identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
C46D87F6BB5C032C84C5B49DFA836EC7.text	C46D87F6BB5C032C84C5B49DFA836EC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha Silvestri 1897	<div><p>Genus  Camptomorpha Silvestri, 1897</p><p>Camptomorpha Silvestri, 1897a: 10, originally proposed invalidly for two species; Brölemann, 1916: 552; Attems, 1931: 44; 1938: 69; Hoffman, 1953: 120, clarification of the genus and tYpe species identitY; 1975: 185, assignment to  Lepturodesmini; 1980: 152; Schubart, 1955: 508; Jeekel, 1971: 252; Bouzan et al., 2018: 11.</p><p>Type species:  C. dOrsAlis Silvestri, 1897a: 10, by subsequent designation of Attems [1931: 44].</p><p>Genus feminine in gender [Jeekel, 1971], even though sometimes erroneously treated as masculine.</p><p>Number of species: 11 described, one new treated below, and several more still undescribed.</p><p>=  PHAntAsmOdesmus Verhoeff, 1927: 509, introduced as a subgenus of  Leptodesmus, synonymized with  Leptodesmus by Attems, 1938: 6, synonymized under  Camptomorpha bY Hoffman, 1953: 120, listed as a generic sYnonYm bY Hoffman, 1980: 152; Kraus, 1956: 142; Jeekel, 1971: 279.</p><p>Type species:  LeptOdesmus pulvillAtus Attems, 1898: 391, by monotypy.</p><p>=  Eucampesmus Chamberlin, 1941: 489, synonymized with PHAntasmodesmus by Kraus, 1956: 141, synonymized with  Camptomorpha bY Hoffman, 1953: 120, listed as a generic sYnonYm bY Hoffman, 1980: 152; Jeekel, 1971: 262.</p><p>Type species:  EucAmpesmus Orites Chamberlin, 1941: 489, by original designation.</p><p>=  Ptyxesmus Chamberlin, 1941: 489,  syn.n., after Hoffman [in litt.].</p><p>Type species:  Ptyxesmus atyphus Chamberlin, 1941: 489, by original designation.</p><p>DIAGNOSIS. A genus of  Lepturodesmini with ♂ coxae 3 usually produced or lobed mesad, almost or fully in contact. Gonopods remarkably complex, sternal remnants strongly reduced or absent; coxites large and globose, devoid of dorsal apophyses; telopodites tripartite, consisting of two large laminae (anterior acropodite and posterior prefemoral process) and a fullY independent solenomere branch between both [Hoffman, 1975, in litt.].</p><p>BRIEF DESCRIPTION. Adults medium-sized (32–50 mm long and 4–6 mm wide), usually dark olive to brown and usually devoid of a bright pattern, more rarely lighter, red-grey with a yellowish axial stripe, yellowish or yellowish white paraterga and reddish-yellow venter and legs. Body with 20 rings, increasingly wide until ring 5, thereafter increasingly attenuated. Antennae rather short and slender to robust, up to only poorly clavate; in length, antennomeres 2–6&gt;&gt;1=7&gt;8. Collum usually broadly rounded laterally, only occasionally triangular and almost pointed. Metatergal surface smooth and shining, often leatherY, verY finelY striolate, with neither transverse sulci nor an axial line. Dorsum moderately convex, especially anteriorly. Sides smooth, onlY occasionallY finelY granulate. Rings 1–4(5) very compact and tightly following one after the other, vs remaining rings, more loose and gradually attenuating towards telson. Paraterga well-developed, on ring 19 smallest, set at about upper 1/2 to 1/4 body height, largely rimmed throughout, broadly and more or less regularly rounded anterolaterally and laterally, typically smooth, only occasionally and only on anterior body rings with a small marginal tooth towards anterior corner, caudal corner usually broadly to narrowly rounded, in poreless paraterga sometimes pointed. Pore formula normal: 5, 7, 9, 10, 12, 13, 15–19. Ozopores placed inside ovoid pits or grooves on distinct ovoid calluses (= peritremata), dorsolateral to lateral in position. Epiproct usuallY short, finger-shaped, tip only occasionally with a paramedian pair of small claws directed ventrocaudally. Hypoproct often with a prominent median projection at caudal margin, longer than paramedian setigerous knobs. Pleurosternal carinae inconspicuous, present only on a few anterior rings.</p><p>Sterna usuallY unmodified, broad, flat, bare and at most with faint cross-impressions. Legs long and slender, anterior ♂ prefemora sometimes with distoventral swellings and most ♂ tibiae with apicoventral pads/chelae subtending the bases of tarsi. ♂ coxae 3 each with a ventral process at apex, usuallY densely setose.</p><p>Gonopodal aperture kidney-shaped, large, taking up most of prozonum 7, but not shifted beyond it, with clearly elevated and rounded lateral sides and a regularly rounded caudal shelf inside (Fig. 12I). Gonopods mostly remarkably complex, characteristic of the genus, each tripartite (see Diagnosis above). Gonopodal coxite large and globose, usually with two macrosetae; cannula not set inside a marginal notch; prefemorite (= a densely setose prefemoral region) proportionately smaller, forming an obtuse angle with telopodite; both acropodite and solenomere set off bY a distinct cingulum formed bY a strong flexure on lateral side.</p><p>The following species of  Camptomorpha are considered valid, all listed in alphabetic order, each accompanied by brief descriptive notes and available meaningful illustrations. The content and qualitY of these notes varY significantlY depending on the amount of published information available. Only the new species is being described and illustrated in due detail.</p></div>	https://treatment.plazi.org/id/C46D87F6BB5C032C84C5B49DFA836EC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB5C032F875EB6D9FDED6E6A.text	C46D87F6BB5C032F875EB6D9FDED6E6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha aberrans Kraus 1957	<div><p>Camptomorpha aberrans Kraus, 1957</p><p>Fig. 1.</p><p>CAmptOmOrpHA AberrAns 
Kraus, 1957: 98, plate 8, figs 11–14. HolotYpe ♂, SMF.</p><p>BRIEF DESCRIPTIVE NOTES (after Kraus [1957]). Width 4.7 mm. Coloration uniformly red-brown. Paraterga broadly and regularly rounded, caudal corner obtuse and rounded; midbodY pore-bearing calluses clearlY set off from anterolateral rim (Fig. 1A). Epiproct simple, short and finger-shaped. ♂ legs ventrallY with prefemora bearing semi-circular bulges distally and with tibiae carrying chelae (= apical soles subtending bases of tarsi).</p><p>Gonopods (Fig. 1B, C) with both prefemoral process (PfP) and acropodite (A) subequal in length, slender and relatively simple, poorly armed; PfP with two small and acuminate projections near its middle, and terminating in an acuminate apex; A divided into two acuminate branches near its apical region; solenomere (sl) rather rod-shaped, curved only apically.</p><p>DISTRIBUTION. Peru, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.30386&amp;materialsCitation.latitude=-9.892722" title="Search Plazi for locations around (long -76.30386/lat -9.892722)">AcomaYo</a> near Huanuco, 9°53′ 33.8″S 76°18′13.9″W, 2700 m [Kraus, 1957]  .</p></div>	https://treatment.plazi.org/id/C46D87F6BB5C032F875EB6D9FDED6E6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB5F032F84C3B60DFC2F6D80.text	C46D87F6BB5F032F84C3B60DFC2F6D80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha atypha (Chamberlin 1941)	<div><p>Camptomorpha atypha (Chamberlin, 1941)</p><p>Fig. 2.</p><p>Ptyxesmus atyphus Chamberlin, 1941: 489, figs 101 and 102. HolotYpe ♂, AMNH, but presentlY misplaced [Hoffman, in litt.].</p><p>BRIEF DESCRIPTIVE NOTES (after Chamberlin [1941]). Width 5.0 mm. Coloration dark olive dorsally, paraterga in part more brownish, legs and antennae dark brown. Collum acutely narrowed on each side (Fig. 2A). Paraterga broadly and regularly rounded, rimmed throughout; caudal corners drawn past rear tergal margins only in caudal body third. Epiproct simple, short and finger-shaped. ♂ legs ventrallY with prefemora bearing semi-circular bulges distally and with tibiae carrying chelae apically.</p><p>Gonopods (Fig. 2B, C) with both prefemoral process (PfP) and acropodite (A) subequal in length, A being relatively simple and leaf-shaped, with several grooves at external margin, apex directed basally; PfP armed and broader, with an irregular internal margin and a small ascending process near middle; solenomere (sl) simple.</p><p>DISTRIBUTION. Peru, Loreto Dept., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.25383&amp;materialsCitation.latitude=-3.7491195" title="Search Plazi for locations around (long -73.25383/lat -3.7491195)">Iquitos</a>, 3°44′56.83″S 73°15′13.79″W [Chamberlin, 1941]  .</p><p>Hoffman [in litt.] provided a sketch of a gonopod of this species from a near-topotype, reproduced here in Fig. 2C. He also recorded  C. atypha from near Tabatinga, Brazil, as well as Rio Yanayacu near Iquitos and Rio Marañon ca 2 km above Nauta, Peru.</p></div>	https://treatment.plazi.org/id/C46D87F6BB5F032F84C3B60DFC2F6D80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB5F032E874AB623FD5D6E13.text	C46D87F6BB5F032E874AB623FD5D6E13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha digitata (Schubart 1947)	<div><p>Camptomorpha digitata (Schubart, 1947)</p><p>Fig. 3.</p><p>Leptodesmus (Desmoleptus) digitAtus Schubart, 1947 b: 43, figs 42–43. Syntypes, MNRJ.</p><p>CAmptOmOrpHA digitAtus (sic!) — Hoffman, 1953: 124.</p><p>Leptodesmus ( PHAntAsmOdesmus)  digitAtus — Kraus, 1956: 142,  Camptomorpha by implication.</p><p>BRIEF DESCRIPTIVE NOTES (after Schubart [1947]). Length 39–40 mm, width 5.2–5.4 mm. Coloration castaneous, clypeus, labrum, sulcus on vertex, basal antennomere, venter, legs and epiproct lighter, sides of collum and rims of paraterga yellowish. Paraterga broadly and regularly rounded, rimmed throughout (Fig. 3A–C); caudal corners increasingly acute, triangular and pointed from ring 14 on. Epiproct simple, short and finger-shaped. ♂ legs ventrallY with prefemora bearing semi-circular bulges distally and with all tibiae but two last pairs carrying chelae apically.</p><p>Gonopods (Fig. 3D, E) with prefemoral process (PfP) clearly longer than acropodite (A), both being rather elaborate and armed; PfP with two small and acuminate projections near middle ending in an irregularly shaped acuminate apex, the latter directed basally; apical margin of A divided into two branches, internal one being slender, vs a broader external one; solenomere (sl) simple and slender, subflagelliform distallY.</p><p>DISTRIBUTION. Brazil, Pará, Aurá near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.397614&amp;materialsCitation.latitude=-1.40825" title="Search Plazi for locations around (long -48.397614/lat -1.40825)">Belem</a>, 1°24′ 29.7″S 48°23′51.4″W [Schubart, 1947]. Hoffman [in litt.] added also Parantins, Amazônas state, almost 800 km W of the type locality  .</p><p>Hoffman [in litt.] revised tYpe material and noted that Schubart’s [1947 b] original drawing of the left gonopod was inaccurate and misleading, providing instead several sketches both of an intact gonopod and several body rings (Fig. 3).</p></div>	https://treatment.plazi.org/id/C46D87F6BB5F032E874AB623FD5D6E13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB5E032984DEB6B5FD106EB0.text	C46D87F6BB5E032984DEB6B5FD106EB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha dorsalis Silvestri 1897	<div><p>Camptomorpha dorsalis Silvestri, 1897</p><p>Fig. 4.</p><p>CAmptOmOrpHA dOrsAlis Silvestri, 1897b: 10, figs 29–31 (sYntYpes of both sexes, ZMT); Attems, 1938: 70; Hoffman, 1953: 123, fig. 1; Jeekel, 1971: 252.</p><p>LeptOdesmus dOrsAlis — Attems, 1899: 427.</p><p>CAmptOmOrpHA perprOximA 
Silvestri, 1897b: 11, figs 32–33, syn.n. (sYntYpes of both sexes, ZMT, after Hoffman [in litt.]).</p><p>LeptOdesmus perprOximA (sic!) — Attems, 1899: 427.</p><p>CAmptOmOrpHA perprOximA — Attems, 1938: 71; Hoffman, 1953: 123.</p><p>BRIEF DESCRIPTIVE NOTES (after Silvestri [1897b] and Hoffman [in litt.]). Length 32–33 mm, width 4.0– 4.6 mm. Coloration red-grey or brown with a yellowish axial stripe from collum to tip of epiproct, sides, entire epiproct and legs sometimes yellow; metaterga, venter and legs yellowish to brown and reddish-yellow; prozona, except anteriormost ones, sometimes with two whitish paramedian spots; paraterga sometimes yellowish white both dorsally and ventrally. Paraterga broadly and regularly rounded, rimmed throughout, paraterga 2 sometimes with a small lateral tooth towards anterolateral corner; caudal corners increasingly acute from ring 5 on (Fig. 4A). Epiproct short and finger-shaped, with a paramedian pair of small apical claws directed caudoventrallY (Fig. 4C). ♂ legs 3–20 ventrallY with prefemora bearing semi-circular bulges distally and with tibiae carrYing chelae apicallY (Fig. 4B). ♂ coxae 2 enlarged and apicallY truncate; ♂ coxae 3 (Fig. 4B) also enlarged and apicallY truncate, denselY setose, almost in contact mediallY; ♂ coxae 4 and 5 produced ventrad into prominent, blunt, rounded lobes.</p><p>Gonopods (Fig. 4D–I) with prefemoral process (PfP) and acropodite (A) subequal in length, both being rather simple, unarmed, with spatuliform apices; PfP strongly twisted; solenomere (sl) simple, helicoid and slender, subflagelliform distallY.</p><p>DISTRIBUTION. For  C. dOrsAlis: Ecuador, San José, 1°29′03.1″S 80°19′59.7″W [Silvestri, 1897b]. Hoffman [in litt.] elucidated the type locality as actually lying at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.33325&amp;materialsCitation.latitude=-1.4841945" title="Search Plazi for locations around (long -80.33325/lat -1.4841945)">Rio San José</a>, ca 25 km NW of Gualaquiza.</p><p>For  C. perprOximA: Ecuador, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.94881&amp;materialsCitation.latitude=-4.062417" title="Search Plazi for locations around (long -78.94881/lat -4.062417)">Valle de Zamora</a> (Santiago Zamora = Zamora Chinchipe), 4°03′44.7″S 78°56′55.7″W [Silvestri, 1897b]  .</p><p>JUSTIFICATION OF SYNONYMY. Hoffman [in litt.] recorded and depicted ♂ material of this species from Cushuene, Pastaza Prov. (Fig. 4A–E) and Hacienda Santa Inés (1º25′ S 78º12′ W) on Rio Pastaza between Rio Mapoto and Rio Topo, Tungurahua Prov. (Fig. 4F, J), Ecuador, comparing it directly with a syntype of  C. perprOximA (Fig. 4G, H). The results unequivocally prove the synonymy and quite a vast distribution of  C dOrsAlis (=  C. perprOximA,  syn.n.) in Ecuador.</p></div>	https://treatment.plazi.org/id/C46D87F6BB5E032984DEB6B5FD106EB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB59032B84C9B6D5FE9A6CCA.text	C46D87F6BB59032B84C9B6D5FE9A6CCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha orites (Chamberlin 1941)	<div><p>Camptomorpha orites (Chamberlin, 1941)</p><p>Fig. 5.</p><p>EucAmpesmus Orites Chamberlin, 1941: 489, figs 97–100 (holotYpe ♂, AMNH); Jeekel 1971: 262.  CAmptOmOrpHA Oreites (sic!) — Hoffman, 1953: 124.  Leptodesmus ( PHAntAsmOdesmus)  Orites — Kraus, 1956: 142,</p><p>Camptomorpha by implication. BRIEF DESCRIPTIVE NOTES (after Chamberlin</p><p>chocolate brown with light brown or yellowish paraterga, and antennae and legs light brown. Paraterga broadly and regularly rounded, lateral rim smooth (Fig. 5A), most anterior ones each with a small tooth towards anterior corner, caudal corner rather narrowly rounded; midbody pore-bearing calluses (= peritremata) set off from anterolateral rim. ♂ legs ventrallY with tibiae carrying apical chelae.</p><p>Gonopods (Fig. 5B–D) with both prefemoral process (PfP) and acropodite (A) subequal in length, both very complex and heavily armed; PfP broad, with two elongated and acute projections along its middle portion, ending in another elongated and acute apex directed laterally; A with an irregular internal margin with several small indentations, apical portion semi-spatulate in shape, with a triangular projection in central portion of apex; solenomere (sl) slender and simple, only slightly curved apically.</p><p>DISTRIBUTION. Peru, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.94092&amp;materialsCitation.latitude=-7.4486666" title="Search Plazi for locations around (long -73.94092/lat -7.4486666)">Contayo Hills</a>, Rio Tapiche, 7°26′55.2″S 73°56′27.3″W, 700 ft [Chamberlin, 1941]  .</p><p>[1941]). Length ca 48 mm, width 6.0 mm. Coloration dark to</p><p>Hoffman [in litt.] located a strict topotYpe ♂ in the AMNH collection and depicted its right gonopod in proper detail (Fig. 5B–D) .</p></div>	https://treatment.plazi.org/id/C46D87F6BB59032B84C9B6D5FE9A6CCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB5B032B851EB4F6FDCC6F3C.text	C46D87F6BB5B032B851EB4F6FDCC6F3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha ortizi Kraus 1959	<div><p>Camptomorpha ortizi Kraus, 1959</p><p>Fig. 6.</p><p>CAmptOmOrpHA Ortizi 
Kraus, 1959:266,figs 4–7. HolotYpe ♂, SMF.</p><p>BRIEF DESCRIPTIVE NOTES (after Kraus [1959]). Width 4.4 mm. Coloration dull red-brown, paraterga and their caudal corners, as well as epiproct and legs yellow-white. Paraterga broadly and regularly rounded, caudal corners mostly rounded, pore-bearing calluses (= peritremata) distinctlY set off from a completely smooth lateral rim (Fig. 6A). Epiproct short and finger-shaped, with a paramedian pair of small apical claws. Hypoproct with an apical tooth between both setigerous knobs. ♂ legs ventrallY with prefemoral apical knobs graduallY reduced towards midbody legs and with all tibiae carrying apical chelae.</p><p>Gonopods (Fig. 6B–D) with prefemoral process (PfP) only slightly longer than acropodite (A), both only moderately complex and poorly armed; PfP spoon-shaped, internal margin near apex with a small invagination subsequently forming a small acute process; A more slender, with a slightly broadened and spatuliform apex, also featuring several small indentations at apical margin; solenomere (sl) unusually slender and bipartite, ventral branch much longer than and flagelliform compared to dorsal branch.</p><p>DISTRIBUTION. Peru, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.726944&amp;materialsCitation.latitude=-7.173333" title="Search Plazi for locations around (long -76.726944/lat -7.173333)">Yuanjui</a> (Juanjuí), Rio Huallaga, 7°10′24.0″S 76°43′37.0″W [Kraus, 1959]  .</p></div>	https://treatment.plazi.org/id/C46D87F6BB5B032B851EB4F6FDCC6F3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB5B03258532B758FE646F33.text	C46D87F6BB5B03258532B758FE646F33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha papillosa (Attems 1931)	<div><p>Camptomorpha papillosa (Attems, 1931)</p><p>Fig. 7.</p><p>Leptodesmus (Leptodesmus) pApillOsus Attems, 1931: 10, figs 1–4 (sYntYpes, ZMH); Attems, 1938: 13, fig. 8; Weidner [1960: 84], listed in type catalogue.</p><p>CAmptOmOrpHA pApillOsus (sic!) — Hoffman, 1953: 123.</p><p>Leptodesmus ( PHAntAsmOdesmus)  pApillOsus — Kraus, 1956: 142,  Camptomorpha by implication.</p><p>Pseudoleptodesmus (Pseudoleptodesmus) cOrdillerAnus 
Attems, 1931: 28, figs 40–42, syn.n. (sYntYpes, ZMH, after Hoffman [in litt.]).</p><p>Leptodesmus (Pseudoleptodesmus) cOrdillerAnus — Attems, 1938: 39, fig. 42.</p><p>CAmptOmOrpHA cOrdillerAnus (sic!) — Hoffman, 1953: 122.</p><p>Leptodesmus ( PHAntAsmOdesmus)  cOrdillerAnus — Kraus, 1956: 142,  Camptomorpha by implication.</p><p>BRIEF DESCRIPTIVE NOTES (after Attems [1931, 1938]). Width 4.0 mm. Coloration dark castaneous brown with a vague, lighter, axial stripe; antennae and legs dark red-brown. Paraterga relatively small and faintly rimmed, caudal corners mostly sharp and dentiform (Fig. 7A). Epiproct simple and finger-shaped, directed slightlY dorsad. HYpoproct subtrapeziform. with a small apical tooth between both thick setigerous knobs. ♂ legs ventrallY with all tibiae but four last pairs carrYing apical chelae; ♂ coxae 3 each with a strong, blunt, denselY setose process directed forward.</p><p>Gonopods (Fig. 7B–F) with both prefemoral process (PfP) and acropodite (A) subequal in length, very complex and strongly armed; PfP also broad, with an irregular internal margin and a small digitiform process near middle; A robust and leaf-shaped, with several grooves at external margin, one of the apical branches being directed basally; solenomere (sl) simple and only slightly curved.</p><p>DISTRIBUTION. Ecuador, Sabanilla, 1°50′45.0″S 80°13′ 04.3″W, East slope of the Cordilleras [Attems, 1931, 1938].</p><p>JUSTIFICATION OF SYNONYMY. Hoffman [in litt.] provided ample evidence of the synonymy of C.  pApillOsA with C. cOrdillerAnA (Fig. 7B–F). Indeed, both these formal species came from the same collection and even the same population, even though Attems [1931] assigned them to different genera! Following Hoffman [in litt.], we choose to use the name C.  papillOsA because it has 26 pages of priority over C. cOrdillerAnA, while both names have been used the same number of times in the primary taxonomic literature.</p></div>	https://treatment.plazi.org/id/C46D87F6BB5B03258532B758FE646F33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB5503248530B756FD086E40.text	C46D87F6BB5503248530B756FD086E40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha pulvillata (Attems 1898)	<div><p>Camptomorpha pulvillata (Attems, 1898)</p><p>Fig. 8.</p><p>LeptOdesmus pulvillAtus Attems, 1898: 391, figs 132–133 (holotYpe  ♂, NHMW); Attems, 1938: 12, fig. 6 .</p><p>CAmptOmOrpHA pulvillAtus (sic!) — Hoffman, 1953: 123.</p><p>LeptOdesmus ( PHAntAsmOdesmus)  pulvillAtus — Kraus, 1956: 142,  Camptomorpha by implication.</p><p>BRIEF DESCRIPTIVE NOTES (after Attems [1898, 1938]). Length 56 mm, width of pro- and metazona 4.0 mm and 6.0 mm, respectively. Coloration yellowish brown; head, sides and anterior margins of metazona castaneous brown; antennae, lateral rims of paraterga and legs yellow; prozona yellowish with a brown axial line. Paraterga unusually narrow, but strongly rimmed, both anterior and caudal corners mostly rounded, caudal corners on rings 4–7 rectangular, thereafter dentiform; lateral rim on rings 1–4 finelY serrate. Epiproct simple and finger-shaped, directed slightlY dorsad. HYpoproct with a small apical tooth between both thick setigerous knobs. ♂ legs ventrally with all prefemora but two last pairs carrying apical swellings, and tibiae with very large apical chelae, tarsi being unusuallY small and slender (Fig. 8A). ♂ coxae 1–3 nearlY in touch medially, each coxa 3 apically with a slender, straight and densely setose ventral process.</p><p>Gonopods (Fig. 8B–E) with prefemoral process (PfP) considerably longer than acropodite (A), both quite complex and strongly armed; PfP broad, with four acute projections at its internal margin, two middle projections being more elongate, PfP ending in another elongated and acute apex directed laterobasally; A with internal margin bearing a process near middle, slightly broadened and acute, apex directed laterally, also wide and acute; solenomere (sl) simple and only slightly curved.</p><p>DISTRIBUTION. Brazil,Amazônas, São Paulo de Olivença, 3°27′23.2″S 68°48′01.3″W [Attems, 1898, 1938; Schubart, 1946]. Attems [1938] erroneously placed that locality in Pará state.</p><p>Hoffman [in litt.] recorded this species from  Padre Cocha and  Rio Yanayacu, both near Iquitos, Peru. To prove the identity, he also depicted the left gonopod of a ♂ (Fig. 8D, E).</p></div>	https://treatment.plazi.org/id/C46D87F6BB5503248530B756FD086E40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB54032484CCB66CFBB66D79.text	C46D87F6BB54032484CCB66CFBB66D79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha titana (Kraus 1956)	<div><p>Camptomorpha titana (Kraus, 1956)</p><p>Fig. 9.</p><p>Leptodesmus ( PHAntAsmOdesmus) titAnus  Kraus, 1956: 142, plate 18, figs 9–12. HolotYpe ♂, SMF; paratYpes, ♂ / ♀ and immatures, SMF.</p><p>CAmptOmOrpHA titAnA — Kraus, 1957: 98.</p><p>BRIEF DESCRIPTIVE NOTES (after Kraus [1957]). Length 47 mm, width 5.8 mm (♂), or 50 mm and 6.4 mm (♀), respectively. Coloration black-brown with a vague, narrow, axial stripe and paraterga white-yellowish, legs horn brown. Paraterga well-developed and strongly rimmed, both anterior and caudal corners mostly clearly rounded even on midbody rings, calluses set off from lateral rim (Fig. 9A). Epiproct short, simple and finger-shaped. HYpoproct with a small apical tooth between both thick setigerous knobs. ♂ legs ventrallY with prefemora carrYing apical swellings, and tibiae with apical chelae.</p><p>Gonopods (Fig. 9B–D) with prefemoral process (PfP) considerably longer than acropodite (A), both quite complex and strongly armed; PfP subrectangular, with one elongated projection in middle portion, ending in a bifid projection; apical portion of PfP bipartite, with two acute branches directed laterally; A wide and irregularly spoon-shaped, internal margin near apex with a small invagination subsequently forming a small subacute process; solenomere (sl) simple, subflagelliform and only slightly curved.</p><p>DISTRIBUTION. Peru, San Martin Dept., Divisoria, Cordillera Azul, divide between Ucayali and Huallaga valleys, 7°50′05.3″S 76°02′53.7″W, 1650 m; Sinchono, Cordillera Azul, 1500 m [Kraus, 1956, 1957].</p></div>	https://treatment.plazi.org/id/C46D87F6BB54032484CCB66CFBB66D79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB5403278767B511FDF96C40.text	C46D87F6BB5403278767B511FDF96C40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha tocachensis (Kraus 1955)	<div><p>Camptomorpha tocachensis (Kraus, 1955)</p><p>Fig. 10.</p><p>PrOletus tOcAcHensis 
Kraus, 1955: 186, plate 18, figs 25–28. HolotYpe ♂, MUSM.</p><p>Leptodesmus ( PHAntAsmOdesmus)  tOcAcHensis — Kraus, 1956: 142,  Camptomorpha by implication.</p><p>BRIEF DESCRIPTIVE NOTES (after Kraus [1955]). Length 45 mm, width 4.8 mm (♂). Coloration light red-brown with a light axial stripe; lateral halves of paraterga white-yellow both dorsally and ventrally, sides red-brown, sterna and legs light brown. Paraterga short and strongly rimmed, anterior corners strongly rounded, lateral rim slightly serrate, caudal corners mostly rounded and slightly drawn past straight rear tergal margins. Epiproct short, slightly attenuated and subtruncate. Hypoproct roundly triangular with two thick setigerous knobs near caudal margin. ♂ legs ventrallY with prefemora carrYing apical swellings, and tibiae with apical chelae (Fig. 10A).</p><p>Gonopods (Fig. 10B–D) with prefemoral process (PfP) much longer than acropodite (A), both highly complex and strongly armed; PfP with a broad and concave apex directed basally, with two processes: one longer and curved, bearing a small secondary branch, and a smaller one positioned more basally; A irregular in shape, resembling a wide lamina, with a well-developed process at internal medial margin; solenomere (sl) simple, subflagelliform and onlY slightlY curved.</p><p>DISTRIBUTION. Peru, Tocache, Rio Huallaga, 8°11′</p><p>24.1″S 76°30′33.9″W, 580 m [Kraus, 1955].</p></div>	https://treatment.plazi.org/id/C46D87F6BB5403278767B511FDF96C40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB57032784D7B45DFEDE6F92.text	C46D87F6BB57032784D7B45DFEDE6F92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha weyrauchi (Chamberlin 1955)	<div><p>Camptomorpha weyrauchi (Chamberlin, 1955)</p><p>Fig. 11.</p><p>Leptodesmus (Pseudoleptodesmus) weyrAucHi 
Chamberlin, 1955: 37, fig. 77. SYntYpes, SMF.</p><p>Leptodesmus ( PHAntAsmOdesmus)  weyrAucHi — Kraus, 1956: 142, plate 18, figs 13–16,  Camptomorpha by implication.</p><p>CAmptOmOrpHA weyrAucHi — Bouzan et al., 2022: 239.</p><p>BRIEF DESCRIPTIVE NOTES (after Chamberlin [1955] and Kraus [1956]). Length 45 mm, width 5.0– 5.3 mm (♂, ♀). Coloration dull red-brown to chocolate brown with contrasting yellow paraterga and a moderately wide, light, axial stripe; legs light brown. Paraterga well-developed and strongly rimmed, anterior corners stronglY rounded, calluses clearlY set off from a smooth lateral rim; caudal corners mostly rounded and only slightly drawn past straight rear tergal margins (Fig. 11A). Epiproct short, cylindrical, slightly curved ventrad. Hypoproct roundly triangular with two thick setigerous knobs at caudal margin. ♂ legs ventrallY with prefemora carrYing apical swellings, and tibiae with apical chelae.</p><p>Gonopods (Fig. 11B–F) with prefemoral process (PfP) considerably longer than acropodite (A), both highly complex and strongly armed; PfP subrectangular in shape, with an elongated projection in its middle portion, ending in a bifid projection; apical portion of PfP slightly bipartite, with two acute branches directed laterally; A broad, with an irregularly shaped apex featuring several indentations; internal margin near apex with a small invagination subsequently forming a small subacute process; solenomere (sl) simple, subflagelliform and only slightly curved.</p><p>DISTRIBUTION. Peru, Dept. Huanuco, Cave “Cueva de las Luchuzas” near Tingo Maria, Rio Huallaga, 9°17′55.0″S 76°00′01.6″W, 670 m; strict topotYpe ♂ [Chamberlin, 1955; Kraus, 1956].</p></div>	https://treatment.plazi.org/id/C46D87F6BB57032784D7B45DFEDE6F92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
C46D87F6BB570320878DB438FDCE698C.text	C46D87F6BB570320878DB438FDCE698C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Camptomorpha hoffmani	<div><p>Camptomorpha hoffmani sp.n.</p><p>Fig. 12.</p><p>HOLOTYPE ♂ (ZMUM), Peru, Junin Dept., Satipo Prov., near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.76639&amp;materialsCitation.latitude=-11.184444" title="Search Plazi for locations around (long -74.76639/lat -11.184444)">Rio Venado</a>, 1100–1400 m a.s.l., 11º11′41″S 74º46′40″W to 11º11′04″S 74º45′59″W, tropical forest, 13–20.XI.2016, I. Melnik leg.</p><p>NAME. GladlY dedicated to the late Richard L. Hoffman (1927–2012), a globally renowned specialist in the systematics of  Diplopoda and an amiable friend, whose unpublished documents have so extensively been used in the present review.</p><p>DIAGNOSIS. Differs from congeners primarilY bY the details of gonopodal structure (see also Key below and Fig. 12).</p><p>DESCRIPTION. Length ca 48 mm, width of midbody metazona 6.3 mm. General colouration in alcohol mostly brown to grey-brown, lateral rims of paraterga and their immediately adjacent parts yellowish to grey-yellow both dorsally and ventrally, legs largely red-brown, antennae and legs somewhat infuscate distally, venter a little lighter, gonopodal telopodites yellow (Fig. 12).</p><p>Head with a rather densely setose clypeolabral region, vertex nearly bare, epicranial suture evident; interantennal isthmus narrow, about half as wide as diameter of antennal socket (Fig. 12A, B). Antennae short and robust, only slightly clavate, in situ barely projecting past ring 2 dorsally; in length, antennomeres 2=6&gt;3–5&gt; 1&gt;7&gt;8.</p><p>Collum broadly rounded laterally and rather broadly rounded caudolaterally, narrowly rimmed. Collum and rigs 2–4 unusually compact and tightly following one after the other, vs remaining rings, these being more loose, as usual, and gradually attenuating towards telson. Dorsum moderately convex, especially anteriorly. Metaterga rather smooth and shining, prozona more dull, sides below paraterga arcuately striolate. Paraterga well-developed, on ring 19 smallest, mostly set at about upper 1/3 body height, clearly rimmed throughout, poriferous calluses (= peritremata) distinct, but not set off from lateral rim. Anterior corner of paraterga invariably broadly rounded, lateral rim smooth, caudal corners first obtuse and well rounded, on rings 4 and 5 subrectangular, thereafter increasingly acute, starting with ring 7 increasingly pointed and drawn past rear tergal margin. Tergal setae missing, setation pattern untraceable. Ozopores inside pits dorsolateral. Strictures between pro- and metazona deep lines. Limbus inconspicuous and entire. Pleurosternal carinae very small ridges on rings 2–5, poor swellings on rings 6 and 7, thereafter virtually missing. Spiracles small and inconspicuous. Epiproct very short, digitiform, subtruncate at apex and devoid of apical claws. Hypoproct subtriangular, with a short central tooth and 1+1 large setigerous knobs at caudal margin (Fig. 12A–H).</p><p>Sterna smooth, flat and bare, cross-impressions slight, sternal cones absent. Legs long and slender, ca 1.2–1.3 times as long as midbody height; coxae 3 each with a rather high, setose, straight, digitiform, apically rounded and well separated ventro-apical process (♂); all prefemora and tibiae but two last pairs with evident ventro-apical swellings and chelae (= sole pads subtending the tarsi basallY), respectivelY (♂). In length, femur (clearly curved ventrad)&gt; prefemur = tibia (with chela)&gt; postfemur = tarsus&gt; coxa &gt;&gt; claw.</p><p>Gonopodal aperture (Fig. 12H, I) transversely reniform, large, taking up most of ventral side of ♂ prozonum 7, not shifted onto metazonum 6, with prominent, lateral, rounded ridges and a rather small, central, caudal shelf inside. Gonopodal sternal elements fully membranous, devoid of sclerites (Fig. 12J). Gonopods complex (Fig. 12H–M). Coxites short, globose, much shorter than caudally curved telopodites, each with two distodorsal macrosetae; cannula as usual, slender and regularly curved. Telopodites tripartite, prefemorites as usual, short and very densely setose. Prefemoral process (PfP) bipartite, but button-shaped at an articulated apex and carrying two spines of different size, one (d) distal and significantlY more stronglY developed than a basal spine b (Fig. 12J–M). Acropodite (A) acuminate, with a considerable rounded lobe orally at midway. PfP much longer than A. Solenomere (sl) simple, rather regularly curved and acuminate.</p><p>♀ unknown.</p><p>DISTRIBUTION. Only the type locality.</p></div>	https://treatment.plazi.org/id/C46D87F6BB570320878DB438FDCE698C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Golovatch, Sergei I.;Bouzan, Rodrigo S.	Golovatch, Sergei I., Bouzan, Rodrigo S. (2025): Review of the South American millipede genus Camptomorpha Sivestri, 1897, with the description of a new species from Peru (Diplopoda: Polydesmida: Chelodesmidae). Arthropoda Selecta 34 (1): 1-15, DOI: 10.15298/arthsel.34.1.01
