), coxa subquadrangular, length more than twice width, anteroventral corner produced, narrowly rounded; basis, with long setae at midpoint of anterior margin, posterior margin lacking setae; ischium and merus short, setose ventrally; carpus, posterior margin setose; propodus, length 1.1x carpus, posterior margin weakly sinuous, setose; dactylus 0.8x propodus. Gnathopod 2 (Fig. 3), coxa, length 1.5x width, anterior margin convex, posterior margin straight, small tooth at posterodistal corner; basis straight, anterior margin with small setules; ischium and merus short, setose ventrally; carpus, posterior margin setose; propodus subovate, length 0.9x carpus, palm very oblique, defined by three stout spines, ventral margin setose; dactylus slender, length 0.5x propodus. Pereopod 3 (Fig. 4) coxa slightly narrowing distally, length greater than width, anterior and ventral margins convex, posterior margin sinuous, anteroventral corner broadly rounded, posteroventral corner with incipient tooth; basis 1.5x length of coxa, with margins slightly setose; merus posterior margin with long setae; carpus and propodus together just longer than merus, posterior margins strongly setose; dactylus slightly shorter than propodus, with unguis. Pereopod 4 (Fig. 4), coxa slightly narrowing distally, length about 1.5x width, anterior and distal margins broadly rounded, posterior margin weakly concave; other articles as in pereopod 3. Pereopod 5 (Fig. 4), coxa width less than twice length, anterolobate; basis, posterior margin with small tooth at 20% length, posterodistal margin weakly concave; merus 0.6x basis; carpus just shorter than merus, 0.5x basis; propodus shorter than carpus; anterior margins of merus to propodus with long setae; dactylus slender, subequal to propodus length. Pereopod 6 (Fig. 4) coxa width about twice length, bilobed, lobes subequal; basis, posterior margin convex proximally, straight distally; distal articles as in pereopod 5. Pereopod 7 (Fig. 4) a little shorter than pereopod 6; coxa oval, width twice length; basis broad, width more than half length, posterior margin convex, with about 10 fine crenulations; merus 0.4x basis, merus and carpus subequal, slightly stouter than in pereopods 5–6; propodus a little shorter than carpus; anterior margins of merus to propodus with clusters of strong spines; dactylus slender, a little shorter than propodus.</p><p>Oostegites (Figs. 3–4) present on gnathopod 2 and pereopods 3–5, long and slender, largest on gnathopod 2 and pereopods 3–4, slightly smaller on pereopod 5, with long curved apical and subapical brood setae ranging in number from 7–8. Gills (Figs. 3–4) suboval on gnathopod 2 and pereopods 3–4, broadly triangular on pereopods 5–6 and small, heart shaped on pereopod 7.</p><p>Epimeron 1 (Fig. 1) broadly rounded. Epimeron 2 (Fig. 1) subquadrangular, anterodistal corner rounded, distal margin straight, posterodistal corner with small acute tooth, posterior margin weakly sinuous. Epimeron 3 (Fig. 1) anterior corner rounded, distal margin straight, posterodistal corner produced into strong acute tooth, posterior margin straight.</p><p>Uropod 1 missing. Uropod 2 (Figs. 1, 5) peduncle stout, subequal to inner ramus, with 8 lateral and 1 distomedial spines; rami lanceolate, outer shorter than inner, both with small inset apical spine; inner ramus as long as peduncle with 6 medial and 3 lateral spines; outer ramus 0.9x peduncle with 3 lateral spines. Uropod 3 (Figs. 1, 5) peduncle very short, about 0.5x rami, with 2 distolateral and distomedial spines; rami of equal length, much longer than peduncle, inner ramus with 6 plumose setae on medial margin, 1 lateral spine; outer ramus biarticulate, second article short, length 0.2x rami, with 2 lateral and 1 medial spine. Telson (Figs. 1, 4–5) subrectangular, length 1.2x breadth, margins with 1 proximolateral spine, cleft 19%, lobes broadly rounded, with 1 small distal spine; surface with prominent microstructure.</p><p>Distribution. Known from eastern Atlantic, Guinea Basin and off the northwest continental margin of Africa.</p><p>Remarks. Vemana cuspidata sp. nov. is separated from all other species in the genus by the form of the telson; subrectangular in shape, shallowly and widely cleft (less than 25%) and with broadly rounded, non-tapered lobes. The new species and V. touzeti Ortiz &amp; Winfield, 2022 both possess a distoventral tooth on antenna 1 peduncle article 2 but the latter is clearly distinguished by the prominent lateral cephalic lobe, gnathopod 2 lacking palm, strongly cusped posteroventral corner of coxa 4, dentate pereopod 7 basis and deeply cleft telson. Ortiz &amp; Winfield (2022, fig. 2) illustrate the distoventral tooth of antenna 1 peduncle article 2, but this distinctive character is not mentioned in their species description.</p><p>Also of note, Ortiz &amp; Winfield (2022, fig 2, also p. 587) reference a callynophore on antenna 1 article 1 flagellum lower margin. However, the callynophore encompasses the whole article, and is not just confined to the lower margin as stated. All Vemana species possess a callynophore in both sexes. In common with many other alicelloids, the callynophore is larger in males and the dorsal boss on urosomite 1 is a little more prominent. Otherwise, the sexes are very similar (see Barnard 1967, p. 190).</p><p>This collection of Vemana cuspidata sp. nov. is also the deepest published occurrence of the genus, extending the depth range to 5060 m.</p></div>	https://treatment.plazi.org/id/BC161708FFBAFFDFFF28FB7825F1FD2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hendrycks, Ed A.;Thurston, Michael H.	Hendrycks, Ed A., Thurston, Michael H. (2025): Two new species of Vemana Barnard, 1964 (Crustacea, Amphipoda, Vemanidae) from abyssal depths in the eastern tropical Atlantic Ocean. Zootaxa 5653 (1): 97-115, DOI: 10.11646/zootaxa.5653.1.5, URL: https://doi.org/10.11646/zootaxa.5653.1.5
BC161708FFB1FFDCFF28FD522542F85B.text	BC161708FFB1FFDCFF28FD522542F85B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vemana hortonae Hendrycks & Thurston 2025	<div><p>Vemana hortonae sp. nov.</p><p>(Figs. 6–9)</p><p>Type material. Holotype: (based on the least damaged specimen). Juvenile female, length 12.6 mm, carcass plus 13 microscope slides (NHMUK 2025.1103); Discovery Cruise 63, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-22.7&amp;materialsCitation.latitude=20.776667" title="Search Plazi for locations around (long -22.7/lat 20.776667)">Station 8524#1</a>, 28 June 1974, 20°45.5’ N 22°42.5’ W to 20°46.6’ N 22°42.0’ W, BN1.5/5C, 4412 m . Paratypes: 1 immature female, 15.6 mm and 1 juvenile female, 12.5 mm, (NHMUK 2025.1105); same data as above . 1 adult female, 16.3 mm, (NHMUK 2025.1104); Discovery Cruise 63, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-22.738333&amp;materialsCitation.latitude=20.748333" title="Search Plazi for locations around (long -22.738333/lat 20.748333)">Station 8524#6</a>, 28 June 1974, 20°44.3’ N 22°44.4’ W to 20°44.9’ N 22°44.3’ W, BN1.5/5C, 4414–4416 m .</p><p>Etymology. Named in honour for Tammy Horton, in recognition of her important contributions on the ecology and taxonomy of amphipod necrophages, particularly in the eastern North Atlantic Ocean. In addition, she has had a major input into the development of the World Register of Marine Species (WoRMS) having chaired the Steering Committee for five years and has entered and checked more amphipod records (currently greater than 22,717) than any other contributor.</p><p>Diagnosis. Lateral cephalic lobe rounded, about as long as rostrum; maxilla 2 outer plate strongly convex and broadened, wider than inner plate; antenna 1 peduncle article 2 lacking distoventral tooth; coxa 1 subrectangular, length and depth subequal, anteroventral angle lacking tooth, anterior margin weakly concave; coxa 4 with prominent rounded posterodistal lobe; gnathopod 1 propodus one-third longer than carpus; gnathopod 2 palm oblique; pereopod 7 posterior margin of basis constricted; epimeron 3 produced posterodistally with small rounded tooth; telson cleft 67%, lobes tapering, subtriangular.</p><p>Description. Holotype, juvenile female, 12.6 mm, NHMUK 2025.1103.</p><p>Body (Fig. 6) vaulted, pereonites broader than deep. Urosome (Fig. 6), urosomite 1 gently rounded posterodorsally, with small boss; urosomite 2 short; urosomite 3 shorter than urosomite 1.</p><p>Head (Fig. 6) eyes absent; rostrum short, acute, not down-curved; lateral cephalic lobe short, triangular, rounded apically. Antenna 1 (Fig. 6) shorter than length of head and pereonites 1–4 combined, peduncle article 2 lacking distoventral tooth, peduncle article 3 shorter than 2; accessory flagellum slender, biarticulate, about half as long as callynophore; flagellum 14-articulate. Antenna 2 (Fig. 6) length 1.2x antenna 1, peduncle article 5 slightly longer than articles 3 and 4 combined; flagellum 19-articulate.</p><p>Mandible (Fig. 7), incisor processes strongly toothed, 6-dentate; left lacinia mobilis 6-dentate, right 4-dentate; accessory spine row, left of about 17 pectinate or setose stout setae, right with 10 stout spines, the five adjacent to the lacinia broadened into blade-like structures; molar process broad, triturative; palp attached just distal to molar, article 1 short, article 2 length 3.5–3.8x width with 18 A2 setae in 2 groups, article 3 length 0.7x article 2 with 21 D3 setae and 2 E3 setae. Lower lip (Fig. 7) distal medial margin with dense fine setules overlying simple, bifurcate and trifurcate setae, mandibular lobes oval. Maxilla 1 (Fig. 7) inner plate tapering distally, strongly setose with 42 plumose setae along inner margin; outer plate with 11 bidentate or simple spine teeth in a 7/4 arrangement, palp article 2 broad with 9 stout spines on mesiodistal margin. Maxilla 2 (Fig. 7) inner plate slightly shorter than outer with parallel margins and strong mediofacial row of 59 plumose setae; outer plate unusually shaped, lateral margin strongly bulged, convex, apically setose, much wider than inner plate. Maxilliped (Fig. 7) inner plate narrow, nearly reaching one-third length of palp article 2, with 2 nodular spine teeth apically and 14 long setae on medial margin; outer plate broad, not reaching end of palp article 2, setose apically, 8 spine teeth distally on medial margin; palp 4-articulate, article 2 longest, stout, setose medially, article 3 broad, 0.7x length of article 2, setose distally, dactyl short, length 0.8x article 3, unguis prominent.</p><p>Gnathopod 1 (Fig. 8) coxa subquadrangular, length and width about equal, anteroventral corner weakly produced, narrowly rounded; basis, with long setae on anterior and posterior margins; ischium and merus short, posterior distal margins setose; carpus, posterior margin strongly setose; propodus, length 1.3x carpus, posterior margin straight, with 11 setae; dactylus long, subequal in length to propodus. Gnathopod 2 (Fig. 8) a little longer than gnathopod 1; coxa, length just greater than width, anterior and ventral margins convex, corners rounded; basis, with long setae on posterior margin; ischium and merus short, posterior distal margins setose; carpus longer than propodus, with 6 posterior marginal clusters of setae; propodus oval, length 0.9x carpus, palm oblique, finely pectinate, defined by 3 stout spines, posterior margin with 4 clusters of long setae; dactylus length 0.5x propodus. Pereopod 3 (Fig. 8) coxa length and width subequal, anterior and ventral margins convex, posterior margin concave, distal corners broadly rounded; merus, length 0.6x basis; propodus longer than carpus, length 0.8x merus; dactylus slender, length 0.9x propodus; merus, carpus and propodus posterior margins with long setae. Pereopod 4 (Fig. 8), coxa length less than width, anterior margin broadly rounded, distal margin straight, oblique, with rounded, strong posterodistal lobe; other articles as in pereopod 3. Pereopod 5 (Fig. 9), coxa width about twice length; basis smooth, length about twice breadth; merus 0.7x basis; carpus just shorter than merus, 0.6x basis; propodus shorter than carpus; anterior margins of merus to propodus setose; dactylus slender, 0.9x propodus. Pereopod 6 (Fig. 9) about 1.2x length of pereopod 5; coxa, width about twice length, bilobed, lobes subequal; basis, posterior margin with obtuse angle at 25% length, distally sinuous; merus 0.6x basis; carpus as long as merus; propodus shorter than carpus; anterior margins of merus to propodus setose; dactylus slender. Pereopod 7 (Fig. 9) slightly shorter than pereopod 6; coxa much wider than long, greatest length anteriorly; basis, posterior margin with obtuse angle at 60% length, distally concave; merus 0.4x basis; carpus 1.3x merus; propodus shorter than carpus; anterior margins of merus to propodus with slender spines; dactylus slender, 0.8x propodus.</p><p>Epimeron 1 (Fig. 6) subquadrate, posterodistal corner angled, posterior margin weakly convex. Epimeron 2 (Fig. 6) anterodistal corner rounded, distal margin convex, posterodistal corner with small, rounded tooth, posterior margin weakly convex. Epimeron 3 (Figs. 6, 9) anterior corner rounded, distal margin straight, posterodistal corner produced into a small subacute tooth, posterior margin straight.</p><p>Uropod 1 (Fig. 9) peduncle stout, with 1 mediodistal spine and 2 proximolateral slender setae; inner ramus 1.6x maximum length of peduncle with 4 medial and 3 lateral spines, tapering to slender apex; outer ramus smooth [broken], but much shorter than inner. Uropod 2 (Fig. 9) peduncle stout with 16 slender lateral and 2 distolateral spines; rami lanceolate, both with small inset apical spine; inner ramus 0.8x peduncle with 6 medial and 5 lateral spines; outer ramus with 2 medial and 8 lateral spines. Uropod 3 (Fig. 9) peduncle short, stout, with 3 distomedial, 5 lateral and 1 distolateral spine; [rami probably damaged and regenerated] inner ramus longer than peduncle, medial margin with 8 plumose setae and 4 spines, lateral margin with 2 spines; outer ramus a little longer than inner, biarticulate, second article short, length 0.2x rami, first article with 2 medial, 4 lateral and 2 distolateral spines. Telson (Fig. 9) length and breadth subequal, proximolateral margins with 1 small spine, cleft 67%, lobes tapering, each with a small apical spine.</p><p>Distribution. Known from eastern Atlantic, off the northwest continental margin of Africa.</p><p>Remarks. Vemana hortonae sp. nov. shares with V. lemuresa Barnard, 1967 the distally constricted basis of pereopod 7 but differs from that species in the strong posterodistal lobation of coxa 4. The only other species in the genus in which coxa 4 is produced is V. touzeti Ortiz &amp; Winfield, 2022 but in this species the projection is a strong posteriorly directed tooth, not a rounded lobe. The new species has a very unusual morphology of maxilla 2, in which the outer plate lateral margin is strongly convex and bowed outward resulting in a much-broadened plate. This character is unique among Vemana species. Vemana hortonae is the largest species recorded in the genus, attaining a length substantially greater than the 10 mm females of V. lemuresa and V. cuspidata sp. nov. herein. Oostegites in the adult female paratype are typical of other alicelloids; long, narrow and with relatively few apical and subapical setae.</p></div>	https://treatment.plazi.org/id/BC161708FFB1FFDCFF28FD522542F85B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hendrycks, Ed A.;Thurston, Michael H.	Hendrycks, Ed A., Thurston, Michael H. (2025): Two new species of Vemana Barnard, 1964 (Crustacea, Amphipoda, Vemanidae) from abyssal depths in the eastern tropical Atlantic Ocean. Zootaxa 5653 (1): 97-115, DOI: 10.11646/zootaxa.5653.1.5, URL: https://doi.org/10.11646/zootaxa.5653.1.5
BC161708FFB7FFD9FF28FF3E2005FD92.text	BC161708FFB7FFD9FF28FF3E2005FD92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vemana compressa J. L. Barnard 1964	<div><p>Vemana compressa Barnard, 1964</p><p>Vemana compressa Barnard, 1964: 39, fig. 30. (type species)</p><p>Diagnosis. Lateral cephalic lobe broadly rounded, about as long as rostrum; antenna 1 peduncle article 2 lacking distoventral tooth; coxa 1 subrectangular, length greater than depth, anteroventral angle lacking tooth, anterior margin concave; coxa 4 lacking posterodistal lobe; gnathopod 1 propodus one-third longer than carpus; gnathopod 2 palm oblique; pereopod 7, basis with posterior margin convex; epimeron 3 produced posteriorly, posterodistal corner rounded; telson cleft less than 50%, lobes tapering, convex laterally.</p><p>Locality. Caribbean Sea, 10°11’ N 78°30’ W, Station V-15-14, 9 November 1958, 1826 m.</p><p>Abundance. Known only from the 5.1 mm male holotype, AMNH No. 12342 .</p></div>	https://treatment.plazi.org/id/BC161708FFB7FFD9FF28FF3E2005FD92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hendrycks, Ed A.;Thurston, Michael H.	Hendrycks, Ed A., Thurston, Michael H. (2025): Two new species of Vemana Barnard, 1964 (Crustacea, Amphipoda, Vemanidae) from abyssal depths in the eastern tropical Atlantic Ocean. Zootaxa 5653 (1): 97-115, DOI: 10.11646/zootaxa.5653.1.5, URL: https://doi.org/10.11646/zootaxa.5653.1.5
BC161708FFB7FFD9FF28FDCF2687FBAC.text	BC161708FFB7FFD9FF28FDCF2687FBAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vemana geyserensis Ledoyer 1986	<div><p>Vemana geyserensis Ledoyer, 1986</p><p>Vemana geyserensis Ledoyer, 1986: 1022–1024, fig. 404.</p><p>Diagnosis. Lateral cephalic lobe narrowly rounded, about as long as rostrum; antenna 1 peduncle article 2 lacking distoventral tooth; coxa 1 strongly produced anteroventrally, apically rounded, length greater than depth, anterior margin concave; coxa 4 lacking posterodistal lobe; gnathopod 1 propodus and carpus subequal; gnathopod 2 palm almost transverse; pereopod 7, basis with posterior margin convex; epimeron 3 angular, weakly produced posterodistally into a small acute tooth; telson cleft about 50%, lobes tapering, subtriangular.</p><p>Locality. Western Indian Ocean, Mozambique Channel, NE of Banc du Geyser, Station DE 12, (Campagne Benthédi), 2500 m.</p><p>Abundance. Known from four specimens, including the 5 mm (sex?) holotype and an 8 mm ovigerous female (only epimeron 3 is drawn of female specimen).</p></div>	https://treatment.plazi.org/id/BC161708FFB7FFD9FF28FDCF2687FBAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hendrycks, Ed A.;Thurston, Michael H.	Hendrycks, Ed A., Thurston, Michael H. (2025): Two new species of Vemana Barnard, 1964 (Crustacea, Amphipoda, Vemanidae) from abyssal depths in the eastern tropical Atlantic Ocean. Zootaxa 5653 (1): 97-115, DOI: 10.11646/zootaxa.5653.1.5, URL: https://doi.org/10.11646/zootaxa.5653.1.5
BC161708FFB7FFD9FF28FBD420A0F99B.text	BC161708FFB7FFD9FF28FBD420A0F99B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vemana lemuresa J. L. Barnard 1967	<div><p>Vemana lemuresa Barnard, 1967</p><p>Vemana lemuresa Barnard, 1967: 188–190, fig. 92.</p><p>Diagnosis. Lateral cephalic lobe narrowly rounded, about as long as rostrum; antenna 1 peduncle article 2 lacking distoventral tooth; coxa 1 strongly produced anteroventrally, apically subacute, length greater than depth, anterior margin concave; coxa 4 lacking posterodistal lobe; gnathopod 1 propodus and carpus subequal; gnathopod 2 palm oblique; pereopod 7, basis with posterior margin constricted; epimeron 3 strongly produced posteriorly, with strong tooth posterodistally; telson cleft about 50%, lobes tapering, subtriangular.</p><p>Locality. East Pacific Ocean, off Baja California. Holotype: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.82111&amp;materialsCitation.latitude=27.62139" title="Search Plazi for locations around (long -115.82111/lat 27.62139)">Station</a> 7249, 27°36’25” N 115° 56’25” W, 4 January 1961, 3705–3745 m. Male: Station 7228, 27°37’17” N 115°49’16” W, 30 December 1960, 3718–4392 m. Abundance. Known from the 10 mm female holotype, AHF No. 6113 and an 8.8 mm male .</p></div>	https://treatment.plazi.org/id/BC161708FFB7FFD9FF28FBD420A0F99B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hendrycks, Ed A.;Thurston, Michael H.	Hendrycks, Ed A., Thurston, Michael H. (2025): Two new species of Vemana Barnard, 1964 (Crustacea, Amphipoda, Vemanidae) from abyssal depths in the eastern tropical Atlantic Ocean. Zootaxa 5653 (1): 97-115, DOI: 10.11646/zootaxa.5653.1.5, URL: https://doi.org/10.11646/zootaxa.5653.1.5
BC161708FFB7FFD9FF28F9E2201CF84E.text	BC161708FFB7FFD9FF28F9E2201CF84E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vemana lizata J. L. Barnard 1964	<div><p>Vemana lizata Barnard, 1964</p><p>Vemana lizata Barnard, 1964: 39–40, fig. 31.</p><p>Diagnosis. Lateral cephalic lobe narrowly rounded, about as long as rostrum; antenna 1 peduncle article 2 lacking distoventral tooth; coxa 1 produced anteroventrally, apically rounded, length greater than depth, anterior margin concave; coxa 4 lacking posterodistal lobe; gnathopod 1 propodus a little longer than carpus; gnathopod 2 palm oblique; pereopod 7, basis with posterior margin convex; epimeron 3 strongly produced posteriorly, with acute tooth posterodistally; telson cleft less than 50%, lobes tapering, convex laterally.</p><p>Locality. Caribbean Sea, 14°05’ N 75°25’ W, Station V-15-10, 7 November 1958, 4077 m.</p><p>Abundance. Known only from the 6 mm (sex?) holotype, AMNH No. 12343 .</p></div>	https://treatment.plazi.org/id/BC161708FFB7FFD9FF28F9E2201CF84E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hendrycks, Ed A.;Thurston, Michael H.	Hendrycks, Ed A., Thurston, Michael H. (2025): Two new species of Vemana Barnard, 1964 (Crustacea, Amphipoda, Vemanidae) from abyssal depths in the eastern tropical Atlantic Ocean. Zootaxa 5653 (1): 97-115, DOI: 10.11646/zootaxa.5653.1.5, URL: https://doi.org/10.11646/zootaxa.5653.1.5
BC161708FFA8FFC6FF28FF3E27C7FD06.text	BC161708FFA8FFC6FF28FF3E27C7FD06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vemana touzeti Ortiz & Winfield 2022	<div><p>Vemana touzeti Ortiz &amp; Winfield, 2022</p><p>Vemana touzeti Ortiz &amp; Winfield, 2022: 585–593, figs. 2–5.</p><p>Diagnosis. Lateral cephalic lobe subacute, much longer than short rostrum; antenna 1 peduncle article 2 with distoventral tooth; coxa 1 subrectangular, depth greater than length, with small tooth at anteroventral angle, anterior margin convex; coxa 4 with prominent acute, posterodistal lobe; gnathopod 1 propodus and carpus subequal; gnathopod 2 lacking palm [authors state that it has a transverse palm, although this condition is not apparent in the illustration]; pereopod 7 posterior margin convex, with 6 teeth, one more prominent than the others; epimeron 3 produced posteriorly, with small acute tooth distally; telson subovate, cleft more than 50% [authors state it is cleft 33%, but as shown in their figure 5D, it appears to be cleft from 41–70%], lobes tapering, subtriangular.</p><p>Locality. SW Gulf of Mexico, Bay of Campeche, cruise site CH-25, 20°15’57.59” N 94°05’34.80” W, 20 September 2010, 1492 m.</p><p>Abundance. Known only from the 7.5 mm male holotype, CNCR36469 .</p></div>	https://treatment.plazi.org/id/BC161708FFA8FFC6FF28FF3E27C7FD06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hendrycks, Ed A.;Thurston, Michael H.	Hendrycks, Ed A., Thurston, Michael H. (2025): Two new species of Vemana Barnard, 1964 (Crustacea, Amphipoda, Vemanidae) from abyssal depths in the eastern tropical Atlantic Ocean. Zootaxa 5653 (1): 97-115, DOI: 10.11646/zootaxa.5653.1.5, URL: https://doi.org/10.11646/zootaxa.5653.1.5
BC161708FFA8FFC7FF28F93721A9FD7E.text	BC161708FFA8FFC7FF28F93721A9FD7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vemana Barnard 1964	<div><p>Key to Vemana species</p><p>1. Epimeron plate 3, posterodistal corner rounded; lateral head lobe broadly rounded, not strongly produced forward................................................................................... V. compressa J.L. Barnard, 1964</p><p>- Epimeron plate 3, posterodistal corner pointed or produced into a tooth; lateral head lobe narrow and produced forward.... 2</p><p>2. Coxa 4, with posterodistal lobe.......................................................................... 3</p><p>- Coxa 4, lacking posterodistal lobe........................................................................ 4</p><p>3. Coxa 1, anteroventral corner rounded; coxa 4, posterodistal lobe blunt, broadly rounded; gnathopod 2, propodus subovate, length 2.4x width; pereopod 7, basis, posterior margin with distinct medial constriction at 60% length, distal half concave, narrower than proximal half, lacking teeth.................................................. V. hortonae sp. nov.</p><p>- Coxa 1, anteroventral corner pointed; coxa 4, posterodistal lobe acute, corner produced into a strong tooth; gnathopod 2, propodus narrowly rectangular, length 3.4x width; pereopod 7, basis broad, posterior margin with six strong teeth, lacking medial constriction........................................................... V. touzeti Ortiz &amp; Winfield, 2022</p><p>4. Gnathopod 1, propodus appearing subchelate, ventral margin sinuous; pereopod 7, basis slender, posterior margin with distinct medial constriction, distal half narrower than proximal half............................ V. lemuresa J.L. Barnard, 1967</p><p>- Gnathopod 1, propodus simple with nearly straight ventral margin; pereopod 7, basis broad, posterior margin not medially constricted........................................................................................... 5</p><p>5. Antenna 1, peduncular article 2 with strong distoventral tooth; coxa 1, anterior margin slightly concave, coxa slightly directed forward; telson broad distally, lobes broadly rounded, gap of cleft wide and shallow, ~ 20% of length... V. cuspidata sp. nov.</p><p>- Antenna 1, peduncular article 2 lacking distoventral tooth; coxa 1, anterior margin strongly concave, coxa distinctly bent forward; telson narrowed distally, lobes narrow, gap of cleft narrow and deep, at least 36% of length................... 6</p><p>6. Epimeron plate 3, posterodistal corner with prolonged strong tooth; gnathopod 1, propodus slightly longer than carpus, dactylus shorter than propodus; gnathopod 2, palm distinctly oblique; pereopod 7, basis broad, ~ 0.64x length... V. lizata J.L. Barnard, 1964</p><p>- Epimeron plate 3, posterodistal corner angular, very weakly produced; gnathopod 1, propodus shorter than carpus, dactylus longer than propodus; gnathopod 2, palm slightly oblique, almost transverse; pereopod 7, basis narrow, ~ 0.44x length................................................................................... V. geyserensis Ledoyer, 1986</p></div>	https://treatment.plazi.org/id/BC161708FFA8FFC7FF28F93721A9FD7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hendrycks, Ed A.;Thurston, Michael H.	Hendrycks, Ed A., Thurston, Michael H. (2025): Two new species of Vemana Barnard, 1964 (Crustacea, Amphipoda, Vemanidae) from abyssal depths in the eastern tropical Atlantic Ocean. Zootaxa 5653 (1): 97-115, DOI: 10.11646/zootaxa.5653.1.5, URL: https://doi.org/10.11646/zootaxa.5653.1.5
