taxonID	type	description	language	source
43EC867ECFDE545A83C94CB08DF980AB.taxon	description	Figs 3 G, 17 A – L, 18 (Japanese name: Nise-yamato-hirata-ariyadori)	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
43EC867ECFDE545A83C94CB08DF980AB.taxon	diagnosis	Diagnosis. This species is distinguished from other species of the genus by the following combination of characteristics: pronotal postero-lateral angle around 90 degrees (Fig. 17 C); metatarsi shorter than half of metatibiae (Fig. 17 F, G); apical lobe of paramere short, broad, and almost straight, length about 1 / 4 of whole paramere, apex distinctly truncate (Fig. 17 K). This species is similar to H. japonica and H. prolongata in general appearance, but it can be distinguished from these two species by its very short metatarsi and truncate apex of the parameral apical lobe.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
43EC867ECFDE545A83C94CB08DF980AB.taxon	description	Description. Body (Fig. 3 G) small, broad; dorsal surface mostly moderately polished. Head large; color brown; eyes small. Antennae (Fig. 17 A, B) short relative length of each antennal segment from first to 11 th, 2.48: 1.96: 1.39: 1.22: 1.04: 1.09: 1.00: 1.15: 1.15: 1.37: 3.26. Thorax. Pronotum (Fig. 17 C) subrectangular, transverse (PW / PL, 1.56 – 1.66), widest near middle, postero-lateral angle around 90 degrees, posterior margin slightly sinuate, reddish brown, surface finely covered with setae and punctures, gently polished and slightly reticulated. Elytra (Fig. 17 C, D) slightly widened posteriad, widest at middle, then narrowed roundly, posterior margins deeply notched near lateral corners, reddish brown, surface finely covered with setae and punctures; gently polished and finely reticulated. Hind wings developed. Mesoventral process (Fig. 17 E) very narrow, with weak medial carina, forming Y-shaped, apex pointed, reaching posterior 1 / 3 of mesocoxal cavities. Metaventral process slightly produced. Abdomen elongate, narrowed posteriad; surface covered with short setae and each posterior margin with long stout setae; finely reticulated. Male: Metatarsi (Fig. 17 F) as long as 1 / 3 of hind tibia. 8 th sternite (Fig. 17 H) produced roundly at postero-medial margin; median lobe of aedeagus (Fig. 17 J) slender, length about 0.61 mm, apical lobe narrow, slightly curved ventrally, apex gently pointed in lateral view, apical valves elongate and mitten-shaped, dorsal arms long and sharpened apically, basal part of copulatory piece small. Paramere (Fig. 17 K) length about 0.73 mm; apical lobe of paramere short and broad, length about 1 / 4 of whole paramere, almost straight, apex distinctly truncate; four setae, b located subapically, c irregularly located more dorsally; velum elongate and narrow. Female: Metatarsi (Fig. 17 G) slightly shorter than half of metatibia. 8 th sternite (Fig. 17 I) slightly rounded posterior margin; spermatheca (Fig. 17 L) apex of basal part slightly swollen, base largely hooklike.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
43EC867ECFDE545A83C94CB08DF980AB.taxon	description	Measurements (in mm) and ratios. Body parts (N = 8): BL ≈ 2.35 – 2.87; AL, 0.79 – 0.88; HW, 0.53 – 0.59; PL, 0.57 – 0.64; PW, 0.9 – 0.99; EL, 0.46 – 0.52; EW, 0.91 – 1.04; HTL, 0.6 – 0.74; PW / PL, 1.56 – 1.66; AL / PL, 1.35 – 1.46; HTL / PL, 1.04 – 1.22. Differences between sexes: HTL (male, N = 4), 0.69 – 0.74; HTL (female, N = 4), 0.6 – 0.66; HTL / PL (male, N = 4), 1.09 – 1.22; HTL / PL (female, N = 4), 1.04 – 1.09.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
43EC867ECFDE545A83C94CB08DF980AB.taxon	distribution	Distribution. Japan (Kii peninsula, Honshu) (Fig. 18).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
43EC867ECFDE545A83C94CB08DF980AB.taxon	etymology	Etymology. The specific name is derived from the Latin adjective, which means “ short footed ” and refers to the short tarsi of this new species.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
0081CFB869745162881E9CA6856EE80C.taxon	description	Figs 3 F, 15 A – I, 16 A – E (Japanese name: Ô-hirata-ariyadori)	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
0081CFB869745162881E9CA6856EE80C.taxon	diagnosis	Diagnosis. This species is distinguished from other species of the genus by the following combination of characteristics: pronotal posterolateral angle distinctly acute (Fig. 15 B); elytra distinctly widened posteriad entirely (Fig. 15 B, C); metaventral process slightly produced (Fig. 15 D); median lobe of aedeagus bulbous, length approximately 0.63 mm (Fig. 15 G). The appearance of this species is similar to H. laevigata and H. prolongata, but it can be distinguished from H. laevigata by its normal long setae of pronotum and elytra, and from H. prolongata by its larger body, aedeagus, and distinct acute posterolateral angle.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
0081CFB869745162881E9CA6856EE80C.taxon	description	Description. Body (Figs 3 F, 16 E) large, broad, somewhat limuloid; dorsal surface mostly moderately polished. Head small; color yellowish brown to brown; eyes small. Antennae (Fig. 15 A) short, relative length of each antennal segment from first to 11 th, 2.91: 2.25: 1.66: 1.27: 1.18: 1.2: 1.23: 1.18: 1.00: 1.45: 4.93. Thorax. Pronotum (Fig. 15 B) semicircular, transverse (PW / PL, 1.62 – 1.80), widest at posterior 1 / 3, postero-lateral angle distinctly acute, posterior margin distinctly sinuate, yellowish brown to brown, sometimes slightly transparent, surface finely covered with long setae and punctures, gently polished and slightly reticulated. Elytra (Fig. 15 B, C) distinctly widened posteriad entirely, posterior margins deeply notched near lateral corners, yellowish brown to brown, surface finely covered with long setae and punctures; gently polished and weakly reticulated. Hind wings developed. Mesoventral process (Fig. 15 D) very narrow, with indistinct medial carina, apex sharply pointed, reaching almost anterior margin of metaventrite. Metaventral process slightly produced (Fig. 15 D). Abdomen elongate, remarkably narrowed posteriad; surface sparsely covered with short setae and each posterior margin with long stout setae; weakly reticulated. Male: 8 th sternite (Fig. 15 E) weakly produced and angled at postero-medial margin; median lobe of aedeagus (Fig. 15 G) bulbous, length about 0.63 mm, apical lobe elogate and slightly curved ventrally, apex round in lateral view, apical valves elongate and mitten-shaped, dorsal arms short, basal part of copulatory piece large. Paramere (Fig. 15 H) length about 0.63 mm; apical lobe of paramere long, length about 1 / 3 of whole paramere, curved ventrally; four setae, b located more basally than c; velum small. Female: 8 th sternite (Fig. 15 F) weakly produced roundly postero-medial margin; spermatheca (Fig. 15 I) S-shaped, apex of basal part slightly sinuate, base largely hooklike.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
0081CFB869745162881E9CA6856EE80C.taxon	description	Measurements (in mm) and ratios. Body parts (N = 10): BL ≈ 2.48 – 4.3; AL, 0.92 – 1.15; HW, 0.55 – 0.64; PL, 0.67 – 0.83; PW, 1.09 – 1.39; EL, 0.51 – 0.62; EW, 1.06 – 1.34; HTL, 0.61 – 0.75; PW / PL, 1.62 – 1.80; AL / PL, 1.22 – 1.46; HTL / PL, 0.89 – 1.02.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
0081CFB869745162881E9CA6856EE80C.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu) (Fig. 16 A).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
0081CFB869745162881E9CA6856EE80C.taxon	etymology	Etymology. The specific name is derived from the Latin adjective, which means “ huge ” and refers to the large size of this new species.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
DAAE202EF1DF5CEAA714ABFE50E7AF80.taxon	description	Figs 2 B, E, 3 C, 9 A – L, 10 A – H (Japanese name: Yamato-hirata-ariyadori)	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
DAAE202EF1DF5CEAA714ABFE50E7AF80.taxon	diagnosis	Diagnosis. This species is distinguished from the other species of the genus by the following combination of characteristics: pronotal posterolateral angle around 90 degrees (Fig. 9 C); elytra distinctly widened posteriad entirely (Fig. 9 C); apical lobe of paramere long, length about half of whole paramere, curved ventrally, apex simply round (Fig. 9 I, J). This species is quite similar to H. longicornis in its general appearance; however, the round apex of the parameral apical lobe can separate it.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
DAAE202EF1DF5CEAA714ABFE50E7AF80.taxon	description	Measurements (in mm) and ratios. Body parts (N = 19): BL ≈ 2.14 – 3.27; AL, 0.81 – 1.03; HW, 0.51 – 0.66; PL, 0.48 – 0.63; PW, 0.78 – 1.03; EL, 0.42 – 0.51; EW, 0.85 – 1.02; HTL, 0.52 – 0.67; PW / PL, 1.59 – 1.71; AL / PL, 1.36 – 1.86; HTL / PL, 0.94 – 1.15.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
DAAE202EF1DF5CEAA714ABFE50E7AF80.taxon	distribution	Distribution. Japan (Honshu, Shikoku, Kyushu) (Fig. 10 E).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
1C8E87D60BD15CAA97F75475F86EDEA9.taxon	description	Figs 2 A, D, 3 B, 7 A – I, 8 A – H (Japanese name: Tsuya-hirata-ariyadori)	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
1C8E87D60BD15CAA97F75475F86EDEA9.taxon	diagnosis	Diagnosis. This species is distinguished from other species of the genus by the following combination of characteristics: body blackish, polished; pronotal postero-lateral angle acute (Fig. 7 B, C); elytra distinctly widened posteriad entirely (Fig. 7 B); pronotum and elytra covered with shortened setae (Fig. 7 B, C); metaventral process hardly produced; male 8 th sternite (Fig. 7 E) slightly emarginated at the middle of the posterior margin; median lobe of aedeagus bulbous; basal part of copulatory piece large. This species is similar to H. gigantea sp. nov. in body size and shape but can be distinguished by its blackish color and shortened setae on the pronotum and elytra.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
1C8E87D60BD15CAA97F75475F86EDEA9.taxon	description	Measurements (in mm) and ratios. Body parts (N = 10): BL ≈ 2.44 – 3.96; AL, 0.91 – 1.01; HW, 0.52 – 0.64; PL, 0.6 – 0.75; PW, 0.94 – 1.15; EL, 0.48 – 0.56; EW, 0.96 – 1.16; HTL, 0.6 – 0.72; PW / PL, 1.53 – 1.57; AL / PL, 1.34 – 1.51; HTL / PL, 0.96 – 1.03. Differences between sexes: AL / PL (male, N = 5), 1.43 – 1.51; AL / PL (female, N = 5), 1.34 – 1.42.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
1C8E87D60BD15CAA97F75475F86EDEA9.taxon	distribution	Distribution. Japan (Honshu) (Fig. 8 A).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
4B546179050E5C999F044986E90FD707.taxon	description	Figs 2 C, F, 3 D, 11 A – K, 12 A – F (Japanese name: Higenaga-hirata-ariyadori)	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
4B546179050E5C999F044986E90FD707.taxon	diagnosis	Diagnosis. This species is distinguished from the other species of the genus by the following combination of characteristics: pronotal posterolateral angle around 90 degrees (Fig. 11 C); elytra distinctly widened entirely posteriad (Fig. 11 C, D); apical lobe of paramere very long, length over half of whole paramere, curved ventrally, apex blade-shaped (Fig. 11 I). This species is quite similar to H. japonica in its general appearance but can be separated by the blade-like apex of its parameral apical lobe. Redescription. Body (Figs 3 D, 12 F) large, broad, stout; dorsal surface mostly moderately polished. Head large; color reddish brown to dark brown; eyes small. Antennae (Fig. 11 A, B) very long. Thorax. Pronotum (Fig. 11 C) subrectangular, transverse (PW / PL, 1.53 – 1.64), widest anterior 1 / 3, postero-lateral angle around 90 degrees, posterior margin slightly sinuate, reddish yellow to reddish brown, surface finely covered with setae and punctures, gently polished and slightly reticulated. Elytra (Fig. 11 C, D) distinctly widened posteriad entirely, posterior margins shallowly notched near lateral corners, reddish yellow to reddish brown, surface finely covered with setae and punctures; gently polished and weakly reticulated. Hind wings developed. Mesoventral process (Fig. 11 E) very narrow, with weak medial carina, forming Y-shaped, apex sharply pointed, reaching posterior 1 / 3 of mesocoxal cavities. Metaventral process weakly produced. Abdomen elongate, narrowed posteriad; surface covered with short setae and each posterior margin with long stout setae; finely reticulated. Male: Relative length of each antennal segment from first to 11 th, 1.66: 1.13: 1.16: 1.07: 1.00: 1.04: 1.00: 1.06: 1.01: 1.13: 2.25. 8 th sternite (Fig. 11 F) weakly rounded at posterior margin; median lobe of aedeagus (Fig. 11 H) slender, length about 0.63 mm, apical lobe elongated, ventral side slightly humped at middle, apex gently pointed in lateral view, apical valves simple and very long, without dorsal arms, basal part of the copulatory piece very small. Paramere (Fig. 11 I) length about 0.79 mm; apical lobe of paramere extremely long, length over half of whole paramere, curved ventrally, dilated apicad, apex blade-shaped; four setae, a far longer than others, b located most apically, velum elongate, broad and produced apically. Female: Relative length of each antennal segment from first to 11 th, 2.37: 1.57: 1.29: 1.09: 1.06: 1.00: 1.06: 1.08: 1.17: 1.40: 2.88. 8 th sternite (Fig. 11 G) truncated or slightly rounded at posterior margin; spermatheca (Fig. 11 J, K) S-shaped, apex of basal part not swollen, base largely hooklike.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
4B546179050E5C999F044986E90FD707.taxon	description	Measurements (in mm) and ratios. Body parts (N = 12): BL ≈ 2.58 – 3.54; AL, 0.98 – 1.3; HW, 0.59 – 0.7; PL, 0.59 – 0.72; PW, 0.97 – 1.13; EL, 0.5 – 0.59; EW, 0.94 – 1.13; HTL, 0.65 – 0.8; PW / PL, 1.53 – 1.64; AL / PL, 1.5 – 1.91; HTL / PL, 0.98 – 1.13. Differences between sexes: AL / PL (male, N = 5), 1.82 – 1.91; AL / PL (female, N = 7), 1.5 – 1.65; HTL / PL (male, N = 5), 1.05 – 1.13; HTL / PL (female, N = 7), 0.98 – 1.04.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
4B546179050E5C999F044986E90FD707.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, Shikoku) (Fig. 12 D).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
49EEBEA60488591F94B6A0D4266766A7.taxon	description	Fig. 6 D – F, H (Japanese name: Hime-hirata-ariyadori)	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
49EEBEA60488591F94B6A0D4266766A7.taxon	diagnosis	Diagnosis. See Nozaki and Maruyama (2022 a).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
49EEBEA60488591F94B6A0D4266766A7.taxon	description	Additional description. Relative length of each antennal segment from first to 11 th, 2.31: 1.98: 1.48: 1.29: 1.17: 1.00: 1.14: 1.14: 1.26: 1.4: 3.36. Hindwings developed. Male. 8 th sternite (Fig. 6 D) gently round at posterior margin; apical lobe of paramere (Fig. 6 F), setae b and c irregularly distributed at the acute corner on dorsal side. Female. 8 th sternite (Fig. 6 E) gently round at posterior margin.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
49EEBEA60488591F94B6A0D4266766A7.taxon	distribution	Distribution. Russia (Primorskyi krai), Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu) (Fig. 6 H).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
B8B9331D8F9F55778EA30E260A3174A6.taxon	description	Figs 3 E, 13 A – I, 14 A – D (Japanese name: Hirata-ariyadori)	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
B8B9331D8F9F55778EA30E260A3174A6.taxon	diagnosis	Diagnosis. This species is distinguished from other species of the genus by the following combination of characteristics: a slightly produced metaventral process (Fig. 13 D), median lobe of aedeagus bulbous with a length of approximately 0.47 mm (Fig. 13 G). This species is similar to H. ovata and H. japonica, but it can be distinguished by a more flattened and semicircular pronotum, less produced metaventral process, and a bulbous median lobe of the male aedeagus.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
B8B9331D8F9F55778EA30E260A3174A6.taxon	description	Measurements (in mm) and ratios. Body parts (N = 13): BL ≈ 1.9 – 2.78; AL, 0.67 – 0.81; HW, 0.43 – 0.52; PL, 0.46 – 0.61; PW, 0.73 – 0.92; EL, 0.37 – 0.51; EW, 0.73 – 0.98; HTL, 0.4 – 0.48; PW / PL, 1.51 – 1.65; AL / PL, 1.22 – 1.53; HTL / PL, 0.8 – 0.9.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
B8B9331D8F9F55778EA30E260A3174A6.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu) (Fig. 14 A).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
E7F592F5B108547CA51BF5BB3E8692DA.taxon	description	Fig. 6 A – C, G (Japanese name: Hoso-hirata-ariyadori)	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
E7F592F5B108547CA51BF5BB3E8692DA.taxon	diagnosis	Diagnosis. See Nozaki and Maruyama (2022 a).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
E7F592F5B108547CA51BF5BB3E8692DA.taxon	description	Additional description. Relative length of each antennal segment from first to 11 th, 2.27: 1.63: 1.37: 1.27: 1.02: 1.02: 1.00: 1.05: 1.07: 1.44: 3.34. Hind wings developed. Male. 8 th sternite (Fig. 6 A) weakly round at posterior margin; four setae of parameral apical lobe (Fig. 6 C) normally arranged. Female. 8 th sternite (Fig. 6 B) gently round at posterior margin.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
E7F592F5B108547CA51BF5BB3E8692DA.taxon	distribution	Distribution. Japan (Honshu, Shikoku, Kyushu) (Fig. 6 G).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
41E2F413A9C851B5B401F51195E80F50.taxon	description	Figs 3 A, 4 A – I, 5 A, B Japanese name: Sukashi-hirata-ariyadori	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
41E2F413A9C851B5B401F51195E80F50.taxon	diagnosis	Diagnosis. This species can be easily distinguished from the other species of the genus by the following combination of characteristics: body color yellowish orange; pronotum strongly transverse (PW / PL, 1.84 – 2.05), postero-lateral angle distinctly acute (Fig. 4 B); metaventral process weakly produced (Fig. 4 D); pronotum and elytra sparsely covered with modified stout setae (Fig. 4 B, C); median lobe of aedeagus apical lobe straight, apex round and slightly humped in lateral view, apical valves very large and cheliform (Fig. 4 G); spermatheca (Fig. 4 I) L-shaped, basal part very long. This species is slightly similar to H. gigantea sp. nov. in color but differs in its pronotal and setal shapes.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
41E2F413A9C851B5B401F51195E80F50.taxon	description	Description. Body (Figs 3 A, 5 B) small, broad; dorsal surface mostly polished. Head large; color yellowish orange; eyes large. Antennae (Fig. 4 A) long; relative length of each antennal segment from first to 11 th, 2.48: 1.96: 1.39: 1.22: 1.04: 1.09: 1.00: 1.15: 1.15: 1.37: 3.26. Thorax. Pronotum (Fig. 4 B) subrectangular, strongly transverse (PW / PL, 1.84 – 2.05), widest near middle, postero-lateral angle distinctly acute, posterior margin distinctly sinuate, color yellowish orange, subtransparent, surface sparsely covered with modified stout setae and punctures, gently polished without reticulation. Elytra (Fig. 4 B, C) slightly widened posteriad, widest at middle, then narrowed roundly, posterior margins shallowly notched near lateral corners, color yellowish orange, subtransparent, surface sparsely covered with modified stout setae and punctures, gently polished without reticulation. Hind wings developed. Mesoventral process (Fig. 4 D) narrow, with weak medial carina, apex gently pointed, reaching posterior 1 / 3 of mesocoxal cavities, Metaventral process weakly produced. Abdomen elongate, narrowed posteriad; surface sparsely covered with short setae and each posterior margin with long stout setae; covered with fine vertical furrows. Male: 8 th sternite (Fig. 4 E) widely protruding at middle of posterior margin; median lobe of aedeagus (Fig. 4 G) elongate, length about 0.57 mm, apical lobe straight, apex round and slightly humped in lateral view, apical valves large and cheliform, dorsal arms boot-shaped; basal part of copulatory piece very large and having claw-shaped projection, basal corner acutely pointed. Paramere (Fig. 4 H) length about 0.59 mm; apical lobe of paramere long, length over 1 / 3 of whole paramere, curved ventrally, slightly dilated apicad; four setae, c located a little more apically than b; velum narrow and membranous. Female: 8 th sternite (Fig. 4 F) slightly protruding at the middle of posterior margin; spermatheca (Fig. 4 I) L-shaped, basal part very long, apex of basal part not swollen, base almost straight and only slightly swollen.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
41E2F413A9C851B5B401F51195E80F50.taxon	description	Measurements (in mm) and ratios. Body parts (N = 10): BL ≈ 2.13 – 2.91; AL, 0.93 – 1.07; HW, 0.51 – 0.57; PL, 0.5 – 0.59; PW, 1 – 1.13; EL, 0.47 – 0.52; EW, 0.98 – 1.16; HTL, 0.69 – 0.76; PW / PL, 1.84 – 2.05; AL / PL, 1.69 – 1.95; HTL / PL, 1.29 – 1.46.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
41E2F413A9C851B5B401F51195E80F50.taxon	distribution	Distribution. Japan (Honshu, Shikoku, Kyushu) (Fig. 5 A).	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
41E2F413A9C851B5B401F51195E80F50.taxon	etymology	Etymology. The specific name is derived from the Latin adjective, which means “ transparent ” and refers to the character of the pronotum of this new species.	en	Nozaki, Tsubasa, Maruyama, Munetoshi (2025): Taxonomy of Homoeusa Kraatz, 1856 (Coleoptera, Staphylinidae, Aleocharinae) from the East Palearctic: II. Revision of Japanese species. Deutsche Entomologische Zeitschrift 72 (2): 525-555, DOI: 10.3897/dez.72.158689
