taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
43EC867ECFDE545A83C94CB08DF980AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483273	https://doi.org/10.3897/dez.72.158689.figure3	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
43EC867ECFDE545A83C94CB08DF980AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483287	https://doi.org/10.3897/dez.72.158689.figure17	Figure 17. Homoeusa brevitarsis sp. nov. A, B. Antenna; A. Male; B. Female; C. Pronotum and elytra; D. Zoomed view of elytron; E. Meso- and metaventrite; F, G. Hind tibia and meta-tarsus; F. Male; G. Female; H, I. Posterior margin of 8 th sternite; H. Male; G. Female; J. Median lobe of aedeagus from lateral view; K. Paramere of aedeagus from lateral view; L. Spermatheca.	Figure 17. Homoeusa brevitarsis sp. nov. A, B. Antenna; A. Male; B. Female; C. Pronotum and elytra; D. Zoomed view of elytron; E. Meso- and metaventrite; F, G. Hind tibia and meta-tarsus; F. Male; G. Female; H, I. Posterior margin of 8 th sternite; H. Male; G. Female; J. Median lobe of aedeagus from lateral view; K. Paramere of aedeagus from lateral view; L. Spermatheca.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
43EC867ECFDE545A83C94CB08DF980AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483288	https://doi.org/10.3897/dez.72.158689.figure18	Figure 18. Homoeusa brevitarsis sp. nov., distribution map; black star – the type locality; white circles – collection data examined in this paper.	Figure 18. Homoeusa brevitarsis sp. nov., distribution map; black star – the type locality; white circles – collection data examined in this paper.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
0081CFB869745162881E9CA6856EE80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483273	https://doi.org/10.3897/dez.72.158689.figure3	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
0081CFB869745162881E9CA6856EE80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483285	https://doi.org/10.3897/dez.72.158689.figure15	Figure 15. Homoeusa gigantea sp. nov. A. Antenna; B. Pronotum and elytra; C. Zoomed view of elytron; D. Meso- and metaventrite; E, F. Posterior margin of 8 th sternite: E. Male; F. Female; G. Median lobe of aedeagus from lateral view; H. Paramere of aedeagus from lateral view; I. Spermatheca.	Figure 15. Homoeusa gigantea sp. nov. A. Antenna; B. Pronotum and elytra; C. Zoomed view of elytron; D. Meso- and metaventrite; E, F. Posterior margin of 8 th sternite: E. Male; F. Female; G. Median lobe of aedeagus from lateral view; H. Paramere of aedeagus from lateral view; I. Spermatheca.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
0081CFB869745162881E9CA6856EE80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483286	https://doi.org/10.3897/dez.72.158689.figure16	Figure 16. Homoeusa gigantea sp. nov. A. Distribution map; black star – the type locality; white circles – collection data examined in this paper; B, C. Living adult in a foraging trail of Lasius umbratus under a stone, Morioka-shi, Iwate-ken, Honshu (photographed by Naoya Nakaya); D. Snapped images from videos shot indoors, adults rubbing abdomen against a host ant worker; E. Clear image of a living adult from Nagano-ken.	Figure 16. Homoeusa gigantea sp. nov. A. Distribution map; black star – the type locality; white circles – collection data examined in this paper; B, C. Living adult in a foraging trail of Lasius umbratus under a stone, Morioka-shi, Iwate-ken, Honshu (photographed by Naoya Nakaya); D. Snapped images from videos shot indoors, adults rubbing abdomen against a host ant worker; E. Clear image of a living adult from Nagano-ken.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
0081CFB869745162881E9CA6856EE80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483271	https://doi.org/10.3897/dez.72.158689.figure1	Figure 1. Explanations on research methods. A. Terminology of aedeagal median lobe (Homoeusa prolongata); al – apical lobe; av – apical valves; cp – copulatory piece; B. Terminology of spermatheca (Homoeusa laevigata); ap – apical part; bp – basal part; C. A simple bait trap set under stone with a swarm of Lasius meridionalis; D. A small camera shooting video of a Lasius fuliginosus group sp. foraging trail; E. Indoor observation setup using a plastic container with plaster.	Figure 1. Explanations on research methods. A. Terminology of aedeagal median lobe (Homoeusa prolongata); al – apical lobe; av – apical valves; cp – copulatory piece; B. Terminology of spermatheca (Homoeusa laevigata); ap – apical part; bp – basal part; C. A simple bait trap set under stone with a swarm of Lasius meridionalis; D. A small camera shooting video of a Lasius fuliginosus group sp. foraging trail; E. Indoor observation setup using a plastic container with plaster.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
DAAE202EF1DF5CEAA714ABFE50E7AF80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483272	https://doi.org/10.3897/dez.72.158689.figure2	Figure 2. Type materials deposited in NHM. A – C. The habiti of the type specimens of three species; A. The holotype of Homoeusa laevigata Sharp, 1888, male; B. The lectotype of Homoeusa japonica Sharp, 1874, male; C. The holotype of Homoeusa longicornis Sharp, 1888, male; D, E. The labels pinned with each specimen (all photographed by Keita Matsumoto).	Figure 2. Type materials deposited in NHM. A – C. The habiti of the type specimens of three species; A. The holotype of Homoeusa laevigata Sharp, 1888, male; B. The lectotype of Homoeusa japonica Sharp, 1874, male; C. The holotype of Homoeusa longicornis Sharp, 1888, male; D, E. The labels pinned with each specimen (all photographed by Keita Matsumoto).	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
DAAE202EF1DF5CEAA714ABFE50E7AF80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483273	https://doi.org/10.3897/dez.72.158689.figure3	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
DAAE202EF1DF5CEAA714ABFE50E7AF80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483279	https://doi.org/10.3897/dez.72.158689.figure9	Figure 9. Homoeusa japonica Sharp, 1874. A, B. Antenna; A. Male; B. Female; C. Pronotum and elytra; D. Zoomed view of elytron; E. Meso- and metaventrite; F, G. Posterior margin of 8 th sternite; F. Male; G. Female; H. Median lobe of aedeagus from lateral view; I, J. Paramere of aedeagus from lateral view: I. Western population; J. Eastern population; K, L. Spermatheca.	Figure 9. Homoeusa japonica Sharp, 1874. A, B. Antenna; A. Male; B. Female; C. Pronotum and elytra; D. Zoomed view of elytron; E. Meso- and metaventrite; F, G. Posterior margin of 8 th sternite; F. Male; G. Female; H. Median lobe of aedeagus from lateral view; I, J. Paramere of aedeagus from lateral view: I. Western population; J. Eastern population; K, L. Spermatheca.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
DAAE202EF1DF5CEAA714ABFE50E7AF80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483280	https://doi.org/10.3897/dez.72.158689.figure10	Figure 10. Homoeusa japonica Sharp, 1874. A – D. Variations of aedeagi; A. Takamatsu-shi, Kagawa-ken, Shikoku; B. Motosu-chô, Gifu-ken, Honshu; C. Ichikai-machi, Tochigi-ken, Honshu; D. Nakanojo-shi, Gunma-ken; E. Distribution map; black star – the type locality (“ Nagasaki ”); white circles – collection data examined in this paper; black circle – dissected male from the western population; red circle – dissected male from the eastern population; F. A walking adult in a foraging trail of Lasius cf. fliginosus on a fallen branch, Mt. Hikosan, Fukuoka-ken, Kyushu; G. Snapped images from videos shot indoors, raising abdomen to an ant; H. Clear image of living adult from Nagano-ken.	Figure 10. Homoeusa japonica Sharp, 1874. A – D. Variations of aedeagi; A. Takamatsu-shi, Kagawa-ken, Shikoku; B. Motosu-chô, Gifu-ken, Honshu; C. Ichikai-machi, Tochigi-ken, Honshu; D. Nakanojo-shi, Gunma-ken; E. Distribution map; black star – the type locality (“ Nagasaki ”); white circles – collection data examined in this paper; black circle – dissected male from the western population; red circle – dissected male from the eastern population; F. A walking adult in a foraging trail of Lasius cf. fliginosus on a fallen branch, Mt. Hikosan, Fukuoka-ken, Kyushu; G. Snapped images from videos shot indoors, raising abdomen to an ant; H. Clear image of living adult from Nagano-ken.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
DAAE202EF1DF5CEAA714ABFE50E7AF80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483271	https://doi.org/10.3897/dez.72.158689.figure1	Figure 1. Explanations on research methods. A. Terminology of aedeagal median lobe (Homoeusa prolongata); al – apical lobe; av – apical valves; cp – copulatory piece; B. Terminology of spermatheca (Homoeusa laevigata); ap – apical part; bp – basal part; C. A simple bait trap set under stone with a swarm of Lasius meridionalis; D. A small camera shooting video of a Lasius fuliginosus group sp. foraging trail; E. Indoor observation setup using a plastic container with plaster.	Figure 1. Explanations on research methods. A. Terminology of aedeagal median lobe (Homoeusa prolongata); al – apical lobe; av – apical valves; cp – copulatory piece; B. Terminology of spermatheca (Homoeusa laevigata); ap – apical part; bp – basal part; C. A simple bait trap set under stone with a swarm of Lasius meridionalis; D. A small camera shooting video of a Lasius fuliginosus group sp. foraging trail; E. Indoor observation setup using a plastic container with plaster.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
A1D83E522C585107A6EAC74C71EAF852.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483279	https://doi.org/10.3897/dez.72.158689.figure9	Figure 9. Homoeusa japonica Sharp, 1874. A, B. Antenna; A. Male; B. Female; C. Pronotum and elytra; D. Zoomed view of elytron; E. Meso- and metaventrite; F, G. Posterior margin of 8 th sternite; F. Male; G. Female; H. Median lobe of aedeagus from lateral view; I, J. Paramere of aedeagus from lateral view: I. Western population; J. Eastern population; K, L. Spermatheca.	Figure 9. Homoeusa japonica Sharp, 1874. A, B. Antenna; A. Male; B. Female; C. Pronotum and elytra; D. Zoomed view of elytron; E. Meso- and metaventrite; F, G. Posterior margin of 8 th sternite; F. Male; G. Female; H. Median lobe of aedeagus from lateral view; I, J. Paramere of aedeagus from lateral view: I. Western population; J. Eastern population; K, L. Spermatheca.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
A1D83E522C585107A6EAC74C71EAF852.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483283	https://doi.org/10.3897/dez.72.158689.figure13	Figure 13. Homoeusa prolongata Sawada, 1970 b. A. Antenna; B. Pronotum and elytra; C. Zoomed view of elytron; D. Meso- and metaventrite; E, F. Posterior margin of 8 th sternite: E. Male; F. Female; G. Median lobe of aedeagus from lateral view; H. Paramere of aedeagus from lateral view; I, J. Spermatheca.	Figure 13. Homoeusa prolongata Sawada, 1970 b. A. Antenna; B. Pronotum and elytra; C. Zoomed view of elytron; D. Meso- and metaventrite; E, F. Posterior margin of 8 th sternite: E. Male; F. Female; G. Median lobe of aedeagus from lateral view; H. Paramere of aedeagus from lateral view; I, J. Spermatheca.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
1C8E87D60BD15CAA97F75475F86EDEA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483272	https://doi.org/10.3897/dez.72.158689.figure2	Figure 2. Type materials deposited in NHM. A – C. The habiti of the type specimens of three species; A. The holotype of Homoeusa laevigata Sharp, 1888, male; B. The lectotype of Homoeusa japonica Sharp, 1874, male; C. The holotype of Homoeusa longicornis Sharp, 1888, male; D, E. The labels pinned with each specimen (all photographed by Keita Matsumoto).	Figure 2. Type materials deposited in NHM. A – C. The habiti of the type specimens of three species; A. The holotype of Homoeusa laevigata Sharp, 1888, male; B. The lectotype of Homoeusa japonica Sharp, 1874, male; C. The holotype of Homoeusa longicornis Sharp, 1888, male; D, E. The labels pinned with each specimen (all photographed by Keita Matsumoto).	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
1C8E87D60BD15CAA97F75475F86EDEA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483273	https://doi.org/10.3897/dez.72.158689.figure3	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
1C8E87D60BD15CAA97F75475F86EDEA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483277	https://doi.org/10.3897/dez.72.158689.figure7	Figure 7. Homoeusa laevigata Sharp, 1888. A. Antenna; B. Pronotum and elytra; C. Zoomed view of elytron; D. Meso- and metaventrite; E, F. Posterior margin of 8 th sternite; E. Male; F. Female; G. Median lobe of aedeagus from lateral view; H. Paramere of aedeagus from lateral view; I, J. Spermatheca.	Figure 7. Homoeusa laevigata Sharp, 1888. A. Antenna; B. Pronotum and elytra; C. Zoomed view of elytron; D. Meso- and metaventrite; E, F. Posterior margin of 8 th sternite; E. Male; F. Female; G. Median lobe of aedeagus from lateral view; H. Paramere of aedeagus from lateral view; I, J. Spermatheca.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
1C8E87D60BD15CAA97F75475F86EDEA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483278	https://doi.org/10.3897/dez.72.158689.figure8	Figure 8. Homoeusa laevigata Sharp, 1888. A. Distribution map; black star – the type locality (“ Seba ”); white circles – collection data examined in this paper; B. A walking adult (red arrow) in a foraging trail of Lasius nipponensis on a trunk, Matsumoto-shi, Nagano-ken, Honshu; C – E. Snapped images from videos shot indoors; C raising abdomen to an ant; D. Later begging for food and succeeding in trophallaxis; E. Licking host ant’s foreleg; F. Another shot of trophallaxis (photographed by Takuto Hashizume); G. Living individual from Kai-shi, Yamanashi-ken (photographed by Taku Shimada); H. Clear image of a living adult from Nagano-ken.	Figure 8. Homoeusa laevigata Sharp, 1888. A. Distribution map; black star – the type locality (“ Seba ”); white circles – collection data examined in this paper; B. A walking adult (red arrow) in a foraging trail of Lasius nipponensis on a trunk, Matsumoto-shi, Nagano-ken, Honshu; C – E. Snapped images from videos shot indoors; C raising abdomen to an ant; D. Later begging for food and succeeding in trophallaxis; E. Licking host ant’s foreleg; F. Another shot of trophallaxis (photographed by Takuto Hashizume); G. Living individual from Kai-shi, Yamanashi-ken (photographed by Taku Shimada); H. Clear image of a living adult from Nagano-ken.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
4B546179050E5C999F044986E90FD707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483272	https://doi.org/10.3897/dez.72.158689.figure2	Figure 2. Type materials deposited in NHM. A – C. The habiti of the type specimens of three species; A. The holotype of Homoeusa laevigata Sharp, 1888, male; B. The lectotype of Homoeusa japonica Sharp, 1874, male; C. The holotype of Homoeusa longicornis Sharp, 1888, male; D, E. The labels pinned with each specimen (all photographed by Keita Matsumoto).	Figure 2. Type materials deposited in NHM. A – C. The habiti of the type specimens of three species; A. The holotype of Homoeusa laevigata Sharp, 1888, male; B. The lectotype of Homoeusa japonica Sharp, 1874, male; C. The holotype of Homoeusa longicornis Sharp, 1888, male; D, E. The labels pinned with each specimen (all photographed by Keita Matsumoto).	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
4B546179050E5C999F044986E90FD707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483273	https://doi.org/10.3897/dez.72.158689.figure3	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
4B546179050E5C999F044986E90FD707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483281	https://doi.org/10.3897/dez.72.158689.figure11	Figure 11. Homoeusa longicornis Sharp, 1888. A, B. Antenna; A. Male; B. Female; C. Pronotum and elytra; D. Zoomed view of elytron; E. Meso- and metaventrite; F, G. Posterior margin of 8 th sternite; F. Male; G. Female; H. Median lobe of aedeagus from lateral view; I. Paramere of aedeagus from lateral view; J, K. Spermatheca.	Figure 11. Homoeusa longicornis Sharp, 1888. A, B. Antenna; A. Male; B. Female; C. Pronotum and elytra; D. Zoomed view of elytron; E. Meso- and metaventrite; F, G. Posterior margin of 8 th sternite; F. Male; G. Female; H. Median lobe of aedeagus from lateral view; I. Paramere of aedeagus from lateral view; J, K. Spermatheca.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
4B546179050E5C999F044986E90FD707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483282	https://doi.org/10.3897/dez.72.158689.figure12	Figure 12. Homoeusa longicornis Sharp, 1888. A – C. Variations of aedeagi; A. Mt. Ishizuchi-san, Ehime-ken, Shikoku; B. Maruseppu-chô, Hokkaido; C. Nakanojo-shi, Gunma-ken; D. Distribution map; black star – the type locality (“ Sapporo ”); white circles – collection data examined in this paper; E. A walking adult in a foraging trail of Lasius cf. fuliginous on a fallen branch, Iimori-toge, Saitama-ken, Honshu; F. Clear image of living adult from Aomori-ken.	Figure 12. Homoeusa longicornis Sharp, 1888. A – C. Variations of aedeagi; A. Mt. Ishizuchi-san, Ehime-ken, Shikoku; B. Maruseppu-chô, Hokkaido; C. Nakanojo-shi, Gunma-ken; D. Distribution map; black star – the type locality (“ Sapporo ”); white circles – collection data examined in this paper; E. A walking adult in a foraging trail of Lasius cf. fuliginous on a fallen branch, Iimori-toge, Saitama-ken, Honshu; F. Clear image of living adult from Aomori-ken.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
49EEBEA60488591F94B6A0D4266766A7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483276	https://doi.org/10.3897/dez.72.158689.figure6	Figure 6. Homoeusa rufescens (Sharp, 1874) and Homoeusa ovata Nozaki & Maruyama, 2022 b. A – C, G. Homoeusa rufescens; D – F, H. Homoeusa ovata. A, B, D, E. Posterior margin of 8 th sternite; A, D. Male; B, E. Female; C, F. Parameral apical lobe (modified from figs 13, 24 in Nozaki and Maruyama 2022 a); G, H. Distribution map; black star – the type locality; white circles – collection data examined in Nozaki and Maruyama (2022 a); black circles – additional data examined in this paper.	Figure 6. Homoeusa rufescens (Sharp, 1874) and Homoeusa ovata Nozaki & Maruyama, 2022 b. A – C, G. Homoeusa rufescens; D – F, H. Homoeusa ovata. A, B, D, E. Posterior margin of 8 th sternite; A, D. Male; B, E. Female; C, F. Parameral apical lobe (modified from figs 13, 24 in Nozaki and Maruyama 2022 a); G, H. Distribution map; black star – the type locality; white circles – collection data examined in Nozaki and Maruyama (2022 a); black circles – additional data examined in this paper.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
B8B9331D8F9F55778EA30E260A3174A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483273	https://doi.org/10.3897/dez.72.158689.figure3	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
B8B9331D8F9F55778EA30E260A3174A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483283	https://doi.org/10.3897/dez.72.158689.figure13	Figure 13. Homoeusa prolongata Sawada, 1970 b. A. Antenna; B. Pronotum and elytra; C. Zoomed view of elytron; D. Meso- and metaventrite; E, F. Posterior margin of 8 th sternite: E. Male; F. Female; G. Median lobe of aedeagus from lateral view; H. Paramere of aedeagus from lateral view; I, J. Spermatheca.	Figure 13. Homoeusa prolongata Sawada, 1970 b. A. Antenna; B. Pronotum and elytra; C. Zoomed view of elytron; D. Meso- and metaventrite; E, F. Posterior margin of 8 th sternite: E. Male; F. Female; G. Median lobe of aedeagus from lateral view; H. Paramere of aedeagus from lateral view; I, J. Spermatheca.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
B8B9331D8F9F55778EA30E260A3174A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483284	https://doi.org/10.3897/dez.72.158689.figure14	Figure 14. Homoeusa prolongata Sawada, 1970 b. A. Distribution map; black star – the type locality (“ Shiga Hights ”); white circles – collection data examined in this paper; B, C. Snapped images from videos shot indoors, adults raising abdomen to an ant; D. Clear image of a living adult from Nagano-ken.	Figure 14. Homoeusa prolongata Sawada, 1970 b. A. Distribution map; black star – the type locality (“ Shiga Hights ”); white circles – collection data examined in this paper; B, C. Snapped images from videos shot indoors, adults raising abdomen to an ant; D. Clear image of a living adult from Nagano-ken.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
E7F592F5B108547CA51BF5BB3E8692DA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483276	https://doi.org/10.3897/dez.72.158689.figure6	Figure 6. Homoeusa rufescens (Sharp, 1874) and Homoeusa ovata Nozaki & Maruyama, 2022 b. A – C, G. Homoeusa rufescens; D – F, H. Homoeusa ovata. A, B, D, E. Posterior margin of 8 th sternite; A, D. Male; B, E. Female; C, F. Parameral apical lobe (modified from figs 13, 24 in Nozaki and Maruyama 2022 a); G, H. Distribution map; black star – the type locality; white circles – collection data examined in Nozaki and Maruyama (2022 a); black circles – additional data examined in this paper.	Figure 6. Homoeusa rufescens (Sharp, 1874) and Homoeusa ovata Nozaki & Maruyama, 2022 b. A – C, G. Homoeusa rufescens; D – F, H. Homoeusa ovata. A, B, D, E. Posterior margin of 8 th sternite; A, D. Male; B, E. Female; C, F. Parameral apical lobe (modified from figs 13, 24 in Nozaki and Maruyama 2022 a); G, H. Distribution map; black star – the type locality; white circles – collection data examined in Nozaki and Maruyama (2022 a); black circles – additional data examined in this paper.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
41E2F413A9C851B5B401F51195E80F50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483273	https://doi.org/10.3897/dez.72.158689.figure3	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	Figure 3. Habiti of Japanese Homoeusa spp. A. Homoeusa translucida sp. nov., holotype, male; B. Homoeusa laevigata Sharp, 1888, male from Nagano-ken; C. Homoeusa japonica Sharp, 1874, male from Fukuoka-ken; D. Homoeusa longicornis Sharp, 1888, male from Hokkaido; E. Homoeusa prolongata Sawada, 1970 b, male from Nagano-ken; F. Homoeusa gigantea sp. nov., holotype, male; G. Homoeusa brevitarsis sp. nov., holotype, male.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
41E2F413A9C851B5B401F51195E80F50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483274	https://doi.org/10.3897/dez.72.158689.figure4	Figure 4. Homoeusa translucida sp. nov. A. Antenna; B. Pronotum and elytra; C. Zoomed view of elytron; D. Meso- and metaventrite; E, F. Posterior margin of 8 th sternite; E. Male; F. Female; G. Median lobe of aedeagus from lateral view; H. Paramere of aedeagus from lateral view; I. Spermatheca.	Figure 4. Homoeusa translucida sp. nov. A. Antenna; B. Pronotum and elytra; C. Zoomed view of elytron; D. Meso- and metaventrite; E, F. Posterior margin of 8 th sternite; E. Male; F. Female; G. Median lobe of aedeagus from lateral view; H. Paramere of aedeagus from lateral view; I. Spermatheca.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
41E2F413A9C851B5B401F51195E80F50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483275	https://doi.org/10.3897/dez.72.158689.figure5	Figure 5. Homoeusa translucida sp. nov. A. Distribution map; black star – the type locality; white circles – collection data examined in this paper; B. A photograph of a living adult collected with Lasius meridionalis at Mt. Hikosan, Fukuoka-ken, Kyushu.	Figure 5. Homoeusa translucida sp. nov. A. Distribution map; black star – the type locality; white circles – collection data examined in this paper; B. A photograph of a living adult collected with Lasius meridionalis at Mt. Hikosan, Fukuoka-ken, Kyushu.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
41E2F413A9C851B5B401F51195E80F50.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1483271	https://doi.org/10.3897/dez.72.158689.figure1	Figure 1. Explanations on research methods. A. Terminology of aedeagal median lobe (Homoeusa prolongata); al – apical lobe; av – apical valves; cp – copulatory piece; B. Terminology of spermatheca (Homoeusa laevigata); ap – apical part; bp – basal part; C. A simple bait trap set under stone with a swarm of Lasius meridionalis; D. A small camera shooting video of a Lasius fuliginosus group sp. foraging trail; E. Indoor observation setup using a plastic container with plaster.	Figure 1. Explanations on research methods. A. Terminology of aedeagal median lobe (Homoeusa prolongata); al – apical lobe; av – apical valves; cp – copulatory piece; B. Terminology of spermatheca (Homoeusa laevigata); ap – apical part; bp – basal part; C. A simple bait trap set under stone with a swarm of Lasius meridionalis; D. A small camera shooting video of a Lasius fuliginosus group sp. foraging trail; E. Indoor observation setup using a plastic container with plaster.	2025-12-03	Nozaki, Tsubasa;Maruyama, Munetoshi		Zenodo	biologists	Nozaki, Tsubasa;Maruyama, Munetoshi			
