identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
FABBEBBC9C46543FB64CAC4C9EA452F3.text	FABBEBBC9C46543FB64CAC4C9EA452F3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia arupii Pykala & Myllys 2025	<div><p>Protoblastenia arupii Pykälä &amp; Myllys sp. nov.</p><p>Fig. 2 A</p><p>Diagnosis.</p><p>Differs from P. dolomitica and P. westbergii in having only a slightly rimose thallus.</p><p>Type.</p><p>Finland • Enontekiön Lappi, Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.85&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.85/lat 69.03333)">Saana</a>, fell, steep NE-slope, dolomite rock outcrop, on NE-facing wall, 820 m a. s. l., 69°02'N, 20°51'E, 11 August 2011, J. Pykälä 44130 (H 9250904 – holotype, GenBank accession number: PV 766636) .</p><p>Description.</p><p>Thallus grey, slightly rimose, ca. 0.02–0.05 mm thick, K-, UV-. Apothecia orange, 0.2–0.5 mm, slightly convex to convex, 1 / 2 - immersed in thallus; ca. 60 apothecia / cm 2. Epihymenium brown-yellow to orange-brown, 12–15 μm thick, K + violet. Hymenium 50–60 μm thick. Hypothecium red-violet, ca. 50–60 μm thick. Paraphyses ca. 1.5–2 μm thick, apex not thickened. Ascospores 0 - septate, (10.3 –) 11.8–12.8 – 13.8 (– 15.1) × (5.3 –) 5.7–6.8 – 7.8 (– 9.4) μm (n = 27).</p><p>Habitat and distribution.</p><p>Only one specimen has been found on the calcareous Saana fell in NW Finland. The species was associated with Polyblastia sp. and Rhizocarpon petraeum (Wulfen) A. Massal.</p><p>Etymology.</p><p>The specific epithet is in honour of Dr Ulf Arup (Lund) for his many important contributions on the taxonomy (particularly Teloschistaceae and Lecanoraceae) and biogeography of especially Fennoscandian lichens.</p><p>Notes.</p><p>Protoblastenia arupii is related to P. dolomitica and P. westbergii in the ITS tree, although the relationship remains unsupported. The only known specimen is characterised by a thin red-violet hypothecium and large spores, some of them particularly broad. Protoblastenia dolomitica and P. westbergii differ in their rimose to areolate thallus and slightly narrower spores. An ITS marker as a DNA barcode may be needed for unambiguous identification.</p></div>	https://treatment.plazi.org/id/FABBEBBC9C46543FB64CAC4C9EA452F3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
9C8CA1D62BD650B2A2EE4956005B2AA6.text	9C8CA1D62BD650B2A2EE4956005B2AA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia borealis Pykala & Myllys 2025	<div><p>Protoblastenia borealis Pykälä &amp; Myllys sp. nov.</p><p>Fig. 2 B</p><p>Diagnosis.</p><p>Differs from P. rupestris in having a mostly rimose thallus.</p><p>Type.</p><p>Finland • Varsinais-Suomi, Lohja, Lylyinen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.95&amp;materialsCitation.latitude=60.25" title="Search Plazi for locations around (long 23.95/lat 60.25)">Sääksniemi</a>, N of summer cottages, small calcareous rock outcrop, on SW-slope, 50 m a. s. l., 60°15'N, 23°57'E, 16 September 2003, J. Pykälä 23710 (H 9250905 – holotype, UPS – isotype, GenBank accession number: PV 766638) .</p><p>Description.</p><p>Thallus white to pale grey, endolithic (rarely), rimose (usually) to areolate, areoles 0.1–0.8 mm, ca. 0.02–0.2 mm thick, K-, C-, UV- to UV + bluish-white, algal cells 5–12 (– 17) μm. Apothecia yellow-orange to dirty brown-orange, 0.2–1.0 mm, slightly convex to strongly convex, ½ - immersed in thallus to superficial, occasionally leaving shallow pits, K + violet; ca. 30–140 apothecia / cm 2. Epihymenium orange-brown, 10–18 μm thick, K + violet. Hymenium 50–90 (– 100) μm thick. Hypothecium colourless to pale yellow, rarely lower part reddish-brown, occasionally with oil droplets, ca. 40–160 μm thick. Paraphyses ca. (1.5 –) 2–2.5 μm thick, apex not or slightly thickened, sparingly branched and anastomosing. Ascospores 0 - septate, (7.6 –) 9.6–11.2 – 12.7 (– 16.1) × (4.8 –) 5.4–6.2 – 7.0 (– 7.7) μm (n = 146).</p><p>Habitat and distribution.</p><p>This species is the most common species of Protoblastenia in Finland. It occurs from the hemi-boreal zone to middle Lapland. It grows on calcareous and calciferous rocks on both sun-exposed and shady habitats, but may prefer the latter. It also often grows in lime quarries. Specimens from concrete probably also belong in this species, but no sequences of them are available. Typical companion species include, for example, Acarospora glaucocarpa (Ach.) Körb., Clauzadea monticola (Schaer.) Hafellner &amp; Bellem., Farnoldia jurana (Schaer.) Hertel, Lathagrium fuscovirens (With.) Otálora, P. M. Jørg. &amp; Wedin, Phaeophyscia sciastra (Ach.) Moberg, Scytinium parvum (Degel.) Otálora, P. M. Jørg. &amp; Wedin, Verrucaria deversa Vain., V. epilithea Vain., V. muralis Ach. and V. nigrescens Pers.</p><p>Etymology.</p><p>The name refers to the geographic distribution of the species in the boreal vegetation zone.</p><p>Notes.</p><p>Protoblastenia borealis is morphologically very variable and may differ in the size and density of apothecia and in the thickness of the hypothecium. The sister species Protoblastenia ekmanii is morphologically similar, but appears to differ in having a thicker hypothecium. The new species is also difficult to distinguish from the closely-related P. rupestris . However, P. rupestris does not have any strongly convex apothecia and has, on average, a thicker thallus, which is mostly areolate. Protoblastenia pseudocompressa often / usually has plane apothecia, but some specimens with convex apothecia are somewhat similar to P. borealis . The two Finnish specimens of Protoblastenia calvella resemble P. borealis in morphology; they have strongly convex apothecia and a thick hypothecium which were also observed in a few P. borealis specimens. Due to these overlaps, ITS is needed for unambiguous identification of P. borealis .</p><p>Other sequenced specimens.</p><p>Finland • Varsinais-Suomi, Karkkila, Haavisto, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.333334&amp;materialsCitation.latitude=60.483334" title="Search Plazi for locations around (long 24.333334/lat 60.483334)">Herneojankallio</a>, pyroxene gneiss rock outcrop, on high SW-facing overhanging wall, scarce, 80 m a. s. l., 60°29'N, 24°20'E, 18. 5. 1996, J. Pykälä 16567 (H) ; • Lohja, Torhola, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.866667&amp;materialsCitation.latitude=60.25" title="Search Plazi for locations around (long 23.866667/lat 60.25)">300 m N-NE of Kallioranta</a>, young pine plantation, S-slope, almost 1 m high rock wall of calcareous rock outcrop, rather scarce, 60 m a. s. l., 60°15'N, 23°52'E, 22. 7. 2004, J. Pykälä 25292 (H) ; • Karjalohja, Karkali, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.816668&amp;materialsCitation.latitude=60.233334" title="Search Plazi for locations around (long 23.816668/lat 60.233334)">200 m SW Karkali farm, on a shore of Lohjanjärvi Lake</a>, small calcareous rock outcrop, on stone, 33 m a. s. l., 60°14'N, 23°49'E, 19. 10. 2004, J. Pykälä 26097 (H) ; • Pohja, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.4&amp;materialsCitation.latitude=60.133335" title="Search Plazi for locations around (long 23.4/lat 60.133335)">Kuovila</a>, 300 m SW of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.4&amp;materialsCitation.latitude=60.133335" title="Search Plazi for locations around (long 23.4/lat 60.133335)">Vihreämäki</a>, abandoned lime quarry, NE-facing wall, 37 m a. s. l., 60°08'N, 23°24'E, 12. 10. 2006, J. Pykälä 29704 (H) ; • Suomusjärvi, Sallittu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.616667&amp;materialsCitation.latitude=60.3" title="Search Plazi for locations around (long 23.616667/lat 60.3)">Huuttavanmäki</a>, calciferous rock outcrop, on W-facing wall, 110 m a. s. l., 60°18'N, 23°37'E, 28. 6. 2008, J. Pykälä 32757 (H) ; • Särkisalo, Förby, E of Vähämaankaula, abandoned lime quarry, on N-facing wall, 7 m a. s. l., 23. 7. 2008, J. Pykälä 33037 (H) ; • Särkisalo, Förby, E of Vähämaankaula, abandoned lime quarry, on NW-facing wall, 7 m a. s. l, 23. 7. 2008, J. Pykälä 33101 (H) ; • Kemiönsaari (Västanfjärd), Finsjö, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.683332&amp;materialsCitation.latitude=60.0" title="Search Plazi for locations around (long 22.683332/lat 60.0)">Verkviksudden</a>, calcareous rock outcrop, on 1.2 m high E-facing wall, 5 m a. s. l., 60°00'N, 22°41'E, 13. 7. 2010, J. Pykälä 38784 (H) ; • Salo (Suomusjärvi), Hinttala, W of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.65&amp;materialsCitation.latitude=60.383335" title="Search Plazi for locations around (long 23.65/lat 60.383335)">Kalattomansuo</a>, Picea abies - dominated herb-rich forest, abandoned lime quarry, on S-facing wall, 110 m a. s. l, 60°23'N, 23°39'E, 28. 9. 2010, J. Pykälä 41355 (H) ; • Etelä-Savo, Kerimäki, Ruokojärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.0&amp;materialsCitation.latitude=61.933334" title="Search Plazi for locations around (long 29.0/lat 61.933334)">Pitkäniemi</a>, close by abandoned lime quarry, young Pinus sylvestris - forest, on calcareous boulder, 90 m a. s. l., 61°56'N, 29°00'E, 15. 9. 2011, J. Pykälä 45759 (H) ; • Pohjois-Karjala, Juankoski, Siikajärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=28.35&amp;materialsCitation.latitude=63.2" title="Search Plazi for locations around (long 28.35/lat 63.2)">Huosiaisniemi, Nature Reserve</a>, mixed herb-rich forest, dolomite rock outcrop, on W-facing wall, scarce, 100 m a. s. l., 63°12'N, 28°21'E, 25. 7. 2011, J. Pykälä 42802 (H) ; • Juankoski, Siikajärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=28.6&amp;materialsCitation.latitude=63.15" title="Search Plazi for locations around (long 28.6/lat 63.15)">Lauantaijoki, Kalliola 250 m N</a>, calciferous serpentine rock outcrop, on NW-facing wall, 132 m a. s. l., 63°09'N, 28°36'E, 9. 9. 2013, J. Pykälä 47217 (H) ; • Keski-Pohjanmaa, Vimpeli, Sääksjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.066668&amp;materialsCitation.latitude=63.15" title="Search Plazi for locations around (long 24.066668/lat 63.15)">Huosianmaankallio</a>, lime quarry, quarry spoil, on calcareous boulder, rather scarce, 135 m a. s. l., 63°09'N, 24°04'E, 1. 9. 2010, J. Pykälä 40695 (H) ; • Kainuu, Hyrynsalmi, Moisiovaara, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.083334&amp;materialsCitation.latitude=64.53333" title="Search Plazi for locations around (long 29.083334/lat 64.53333)">Hyyryläinen SW</a>, Pinus sylvestris - dominated heath forest, VMT site type, calciferous serpentine rock outcrop, on 1 m high W-facing wall, 190 m a. s. l., 64°32'N, 29°05'E, 30. 5. 2021, J. Pykälä 57501 (H) ; • Koillismaa, Kuusamo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.316668&amp;materialsCitation.latitude=66.36667" title="Search Plazi for locations around (long 29.316668/lat 66.36667)">Oulanka National Park, Kiutaköngäs 400 m N</a>, Pinus sylvestris - herb-rich forest, small dolomite rock outcrop, on SE-slope, rather scarce, 175 m a. s. l., 66°22'N, 29°19'E, 3. 8. 2010, J. Pykälä 39151 ; • Kuusamo, Juuma, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.45&amp;materialsCitation.latitude=66.25" title="Search Plazi for locations around (long 29.45/lat 66.25)">Oulanka National Park, Hautaniitynvuoma</a>, gorge, calciferous (dolomite) schistose rock outcrop, on SW-facing wall, 195 m a. s. l, 66°15'N, 29°27'E, 14. 8. 2010, J. Pykälä 39942 (H) ; • Kuusamo, Juuma, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.433332&amp;materialsCitation.latitude=66.25" title="Search Plazi for locations around (long 29.433332/lat 66.25)">Oulanka National Park, Hautaniitynvuoma</a>, gorge, NE-slope, calciferous (dolomite) schistose rock outcrop, on NE-facing wall, 179 m a. s. l., 66°15'N, 29°26'E, 21. 8. 2011, J. Pykälä 44641 (H) ; • Salla, Oulanka National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.183332&amp;materialsCitation.latitude=66.433334" title="Search Plazi for locations around (long 29.183332/lat 66.433334)">Savilampi 1.4 km NE, bank of Savinajoki River</a>, cliff, dolomite rock outcrop, on SE-facing wall, 185 m a. s. l., 66°26'N, 29°11'E, 23. 8. 2011, J. Pykälä 44906 (H) ; • Kuusamo, Kallunki, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.05&amp;materialsCitation.latitude=66.36667" title="Search Plazi for locations around (long 29.05/lat 66.36667)">Juhonlampi</a>, young Pinus sylvestris - dominated heath forest, calcareous rock outcrop, SW-slope, on 80 cm high SW-facing wall, 236 m a. s. l., 66°22'N, 29°03'E, 5. 8. 2021, J. Pykälä 58196 (H) ; • Kuusamo, Vasaraperä, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=28.733334&amp;materialsCitation.latitude=66.1" title="Search Plazi for locations around (long 28.733334/lat 66.1)">Havukkalampi N</a>, mixed herb-rich heath forest, calcareous rock outcrop, on 100 cm high SW-facing wall, 246 m a. s. l., 66°06'N, 28°44'E, 8. 8. 2021, J. Pykälä 58392 (H) ; • Kittilän Lappi, Kittilä, Tepsa, Herravaara SE, clear-cut heath forest, small calcareous rock outcrop, on 1.5 m high SE-facing wall, scattered, 15. 9. 2022 J. Pykälä &amp; I. Kuusisto 60511 (H) .</p></div>	https://treatment.plazi.org/id/9C8CA1D62BD650B2A2EE4956005B2AA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
F2FB6207636D5608A655A6F95C6B5CAC.text	F2FB6207636D5608A655A6F95C6B5CAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia calvella Kainz & Rambold	<div><p>Protoblastenia calvella Kainz &amp; Rambold, Bibl. Lichenol. 88: 290 (2004)</p><p>Description.</p><p>Thallus white to whitish-grey, rimose to areolate, areoles 0.2–0.5 mm, ca. 0.02–0.2 mm thick, K-, C-, UV-, algal cells 5–8 μm. Apothecia orange-yellow to orange, 0.3–0.6 mm, strongly convex, superficial, not leaving pits to leaving shallow pits, K + violet; ca. 40–100 apothecia / cm 2. Epihymenium dark orange-brown, 12–15 μm thick, K + violet. Hymenium 80–100 μm thick. Hypothecium colourless to pale yellow, ca. 100–180 μm thick. Paraphyses ca. 2–2.5 μm thick, apex slightly thickened, often branching. Ascospores 0 - septate, (7.6 –) 9.4–10.7 – 12.0 (– 12.7) × (4.4 –) 5.2–5.9 – 6.7 (– 7.8) μm (n = 29).</p><p>Habitat and distribution.</p><p>Only two specimens are known from northern Finland (Pykälä 2023). The habitats are sun-exposed calcareous and calciferous rock outcrops where the species grows on rock or on pebbles. The species is associated with Enchylium polycarpon (Hoffm.) Otálora, P. M. Jørg. &amp; Wedin and Rhizocarpon petraeum .</p><p>Notes.</p><p>In the ITS phylogeny, the P. calvella specimens are divided into three strongly-supported lineages: the first one includes three specimens collected in Austria, Germany and Sweden; the second group includes two Finnish specimens and the third group consists of specimens collected in Croatia and Germany (Fig. 1). More material is needed to determine whether P. calvella represents several species. Protoblastenia calvella is a cryptic species which cannot be identified without molecular data (Wirth et al. 2013). Older apothecia are often brown-orange to dark red-brown (Svensson et al. 2024), but such colouration was not observed in the Finnish specimens. It may be confused with several other Protoblastenia species. A few specimens of P. borealis have a thick hypothecium and / or hymenium and thus resemble the Finnish specimens of P. calvella . The Finnish specimens of P. rupestris have a thinner hymenium and hypothecium compared to P. calvella . Furthermore, the apothecia tend to be predominantly orange, while in P. calvella, they are orange yellow. Protoblastenia ekmanii may differ in having darker orange apothecia. Protoblastenia fennoarctica can be rather similar to P. calvella, but its apothecia vary from plane to strongly convex and its hymenium is thinner (ca. 50–80 μm thick).</p><p>Specimens examined.</p><p>• Enontekiön Lappi, Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.5&amp;materialsCitation.latitude=69.183334" title="Search Plazi for locations around (long 21.5/lat 69.183334)">Toskalharji, Toskalpahta</a>, fell, dolomite rock outcrop, on steep SW-slope, 820 m a. s. l., 69°11'N, 21°30'E, 4. 8. 2011, J. Pykälä 43503 (H) ; • Koillismaa, Kuusamo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.4&amp;materialsCitation.latitude=66.26667" title="Search Plazi for locations around (long 29.4/lat 66.26667)">Juuma, Niskakoski, river bank</a>, calciferous (dolomite) schistose rock outcrop, on pebbles, 224 m a. s. l., 66°16'N, 29°24'E, 22. 8. 2011, J. Pykälä 44807 (H) .</p></div>	https://treatment.plazi.org/id/F2FB6207636D5608A655A6F95C6B5CAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
05B49C280BF1579BB75280480A65D4C3.text	05B49C280BF1579BB75280480A65D4C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia compressa Pykala & Myllys 2025	<div><p>Protoblastenia compressa Pykälä &amp; Myllys sp. nov.</p><p>Fig. 2 C</p><p>Diagnosis.</p><p>Differs from P. pseudocompressa in having plane to slightly convex apothecia.</p><p>Type.</p><p>Finland • Varsinais-Suomi, Parainen (Korppoo), Elfsjö, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.4&amp;materialsCitation.latitude=60.133335" title="Search Plazi for locations around (long 21.4/lat 60.133335)">Hummelskär Island, close by shore of the Baltic Sea</a>, calcareous rock outcrop, on shady 1 m high NE-facing wall, 5 m a. s. l., 60°08'N, 21°24'E, 31 July 2009, J. Pykälä 35585 (H 9250940 – holotype, GenBank accession number: PV 766659) .</p><p>Description.</p><p>Thallus white to pale grey, continuous, rimose to rarely areolate, areoles 0.2–0.8 mm, ca. 0.01–0.15 mm thick, K-, C-, UV- to UV + bluish-white, algal cells 4–9 μm. Apothecia yellow, yellow-orange to orange, 0.2–0.8 mm, plane to slightly convex, 1 / 4–1 - immersed in thallus, occasionally leaving shallow pits, K + violet; ca. 20–190 apothecia / cm 2. Epihymenium orange-brown, 12–18 μm thick, K + violet. Hymenium 50–60 μm thick. Hypothecium colourless to yellow, ca. 40–100 μm thick. Paraphyses ca. 2–2.5 μm thick, apex not thickened to slightly thickened, sparingly branched and anastomosing. Ascospores 0 - septate, (8.6 –) 9.8–11.4 – 12.6 (– 13.5) × (4.8 –) 5.1–5.9 – 6.7 (– 7.4) μm (n = 45).</p><p>Habitat and distribution.</p><p>This species has ca. 40 known localities in the hemi-boreal vegetation zone in SW Finland (seven localities confirmed by ITS). It grows on calcareous rocks and in lime quarries, often on N-facing walls. It seems to prefer shady habitats. In the SW archipelago of Finland, P. compressa may be the most common Protoblastenia species. Typical companion species are Bagliettoa baldensis (A. Massal.) Vězda s. lat, Clauzadea monticola, Lepraria finkii (B. de Lesd.) R. C. Harris, Polyblastia abscondita (Nyl.) Arnold, Sagiolechia protuberans (Ach.) A. Massal., Thelidium decipiens (Nyl.) Kremp., T. incavatum Mudd, Verrucaria foveolata (Flörke) A. Massal., V. muralis, V. nigrescens s. lat. and V. viridula (Schrad.) Ach.</p><p>Etymology.</p><p>The name refers to the flattened apothecia of the species.</p><p>Notes.</p><p>The species is characterised by plane to slightly convex apothecia. Protoblastenia pseudocompressa is a sister species with a rather similar morphology. However, P. pseudocompressa often has convex apothecia which have not been seen in P. compressa . The occurrence of Protoblastenia pseudocompressa has been confirmed only in the Kuusamo-Salla area ca. 800 kilometres north from the distribution area of P. compressa . Protoblastenia incrustans differs in having apothecia that leave pits ( P. compressa rarely leaves shallow pits). Protoblastenia coniasis differs in having a yellow thallus that gradually turns brownish-rose in the herbarium (Poelt 1957). Protoblastenia compressa may also be difficult to separate from P. rupestris, even though they are not closely related. However, P. compressa usually has thinner apothecia and a thinner thallus which is rarely areolate (mostly areolate in P. rupestris). ITS may sometimes be needed for unambiguous identification of P. compressa .</p><p>Other sequenced specimens.</p><p>Finland • Varsinais-Suomi, Kemiönsaari (Västanfjärd), Södersundvik, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.766666&amp;materialsCitation.latitude=60.05" title="Search Plazi for locations around (long 22.766666/lat 60.05)">Näsudden</a>, calcareous rock outcrop on shore of the Baltic Sea, cliff, on N-facing wall, rather scarce, 12 m a. s. l., 60°03'N, 22°46'E, 25. 8. 2009, J. Pykälä 36521 (H) ; • Parainen (Korppoo), Elfsjö, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.433332&amp;materialsCitation.latitude=60.15" title="Search Plazi for locations around (long 21.433332/lat 60.15)">Stora Limskär Island</a>, abandoned lime quarry, beneath N-facing wall, bottom, on pebbles, 4 m a. s. l., 60°09'N, 21°26'E, 22. 6. 2010, J. Pykälä 38349 (H) ; • Parainen (Korppoo), Åfvensår, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.533333&amp;materialsCitation.latitude=60.3" title="Search Plazi for locations around (long 21.533333/lat 60.3)">Ronudden</a>, abandoned lime quarry, on NW-facing wall, 2 m a. s. l., 60°18'N, 21°32'E, 2. 7. 2010, J. Pykälä 38646 (H) ; • Parainen (Korppoo), Åfvensår, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.533333&amp;materialsCitation.latitude=60.283333" title="Search Plazi for locations around (long 21.533333/lat 60.283333)">Ronudden</a>, abandoned lime quarry on shore of the Baltic Sea, on overhanging NW-facing wall, 2 m a. s. l., 60°17'N, 21°32'E, 15. 7. 2010, J. Pykälä 38965 (H) ; • Salo (Suomusjärvi), Rautsuo, 200 m NE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.616667&amp;materialsCitation.latitude=60.35" title="Search Plazi for locations around (long 23.616667/lat 60.35)">Kukutin</a>, herb-rich forest, abandoned lime quarry, on NW-facing wall, 85 m a. s. l., 60°21'N, 23°37'E, 28. 9. 2010, J. Pykälä 41376 (H) ; • Parainen, Skräbböle, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.283333&amp;materialsCitation.latitude=60.283333" title="Search Plazi for locations around (long 22.283333/lat 60.283333)">Skräbböle</a> lime quarry, by the lime quarry, N-slope, on stone, 24 m a. s. l., 60°17'N, 22°17'E, 5. 10. 2010, J. Pykälä 41511 b (H) ; • Parainen, Attu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.3&amp;materialsCitation.latitude=60.183334" title="Search Plazi for locations around (long 22.3/lat 60.183334)">Skötudden SW</a>, abandoned lime quarry on shore of the Baltic Sea, on NE-facing wall, 2 m a. s. l., 60°11'N, 22°18'E, 16. 6. 2011, J. Pykälä 42124 (H) .</p></div>	https://treatment.plazi.org/id/05B49C280BF1579BB75280480A65D4C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
B5E2DF0491C05E7B94AC65B8593D8E2E.text	B5E2DF0491C05E7B94AC65B8593D8E2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia dolomitica Pykala & Myllys 2025	<div><p>Protoblastenia dolomitica Pykälä &amp; Myllys sp. nov.</p><p>Fig. 2 D</p><p>Diagnosis.</p><p>Differs from P. siebenhaariana in having more densely occurring apothecia and from P. arupii in having a rimose to areolate thallus and often olive-yellow apothecia.</p><p>Type.</p><p>Finland • Enontekiön Lappi, Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.433332&amp;materialsCitation.latitude=69.183334" title="Search Plazi for locations around (long 21.433332/lat 69.183334)">Toskalharji, Toskaljärvi N</a>, fell, brook, W-shore, dolomite rock outcrop, on N-slope, 710 m a. s. l., 69°11'N, 21°26'E, 3 August 2011, J. Pykälä 43466 (H 9235018 – holotype, GenBank accession number: PV 766671) .</p><p>Description.</p><p>Thallus white to whitish-grey, rarely pale brown, rimose to areolate, areoles 0.2–0.9 mm, ca. 0.03–0.25 mm thick, K-, C-, UV-, one specimen UV + bluish-white, algal cells 4–8 (– 12) μm. Apothecia yellow to dirty orange-yellow, sometimes olive-yellow, 0.15–0.7 mm, plane, slightly convex to convex, 1 / 2 - immersed in thallus; ca. 40–160 apothecia / cm 2. Epihymenium dirty olive-yellow to orange-brown, 12–20 μm thick, K + violet. Hymenium 50–70 μm thick. Hypothecium dark brown, reddish-brown or usually violet, ca. 40–100 μm thick. Paraphyses ca. 1.5–2 μm thick, apex not thickened to slightly thickened, branching. Ascospores 0 - septate, (7.6 –) 11.0–12.8 – 14.7 (– 19.4) × (4.4 –) 4.8–5.4 – 5.9 (– 7.4) μm (n = 117).</p><p>Habitat and distribution.</p><p>This species has been found only in the parishes of Salla (in the biogeographical Province Ks) and Enontekiö (biogeographical Province EnL) in N Finland. It grows on dolomite rock outcrops, once collected from dolomite stones and pebbles. The species may prefer sun-exposed and half-shady sites on fells and in gorges. Companion species include, for example, Acarospora glaucocarpa, Hymenelia rhodopis (Sommerf.) Lutzoni, Opegrapha dolomitica (Arnold) Clauzade &amp; Cl. Roux ex Torrente &amp; Egea, Polyblastia spp., Protoblastenia fennoarctica, Synalissa ramulosa (Bernh.) Körb., Thelidium auruntii (A. Massal.) Kremp. and T. declivum Pykälä &amp; Myllys.</p><p>Etymology.</p><p>The species grows on dolomite rocks.</p><p>Notes.</p><p>Protoblastenia dolomitica is closely related to P. westbergii, which has slightly larger apothecia, broader spores and sometimes a thicker hypothecium. The specimens have been previously identified as P. siebenhaariana in Finland. However, this species has only 1–10 apothecia / cm 2 (Kainz and Rambold 2004) and the Finnish specimens are not closely related to a GenBank specimen of P. siebenhaariana . Protoblastenia dolomitica is characterised by rather small, often dirty olive-yellow apothecia and a thin hypothecium. Protoblastenia arupii, based on the inspection of only one known specimen, has orange apothecia, on average, broader spores and an only slightly thinner rimose thallus. More material from the morphologically related species is needed to determine whether P. dolomitica can be identified by morphology only.</p><p>Other specimens examined.</p><p>Finland • Koillismaa, Salla, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.15&amp;materialsCitation.latitude=66.416664" title="Search Plazi for locations around (long 29.15/lat 66.416664)">Oulanka National Park, Pikkuköngäs, bank of River Oulankajoki</a>, high cliff, calciferous (dolomite) schistose rock, on overhanging SW-facing wall, 180 m a. s. l., 66°25'N, 29°09'E, 10. 8. 2009, J. Pykälä 36072 (H) ; • Salla, Oulanka National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.133333&amp;materialsCitation.latitude=66.416664" title="Search Plazi for locations around (long 29.133333/lat 66.416664)">Pikkuköngäs, bank of River Oulankajoki</a>, high cliff, calciferous (dolomite) schistose rock outcrop, on SE-facing wall, 182 m a. s. l., 66°25'N, 29°08'E, 4. 8. 2010, J. Pykälä 39217 (H) ; • Salla, Oulanka National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.133333&amp;materialsCitation.latitude=66.416664" title="Search Plazi for locations around (long 29.133333/lat 66.416664)">Pikkuköngäs, bank of River Oulankajoki</a>, high cliff, calciferous (dolomite) schistose rock outcrop, beneath SW-facing wall, on pebble, 180 m a. s. l., 66°25'N, 29°08'E, 13. 8. 2010, J. Pykälä 39851 (H) ; • Enontekiön Lappi, Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.433332&amp;materialsCitation.latitude=69.2" title="Search Plazi for locations around (long 21.433332/lat 69.2)">Toskalharji, Toskaljärvi N</a>, fell, dolomite rock outcrop, gentle SE-slope, on dolomite pebbles, 730 m a. s. l., 69°12'N, 21°26'E, 2. 8. 2011, J. Pykälä 43340 (H) ; • Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.433332&amp;materialsCitation.latitude=69.183334" title="Search Plazi for locations around (long 21.433332/lat 69.183334)">Toskalharji, Toskaljärvi N</a>, lake shore, dolomite scree, on dolomite stones, 705 m a. s. l., 69°11'N, 21°26'E, 7. 8. 2011, J. Pykälä 43855 (H) ; • Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.85&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.85/lat 69.03333)">Saana</a>, fell, steep NE-slope, dolomite rock outcrop, on NE-facing wall, 820 m a. s. l., 69°02'N, 20°51'E, 11. 8. 2011, J. Pykälä 44115 (H) .</p></div>	https://treatment.plazi.org/id/B5E2DF0491C05E7B94AC65B8593D8E2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
D403566327F65D71B1AEEBE6C97A9B99.text	D403566327F65D71B1AEEBE6C97A9B99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia ekmanii Pykala & Myllys 2025	<div><p>Protoblastenia ekmanii Pykälä &amp; Myllys sp. nov.</p><p>Fig. 3 A</p><p>Diagnosis.</p><p>Differs from P. borealis in having a thicker hypothecium.</p><p>Type.</p><p>Finland • Varsinais-Suomi, Salo (Kisko), Kirkonkylä, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.516666&amp;materialsCitation.latitude=60.2" title="Search Plazi for locations around (long 23.516666/lat 60.2)">450 m NW of Kolko, shore of Määrjärvi Lake</a>, calcareous rock outcrop, on W-slope, 43 m a. s. l., 60°12'N, 23°31'E, 20 May 2010, J. Pykälä 37508 (H 9224071 – holotype, GenBank accession number: PV 766674) .</p><p>Description.</p><p>Thallus white to greyish-white, rimose to areolate, areoles 0.3–0.6 mm, ca. 0.05–0.2 mm thick, K-, UV + bluish-white, algal cells 5–8 μm. Apothecia dirty brown-orange, 0.4–0.8 mm, convex to strongly convex, superficial; ca. 30–50 apothecia / cm 2. Epihymenium dark orange-brown, 10–20 μm thick, K + violet. Hymenium 50–70 μm thick. Hypothecium yellowish-white, ca. 120–200 μm thick, in some apothecia base patchily red-violet. Paraphyses ca. 2–2.5 μm thick, apex not or slightly thickened. Ascospores 0 - septate, (8.1 –) 8.7–10.2 – 11.6 (– 12.5) × (5.0 –) 5.1–5.9 – 6.6 (– 7.2) (n = 12).</p><p>Habitat and distribution.</p><p>Only one specimen is known from a calcareous rock outcrop on a lake shore in SW Finland. The sun-exposed site suggests that the species may have a southern distribution. The species is apparently very rare in Finland. Companion species include Acarospora glaucocarpa, Clauzadea monticola, Placynthium nigrum (Huds.) Gray and Thelidium incavatum .</p><p>Etymology.</p><p>The name is in honour of Dr Stefan Ekman (Uppsala) for his many important contributions on the phylogeny and taxonomy of lichens in Fennoscandia and elsewhere.</p><p>Notes.</p><p>The species is closely related to P. borealis (only 2.5 % difference in ITS), but seems to differ in having a thicker hypothecium. The hypothecium of P. borealis is very variable, but in no specimens does it reach the upper limit of the thickness in the P. ekmanii specimen. The Finnish specimens of P. calvella have orange-yellow to orange apothecia and thus resemble P. ekmanii . More material is needed to determine whether the species can be unambiguously morphologically distinguished from P. calvella .</p></div>	https://treatment.plazi.org/id/D403566327F65D71B1AEEBE6C97A9B99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
EC602A5E49655BDD8DC0FBF1BFE0AD57.text	EC602A5E49655BDD8DC0FBF1BFE0AD57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia fennoarctica Pykala & Myllys 2025	<div><p>Protoblastenia fennoarctica Pykälä &amp; Myllys sp. nov.</p><p>Fig. 3 B</p><p>Diagnosis.</p><p>Differs from P. calvella in having thinner hymenium, from P. minuta in having rimose to areolate thallus and from P. rikkinenii in usually having more densely occurring apothecia.</p><p>Type.</p><p>Finland • Enontekiön Lappi, Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.466667&amp;materialsCitation.latitude=69.2" title="Search Plazi for locations around (long 21.466667/lat 69.2)">Toskalharji, 1.2 km NE of Toskaljärvi</a>, fell, SW-slope, scree, on dolomite boulder, 875 m a. s. l., 69°12'N, 21°28'E, 5 August 2011, J. Pykälä 43656 (H 9220129 – holotype, GenBank accession number: PV 766677) .</p><p>Description.</p><p>Thallus white to pale greyish-brown, rimose to areolate, areoles 0.15–1.0 mm, ca. 0.03–0.25 mm thick, K-, C-, UV-, algal cells 5–10 μm. Apothecia yellow, orange-yellow to orange, 0.3–0.9 mm, plane to strongly convex, immersed in thallus to superficial; ca. 20–110 apothecia / cm 2. Epihymenium orange-yellow to brown, 12–20 μm thick, K + violet. Hymenium 50–80 μm thick. Hypothecium colourless to yellow, base sometimes patchily reddish-brown, ca. 60–280 μm thick. Paraphyses ca. 2–3 μm thick, cells 5–8 μm long, apex not thickened to slightly thickened, sparingly branched and anastomosing. Ascospores 0 - septate, (6.6 –) 8.7–10.0 – 11.3 (– 12.5) × (4.2 –) 4.8–5.3 – 5.9 (– 7.2) μm (n = 93).</p><p>Habitat and distribution.</p><p>This species has been found only in the Parish Enontekiö (biogeographical Province EnL in NW Finland). It grows on fells on dolomite rock outcrops, boulders and pebbles. Companion species include Enchylium polycarpon, Farnoldia jurana, Physcia caesia (Hoffm.) Fürnr., Polyblastia spp., Protoblastenia dolomitica and Verrucaria vacillans Pykälä &amp; Myllys.</p><p>Etymology.</p><p>The species occurs in the most northern, almost arctic (oroarctic) sites in Finland (Fennia).</p><p>Notes.</p><p>Protoblastenia fennoarctica morphologically resembles P. calvella, P. minuta, P. remota and P. rikkinenii in having a pale hypothecium and small spores. In the ITS phylogeny, the species is most closely related to P. rikkinenii (Fig. 1). Protoblastenia rikkinenii differs in having more sparsely occurring apothecia, but more material is needed to confirm whether the two species can be identified by morphology. Protoblastenia calvella has a thicker hymenium (80–100 μm thick). Protoblastenia minuta and P. remota have an endolithic to thin thallus and P. minuta has smaller spores compared to P. fennoarctica, while P. remota may have slightly larger spores.</p><p>Other specimens examined.</p><p>Finland • Enontekiön Lappi, Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.483334&amp;materialsCitation.latitude=69.183334" title="Search Plazi for locations around (long 21.483334/lat 69.183334)">Toskalharji, Toskalpahta</a> fell, SW-slope, scree, on dolomite pebbles, 795 m a. s. l., 69°11'N, 21°29'E, 1. 8. 2011, J. Pykälä 43105 (H) ; • Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.433332&amp;materialsCitation.latitude=69.183334" title="Search Plazi for locations around (long 21.433332/lat 69.183334)">Toskalharji, Toskaljärvi N</a>, fell, dolomite scree, gentle SE-slope, on dolomite boulder, 715 m a. s. l., 69°11'N, 21°26'E, 2. 8. 2011, J. Pykälä 43194 (H) ; • Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.85&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.85/lat 69.03333)">Saana</a>, fell, steep NE-slope, dolomite rock outcrop, on NE-facing wall, 820 m a. s. l., 69°02'N, 20°51'E, 11. 8. 2011, J. Pykälä 44116 (H) .</p></div>	https://treatment.plazi.org/id/EC602A5E49655BDD8DC0FBF1BFE0AD57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
26F7C1B6407055EBA01BCB72E0DFB756.text	26F7C1B6407055EBA01BCB72E0DFB756.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia lilacina Poelt & Vezda	<div><p>Protoblastenia lilacina Poelt &amp; Vězda, Čas. slezsk. Mus. Opavě, Ser. A 19: 26 (1970)</p><p>Description.</p><p>Thallus white to grey, endolithic, farinose to small areoles, K-, C-, UV- to UV + bluish-white, algal cells 3–8 μm. Apothecia orange to dark orange, 0.2–0.6 mm, slightly convex to strongly convex, 1 / 4 - immersed to superficial, sometimes leaving shallow pits, K- or K + fairly weakly red; ca. 40–100 apothecia / cm 2. Epihymenium (yellowish) orange-brown, 12–20 μm thick, K- to K + orange or carmine red, not into solution. Hymenium 50–120 μm thick, with oil droplets. Hypothecium pale yellow, sometimes patchily violet, ca. 60–120 μm thick. Paraphyses ca. 2–3 μm thick, apex not thickened to slightly thickened, sparingly branched and anastomosing. Ascospores 0 - septate, (8.4 –) 10.7–12.4 – 14.2 (– 16.6) × (5.2 –) 6.0–6.7 – 7.3 (– 7.8) μm (n = 30).</p><p>Habitat and distribution.</p><p>The species is rare and occurs only on calcareous rock outcrops and boulders in the hemi-boreal zone in SW Finland. This seems to be the northern limit of this predominantly temperate species. Protoblastenia lilacina grows both on fairly sun-exposed and shady habitats. It may prefer rather dry calcareous rocks. Companion species include, for example, Bagliettoa baldensis s. lat, B. calciseda (DC.) Gueidan &amp; Cl. Roux, Catillaria lenticularis (Ach.) Th. Fr., Hymenelia spp., Lecidella stigmatea (Ach.) Hertel &amp; Leuckert, Staurothele rupifraga (A. Massal.) Arnold, Verrucaria deversa Vain., V. foveolata and V. viridula .</p><p>Notes.</p><p>The species differs from most species of Protoblastenia in having K- or weakly K + red apothecia. The closely-related Protoblastenia oulankaensis is the only other species with this characteristic. However, the two species have different distribution areas as Protoblastenia oulankaensis is only known from the Oulanka area in NE Finland. Morphologically, P. oulankaensis differs in having a thicker hypothecium and often more sparsely occurring apothecia (in most specimens 10–30 apothecia / cm 2).</p><p>Specimens examined. Finland • Varsinais-Suomi, Kemiönsaari (Västanfjärd), Vesterillo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.733334&amp;materialsCitation.latitude=60.0" title="Search Plazi for locations around (long 22.733334/lat 60.0)">Strömmen 200–400 m SE</a>, Pinus sylvestris - dominated forest, calcareous rock outcrop, on 70 cm high E-facing wall, 17 m a. s. l., 60°00'N, 22°44'E, 9. 9. 2009, J. Pykälä 37012 (H) ; • Lohja, Moisio, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.1&amp;materialsCitation.latitude=60.25" title="Search Plazi for locations around (long 24.1/lat 60.25)">between Lohjanharjuntie and Nummentie roads, NW-slope of Lohjanharju esker</a>, Pinus sylvestris - heath forest, on calcareous boulder, 82 m a. s. l., 60°15'N, 24°06'E, 18. 6. 2005, J. Pykälä 27009 H ); • Lohja, Lohja, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.033333&amp;materialsCitation.latitude=60.25" title="Search Plazi for locations around (long 24.033333/lat 60.25)">Pitkäniemi industrial area, 5 m from shore of Lohjanjärvi Lake</a>, deciduous forest, calcareous rock outcrop, on NW-facing 2 m high rock wall, 35 m a. s. l., 60°15'N, 24°02'E, 19. 8. 2005, J. Pykälä 27629 (H) ; • Lohja, Lohja, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.033333&amp;materialsCitation.latitude=60.25" title="Search Plazi for locations around (long 24.033333/lat 60.25)">Pitkäniemi industrial area, deciduous forest ca. 5 m from shore of Lohjanjärvi Lake</a>, calcareous rock outcrop, on 1.7 m high overhanging NW-facing wall, 35 m a. s. l., 60°15'N, 24°02'E, 12. 7. 2021, J. Pykälä 57976 (H) .</p></div>	https://treatment.plazi.org/id/26F7C1B6407055EBA01BCB72E0DFB756	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
A4AF1B5F60E45D47BAD7F03E6789169D.text	A4AF1B5F60E45D47BAD7F03E6789169D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia minuta Pykala & Myllys 2025	<div><p>Protoblastenia minuta Pykälä &amp; Myllys sp. nov.</p><p>Fig. 3 C</p><p>Diagnosis.</p><p>Differs from the other Protoblastenia species with a pale hypothecium in having small apothecia and spores and apothecia which leave pits.</p><p>Type.</p><p>Finland • Enontekiön Lappi, Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.85&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.85/lat 69.03333)">Saana Nature Reserve</a>, E-part, fell, steep SW-slope, dolomite rock outcrop, on S-facing wall, scarce, 715 m a. s. l, 69°02'N, 20°51'E, 12 August 2011, J. Pykälä 44275 (H 9250941 – holotype, GenBank accession number: PV 766683) .</p><p>Description.</p><p>Thallus ochraceous, endolithic, K-, UV-. Apothecia orange, 0.2–0.4 mm, slightly convex to strongly convex, immersed in rock to superficial, leaving shallow to fairly deep pits; ca. 80 apothecia / cm 2. Epihymenium brown-yellow, 10–12 μm thick, K + violet. Hymenium 45–55 μm thick. Hypothecium colourless to yellow, ca. 70–100 μm thick. Paraphyses ca. 2–3 μm thick, apex not thickened to slightly thickened. Ascospores 0 - septate, (7.3 –) 7.6–7.9 – 8.3 (– 8.8) × (4.1 –) 4.3–4.5 – 4.7 (– 4.8) μm (n = 9).</p><p>Habitat and distribution.</p><p>Only one specimen has been found on the calcareous Saana fell in NE Finland, on a dolomite rock outcrop. Companion species include Acarospora macrospora (Hepp) A. Massal. ex Bagl., Enchylium polycarpon, Placynthium asperellum (Ach.) Trevis. s. lat, Romjularia lurida (Ach.) Timdal and Thalloidima alutaceum Anzi.</p><p>Etymology.</p><p>The name refers to the small size of the apothecia and spores.</p><p>Notes.</p><p>The species is rather isolated amongst the sequenced species of Protoblastenia (minimum difference from other species over 9 % in ITS). The species has smaller apothecia and spores than most other Protoblastenia species with a pale hypothecium. Furthermore, other Finnish Protoblastenia species do not leave pits or only rarely leave shallow pits. However, only one specimen is known and the species may be expected to be morphologically more variable. More material is needed to evaluate whether the species can be identified by morphology only. Protoblastenia saanaensis has a rimose thallus and the apothecia do not leave pits. Protoblastenia szaferi differs in having larger apothecia and spores (see Hafellner (2006)).</p></div>	https://treatment.plazi.org/id/A4AF1B5F60E45D47BAD7F03E6789169D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
FE5B4688BA79556AB55401286B667578.text	FE5B4688BA79556AB55401286B667578.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia oulankaensis Pykala & Myllys 2025	<div><p>Protoblastenia oulankaensis Pykälä &amp; Myllys sp. nov.</p><p>Fig. 3 D</p><p>Diagnosis.</p><p>Differs from P. lilacina in often having slightly larger and more sparse apothecia and an often thicker hypothecium.</p><p>Type.</p><p>Finland • Koillismaa, Salla, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.166666&amp;materialsCitation.latitude=66.416664" title="Search Plazi for locations around (long 29.166666/lat 66.416664)">Oulanka National Park, W of Savikoski</a>, cliff, dolomite rock outcrop, on NE-facing wall, 190 m a. s. l., 66°25'N, 29°10'E, 17 August 2010, J. Pykälä 40201 (H 9224130 – holotype, UPS – isotype, GenBank accession number: PV 766686) .</p><p>Description.</p><p>Thallus white to whitish-grey, endolithic, farinose or rimose, ca. 0–0.1 mm thick, K-, C-, UV- to UV + bluish-white, algal cells 5–9 μm. Apothecia orange to dark orange, 0.3–0.9 mm, convex to strongly convex, 1 / 4 - immersed to superficial, often leaving shallow pits, K + weakly red; ca. 10–50 apothecia / cm 2 (in one specimen, ca. 50–80 apothecia / cm 2). Epihymenium dark orange-brown, 12–25 μm thick, K- to K + orange or carmine red. Hymenium 70–120 μm thick, with oil droplets, in one specimen, partly violet in few apothecia. Hypothecium colourless to yellow, more rarely patchily red-brown, ca. 60–220 μm thick. Paraphyses ca. 2–3 μm thick, apex not thickened to slightly thickened, sparingly branched and anastomosing. Ascospores 0 - septate, (8.8 –) 11.8–13.9 – 16.0 (– 18.8) × (5.2 –) 6.6–7.4 – 8.2 (– 9.6) μm (n = 92).</p><p>Habitat and distribution.</p><p>The species occurs only in the Oulanka area in the parishes of Kuusamo and Salla in NE Finland. It grows on N- and S-facing walls of dolomite rock outcrops. Companion species include, for example, Diploschistes gypsaceus (Ach.) Zahlbr., Gyalecta jenensis (Batsch) Zahlbr., Lempholemma isidioides (Nyl. ex Arnold) H. Magn., Protoblastenia borealis, Sagiolechia protuberans, Scytinium parvum, Verrucaria foveolata and Xanthocarpia crenulatella (Nyl.) Frödén, Arup &amp; Søchting s. lat. Based on the ITS phylogeny, one GenBank specimen of P. lilacina from northern Italy (AY 425629) belongs to P. oulankaensis (Fig. 1).</p><p>Etymology. In Finland, the species occurs only in the Oulanka area in the northeast.</p><p>Notes.</p><p>Protoblastenia oulankaensis is closely related to P. lilacina, but differs clearly from that species in its ITS profile (10 % difference). It is also morphologically rather similar to P. lilacina, having a weak K + reaction of apothecia and large spores. Protoblastenia oulankaensis tends to have slightly larger apothecia, which usually occur more sparsely (in most specimens ca. 10–50 apothecia / cm 2) and, on average, a thicker hypothecium. Protoblastenia lilacina is only known from the hemi-boreal zone of SW Finland, ca. 800 kilometres south to south-west.</p><p>Other specimens examined.</p><p>Finland • Koillismaa, Salla, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.166666&amp;materialsCitation.latitude=66.416664" title="Search Plazi for locations around (long 29.166666/lat 66.416664)">Oulanka National Park, Savikoski 300 m N</a>, Pinus sylvestris - forest, steep N-slope, dolomite rock outcrop, on NW-facing wall, st pc – sp, 180 m a. s. l., 66°25'N, 29°10'E, 10. 8. 2010, J. Pykälä 39629 (H) ; • Salla, Oulanka National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.166666&amp;materialsCitation.latitude=66.416664" title="Search Plazi for locations around (long 29.166666/lat 66.416664)">Savilamminniemi, shore of Savilampi Lake</a>, cliff, dolomite rock outcrop, on E-facing wall, sp, 180 m a. s. l., 66°25'N, 29°10'E, 12. 8. 2010, J. Pykälä 39810 (H) ; • Salla, Oulanka National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.183332&amp;materialsCitation.latitude=66.433334" title="Search Plazi for locations around (long 29.183332/lat 66.433334)">Savilampi 1.4 km NE, bank of Savinajoki river</a>, cliff, dolomite rock outcrop, upper slope, on NE-facing wall, 192 m a. s. l., 66°26'N, 29°11'E, 23. 8. 2011, J. Pykälä 44928 (H) ; • Salla, Hautajärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.15&amp;materialsCitation.latitude=66.433334" title="Search Plazi for locations around (long 29.15/lat 66.433334)">Kurtinniittykuru</a>, dolomite rock outcrop, on SW-facing wall, 195 m a. s. l., 66°26'N, 29°09'E, 24. 8. 2011, J. Pykälä 44949 (H) ; • Salla, Hautajärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.15&amp;materialsCitation.latitude=66.433334" title="Search Plazi for locations around (long 29.15/lat 66.433334)">Kurtinniittykuru</a>, dolomite rock outcrop, on SE-facing wall, 195 m a. s. l., 66°26'N, 29°09'E, 24. 8. 2011, J. Pykälä 44953 (H) ; • Kuusamo, Juuma, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.366667&amp;materialsCitation.latitude=66.26667" title="Search Plazi for locations around (long 29.366667/lat 66.26667)">Myllyniemi</a>, dolomite rock outcrop, on NE-facing wall, 235 m a. s. l., 66°16'N, 29°22'E, 11. 8. 2020, J. Pykälä 55995 (H) .</p></div>	https://treatment.plazi.org/id/FE5B4688BA79556AB55401286B667578	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
2B50F168D1BC5381A71F8B819B54D36C.text	2B50F168D1BC5381A71F8B819B54D36C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia pseudocompressa Pykala & Myllys 2025	<div><p>Protoblastenia pseudocompressa Pykälä &amp; Myllys sp. nov.</p><p>Fig. 4 A</p><p>Diagnosis.</p><p>Differs from P. compressa in often having convex apothecia.</p><p>Type.</p><p>Finland • Koillismaa, Salla, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=28.916666&amp;materialsCitation.latitude=66.7" title="Search Plazi for locations around (long 28.916666/lat 66.7)">Onkamo, Latvarova SW</a>, clear-cut heath forest, on calcareous boulder, 258 m a. s. l., 66°42'N, 28°055'E, 9 August 2021, J. Pykälä 58457 (H 9250942 – holotype, UPS – isotype, GenBank accession number: PV 766696) .</p><p>Description.</p><p>Thallus white to grey, continuous, rimose (usually) to areolate, rarely endolithic, ca. 0–0.15 mm thick, K-, C-, UV- to UV + bluish-white, algal cells 6–12 μm. Apothecia orange-yellow to orange, 0.2–0.7 mm, plane to strongly convex, 1 / 4–1 - immersed in thallus or rock to superficial, sometimes leaving shallow pits; ca. 20–120 apothecia / cm 2. Epihymenium yellow-brown to orange-brown, 12–18 μm thick, K + violet. Hymenium 50–60 μm thick. Hypothecium colourless to pale yellow, rarely pink (in one specimen), ca. 50–120 μm thick. Paraphyses ca. 1.5–2.5 μm thick, sparsely branched and anastomosing. Ascospores 0 - septate, (6.8 –) 9.1–10.6 – 12.1 (– 13.0) × (4.2 –) 4.7–5.5 – 6.2 (– 8.0) μm (n = 51).</p><p>Habitat and distribution.</p><p>Protoblastenia pseudocompressa has been verified by ITS only from the Oulanka area (parishes Kuusamo and Salla) in NE Finland. However, a morphologically similar specimen was found in the neighbouring biogeographical Province of Kainuu. The species cannot be identified with certainty by morphology only and it is likely that more populations occur in the Oulanka area. It mainly grows on dolomite rock outcrops (one specimen was collected from a dolomite boulder) and may prefer half-shady habitats. Typical companion species are Acarospora glaucocarpa, Clauzadea monticola, Farnoldia jurana, Hymenelia prevostii (Duby) Kremp., H. rhodopis, Lathagrium fuscovirens, Placynthium nigrum, Thelidium declivum, Verrucaria deversa, V. muralis, V. kuusamoensis Pykälä, Kantelinen &amp; Myllys and V. subtilis .</p><p>Etymology.</p><p>The name refers to the close relationship in ITS and morphological similarity to P. compressa .</p><p>Notes.</p><p>Protoblastenia pseudocompressa is closely related to P. compressa and the two species are also morphologically similar. Protoblastenia pseudocompressa has, on average, slightly smaller spores than P. compressa, but there is a wide overlap. Furthermore, P. pseudocompressa specimens with plane to slightly convex apothecia may be confused with P. compressa . However, the apothecia of P. pseudocompressa are often convex, while the apothecia of P. compressa are always plane to slightly convex. Based on the distribution of sequenced specimens, the two species are geographically well separated. Protoblastenia pseudocompressa specimens with convex apothecia may also be confused with P. borealis, which, however, often has a thicker hymenium.</p><p>Other specimens examined.</p><p>Finland • Koillismaa, Kuusamo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.316668&amp;materialsCitation.latitude=66.35" title="Search Plazi for locations around (long 29.316668/lat 66.35)">Oulanka National Park, Pikkukönkäänkuru</a>, dolomite rock outcrop, small W-facing wall, 175 m a. s. l., 66°21'N, 29°19'E, 8. 8. 2009, J. Pykälä 35901 (H) ; • Salla, Oulanka National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.166666&amp;materialsCitation.latitude=66.416664" title="Search Plazi for locations around (long 29.166666/lat 66.416664)">400 m N of Savilampi, bank of River Savinajoki</a>, dolomite rock outcrop, on NW-facing wall, 178 m a. s. l., 66°25'N, 29°10'E, 13. 8. 2009, J. Pykälä 36325 (H) ; • Kuusamo, Oulanka National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.316668&amp;materialsCitation.latitude=66.36667" title="Search Plazi for locations around (long 29.316668/lat 66.36667)">Kiutaköngäs 400 m N</a>, SW-slope, Pinus sylvestris - dominated forest, small dolomite rock outcrop, on SW-facing wall, scarce, 175 m a. s. l., 66°22'N, 29°19'E, 6. 8. 2010, J. Pykälä 39367 (H) ; • Kuusamo, Oulanka National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.333334&amp;materialsCitation.latitude=66.36667" title="Search Plazi for locations around (long 29.333334/lat 66.36667)">Kiutaköngäs, bank of Oulankajoki River</a>, by the rapids, calciferous (dolomite) schistose rock outcrop, on rather flat rock, 150 m a. s. l., 66°22'N, 29°20'E, 10. 8. 2010, J. Pykälä 39693 (H) ; • Kuusamo, Oulanka, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.416666&amp;materialsCitation.latitude=66.36667" title="Search Plazi for locations around (long 29.416666/lat 66.36667)">Putaanoja, 500 m W-NW of Hautala</a>, dolomite rock outcrop, on E-facing wall, scarce, 232 m a. s. l., 66°22'N, 29°25'E, 15. 8. 2010, J. Pykälä 39987 (H) .</p></div>	https://treatment.plazi.org/id/2B50F168D1BC5381A71F8B819B54D36C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
7265A0B018825346AC78143B76781C9F.text	7265A0B018825346AC78143B76781C9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia pseudoterricola Pykala & Myllys 2025	<div><p>Protoblastenia pseudoterricola Pykälä &amp; Myllys sp. nov.</p><p>Fig. 4 B</p><p>Diagnosis.</p><p>Differs from P. terricola in having a thinner thallus and hypothecium.</p><p>Type.</p><p>Finland • Enontekiön Lappi, Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.85&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.85/lat 69.03333)">Saana</a>, fell, steep NE-slope, dolomite rock outcrop, beneath NE-facing wall, on dolomite pebbles, 820 m a. s. l., 69°02'N, 20°51'E, 11 August 2011, J. Pykälä 44157 (H 9235068 – holotype, GenBank accession number: PV 766697) .</p><p>Description.</p><p>Thallus white, rimose, ca. 0.05–0.1 mm thick, K-, UV + bluish-white. Apothecia orange (young apothecia) to dirty orange-brown, 0.3–0.8 mm, convex to strongly convex, superficial; ca. 30 apothecia / cm 2. Epihymenium orange-brown, 10–15 μm thick, K + violet. Hymenium 70 μm thick. Hypothecium dark reddish-brown to violet, ca. 60–100 μm thick. Ascospores 0 - septate, (7.9 –) 8.6–9.9 – 11.2 (– 12.4) × (4.5 –) 4.8–5.5 – 6.2 (– 7.0) μm (n = 20).</p><p>Habitat and distribution.</p><p>Only one specimen of this species is known. It was collected from dolomite pebbles beneath an NE-facing wall of a dolomite rock outcrop on the Saana fell (NW Finland). Companion species include Farnoldia jurana, Polyblastia inconspicua Savić &amp; Tibell and Thelidium huuskonenii Pykälä &amp; Myllys.</p><p>Etymology.</p><p>The name refers to the similarities in morphology and ITS to P. terricola .</p><p>Notes.</p><p>The species is closely related to P. terricola and P. violacea . Protoblastenia terricola has rimose to areolate thallus (when growing on rock) mainly exceeding 0.1 mm in thickness and an often thicker hypothecium. Protoblastenia violacea has orange apothecia and smaller spores. More material is needed to confirm whether these three species can be distinguished by morphology only. Protoblastenia westbergii resembles the new species, but has more densely occurring apothecia (ca. 40–160 apothecia / cm 2) and larger spores.</p></div>	https://treatment.plazi.org/id/7265A0B018825346AC78143B76781C9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
BDE342D3B7B8547D84F9CF3E67DE85D5.text	BDE342D3B7B8547D84F9CF3E67DE85D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia remota Pykala & Myllys 2025	<div><p>Protoblastenia remota Pykälä &amp; Myllys sp. nov.</p><p>Fig. 4 C</p><p>Diagnosis.</p><p>Differs from P. borealis and P. pseudocompressa by often smaller more convex apothecia and a smaller thallus.</p><p>Type.</p><p>Finland • Varsinais-Suomi, Perniö, Lupaja, Alhonmäki, abandoned lime quarry, on W-slope, 9 August 1991, J. Pykälä 8547 (H 9250943 – holotype, GenBank accession number: PV 766698) .</p><p>Description.</p><p>Thallus white to grey, endolithic to sparse small areoles, 0.2–0.5 mm wide, ca. 0–0.15 mm thick, K-, C-, UV- to UV + bluish-white. Apothecia orange, 0.2–0.6 mm, convex to strongly convex, 1 / 2 - immersed to mainly superficial; ca. 15–80 apothecia / cm 2. Epihymenium yellow-brown to dark orange, 12–20 μm thick, K + violet. Hymenium 50–80 μm thick. Hypothecium colourless to pale yellow, ca. 100–200 μm thick. Paraphyses ca. 2–2.5 μm thick, apex not thickened to thickened to ca. 3–5 μm thick. Ascospores 0 - septate, (8.4 –) 9.8–11.4 – 12.9 (– 14.4) × (4.8 –) 5.0–5.6 – 6.1 (– 7.1) μm (n = 36).</p><p>Habitat and distribution.</p><p>Two specimens of this species are known: one from a lime quarry in SW Fnland and one from a calciferous schistose cliff in NE Finland. Companion species include Lathagrium fuscovirens, Mycobilimbia tetramera (De Not.) Vitik., Ahti, Kuusinen, Lommi &amp; T. Ulvinen ex Hafellner &amp; Türk and Verrucaria deversa . Furthermore, two closely-related specimens from North America and previously identified as Protoblastenia aff. rupestris (MZ 922194, MZ 922195) (Munger et al. 2022) may belong to P. remota (Fig. 1).</p><p>Etymology.</p><p>The name refers to the large difference in ITS of the species compared to other Protoblastenia species (differs by ca. 15–18 % from other species).</p><p>Notes.</p><p>Based on ITS sequences, this species is not closely related to any sequenced Protoblastenia species. The species is morphologically a typical Protoblastenia and very difficult to distinguish from several species, particularly from P. borealis and P. pseudocompressa . However, these two species have, on average, less convex apothecia, a thinner hypothecium and often a better developed thallus, but there is a wide overlap. In addition, Protoblastenia borealis often has larger apothecia (up to 1 mm).</p><p>Other specimens examined.</p><p>Finland • Koillismaa, Kuusamo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.516666&amp;materialsCitation.latitude=66.26667" title="Search Plazi for locations around (long 29.516666/lat 66.26667)">Oulanka National Park, Päähkänänkallio</a>, high cliff, calciferous (dolomite) schistose rock outcrop, on SE-facing wall, st pc, 195 m a. s. l., 66°16'N, 29°31'E, 7. 8. 2010, J. Pykälä 39455 (H) .</p></div>	https://treatment.plazi.org/id/BDE342D3B7B8547D84F9CF3E67DE85D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
3E73610245705DF6B9E335C87959A1E5.text	3E73610245705DF6B9E335C87959A1E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia rikkinenii Pykala & Myllys 2025	<div><p>Protoblastenia rikkinenii Pykälä &amp; Myllys sp. nov.</p><p>Fig. 4 D</p><p>Diagnosis.</p><p>Differs from P. fennoarctica in always having convex to strongly convex apothecia and from P. saanaensis and P. timdalii in having sparsely occurring apothecia and a thick hypothecium.</p><p>Type.</p><p>Finland • Enontekiön Lappi, Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.433332&amp;materialsCitation.latitude=69.2" title="Search Plazi for locations around (long 21.433332/lat 69.2)">Toskalharji, Toskaljärvi N</a>, fell, dolomite rock outcrop, on dolomite pebbles, 735 m a. s. l., 69°12'N, 21°26'E, 2 August 2011, J. Pykälä 43252 (H 9250944 – holotype, GenBank accession number: PV 766700) .</p><p>Description.</p><p>Thallus white, grey to pale brown, rimose to areolate, ca. 0.05–0.2 mm thick, K-, C-, UV-, algal cells 5–9 μm. Apothecia yellow-orange, 0.3–0.8 mm, convex to strongly convex, 1 / 2 - immersed to superficial; ca. 20–40 apothecia / cm 2. Epihymenium dirty yellow to orange-yellow, 12–18 μm thick, K + violet. Hymenium 50–70 μm thick. Hypothecium pale yellow, ca. 80–300 μm thick. Paraphyses ca. 2–2.5 μm thick, apex slightly thickened to 3 μm thick. Ascospores 0 - septate, (7.4 –) 8.5–9.6 – 10.7 (– 12.6) × (4.4 –) 4.7–5.1 – 5.5 (– 6.3) μm (n = 52).</p><p>Habitat and distribution.</p><p>The species occurs on dolomite rock outcrops, dolomite stone and dolomite pebbles on the fells. It may prefer sun-exposed sites. Companion species include Halecania alpivaga (Th. Fr.) M. Mayrhofer, Hymenelia heteromorpha (Kremp.) Lutzoni, H. prevostii, H. rhodopis, Lecidea polycocca Sommerf., Polyblastia sp., Rhizocarpon petraeum, Thelidium huuskonenii and Verrucaria lapponica Pykälä. One GenBank sequence (KY 266981) originates from northern Norway.</p><p>Etymology.</p><p>The name is in honour of Prof Jouko Rikkinen (Helsinki) for his innovative work on various fields of lichenology.</p><p>Notes.</p><p>The species is related to P. fennoarctica, which often has more densely occurring, often plane and / or yellow apothecia. However, an ITS sequence may be needed for unambiguous identification of specimens. Protoblastenia saanaensis and P. timdalii have a thinner hypothecium and more densely occurring apothecia. Protoblastenia calvella has more densely occurring apothecia and slightly larger spores.</p><p>Other specimens examined.</p><p>Finland • Enontekiön Lappi, Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.8&amp;materialsCitation.latitude=69.05" title="Search Plazi for locations around (long 20.8/lat 69.05)">Saana Nature Reserve, W-part</a>, fell, dolomite rock outcrop, on W-facing wall, 700 m a. s. l., 69°03'N, 20°48'E, 9. 8. 2011, J. Pykälä 43910 (H) ; • Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.833334&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.833334/lat 69.03333)">Saana Nature Reserve</a>, fell, steep SW-slope, dolomite rock outcrop, beneath SW-facing wall, on dolomite stone, 840 m a. s. l., 69°02'N, 20°50'E, 14. 8. 2011, J. Pykälä 44387 (H) .</p></div>	https://treatment.plazi.org/id/3E73610245705DF6B9E335C87959A1E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
CE2CF32D1453527DAFB5185219F1D87F.text	CE2CF32D1453527DAFB5185219F1D87F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia rupestris (Scop.) J. Steiner	<div><p>Protoblastenia rupestris (Scop.) J. Steiner, Verh. Kaiserl.-Königl. zool.-bot. Ges. Wien 61: 47 (1911)</p><p>Lichen rupestris Scop., Fl. carniol., Edn 2 (Wien) 2: 363 (1772).</p><p>Description.</p><p>Thallus whitish-grey to rarely pale brown, rimose to predominantly areolate, areoles 0.1–0.8 mm, ca. 0.1–0.2 mm thick, K-, C-, UV- to UV + bluish-white, algal cells 4–12 μm. Apothecia yellow-orange to orange, 0.2–0.9 mm, slightly convex to convex, 3 / 4 - immersed in thallus to superficial; ca. 40–100 apothecia / cm 2. Epihymenium orange-brown, 12–20 μm thick, K + violet. Hymenium 50–80 μm thick. Hypothecium pale yellow, ca. 60–80 μm thick. Paraphyses ca. 1.5–2.5 μm thick, apex not thickened to slightly thickened to 2–3 μm thick, sparingly branched and anastomosing. Ascospores 0 - septate, (8.2 –) 9.1–10.3 – 11.6 (– 13.0) × (5.2 –) 5.6–6.3 – 7.1 (– 7.6) μm (n = 43).</p><p>Habitat and distribution.</p><p>Sequenced specimens were collected from rather sun-exposed calcareous rock outcrops in the hemi-boreal zone in Finland. The species may prefer calcareous rock outcrops close to shores.</p><p>Notes.</p><p>Protoblastenia rupestris sequences available in GenBank are heterogeneous and distributed in three different groups in our ITS phylogeny (Fig. 1). Specimens similar to those in Kainz and Rambold (2004) are likely to belong to P. rupestris s. stricto as the species was described from Central Europe. The sequence MZ 159571 (UK) is related to P. incrustans and the sequence KY 266981 (Norway) belongs in P. rikkinenii . Morphologically, the Finnish specimens of P. rupestris may be confused with the closely-related P. borealis . However, the P. rupestris specimens have a predominantly areolate thallus, while the thalli of P. borealis are more often rimose and usually do not exceed 0.1 mm in thickness. Furthermore, P. borealis often has a thicker hypothecium. Despite these minor differences, the two species often cannot be distinguished with certainty by morphology only.</p><p>Specimens examined.</p><p>Finland • Varsinais-Suomi, Lohja, Skraatila, Kirkkovuori E, cliff, calcareous rock outcrop, on S-slope, 8. 8. 1990, J. Pykälä 5963 (H) ; • Lohja, Lohja, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.033333&amp;materialsCitation.latitude=60.25" title="Search Plazi for locations around (long 24.033333/lat 60.25)">Pitkäniemi industrial area, by the sauna, on shore of Lohjanjärvi Lake</a>, flat calcareous rock outcrop, 33 m a. s. l., 60°15'N, 24°02'E, 19. 8. 2005, J. Pykälä 27569 (H) ; • Karjalohja, Saarenpää, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.783333&amp;materialsCitation.latitude=60.216667" title="Search Plazi for locations around (long 23.783333/lat 60.216667)">Saarenpäänniemi</a>, calcareous rock outcrop on shore of Lohjanjärvi Lake, on small SW-facing wall, 35 m a. s. l., 60°13'N, 23°47'E, 7. 10. 2008, J. Pykälä 34043 (H) ; • Salo (Särkisalo), Kaukosalo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.966667&amp;materialsCitation.latitude=60.1" title="Search Plazi for locations around (long 22.966667/lat 60.1)">Klintinmäki</a>, cliff, calciferous schistose rock outcrop, on SW-facing wall, st pc, 35 m a. s. l., 60°06'N, 22°58'E, 4. 10. 2011, J. Pykälä 46413 (H) .</p></div>	https://treatment.plazi.org/id/CE2CF32D1453527DAFB5185219F1D87F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
3817FD07D2215FE9B64B2ABC5DC32B13.text	3817FD07D2215FE9B64B2ABC5DC32B13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia saanaensis Pykala & Myllys 2025	<div><p>Protoblastenia saanaensis Pykälä &amp; Myllys sp. nov.</p><p>Fig. 5 A</p><p>Diagnosis.</p><p>Differs from P. minuta in apothecia not leaving pits, from P. rikkinenii in having more densely occurring apothecia, from P. fennoarctica in having a thinner hypothecium and from P. timdalii in often having smaller apothecia.</p><p>Type.</p><p>Finland • Enontekiön Lappi, Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.833334&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.833334/lat 69.03333)">Saana Nature Reserve</a>, fell, steep SW-slope, dolomite rock outcrop, SW-slope, on dolomite pebbles, 835 m a. s. l., 69°02'N, 20°50'E, 14 August 2011, J. Pykälä 44431 (H 9250945 – holotype, GenBank accession number: PV 766707) .</p><p>Description.</p><p>Thallus white, grey or pale brown, rimose to areolate, areoles 0.3–0.8 mm, ca. 0.03–0.1 mm thick, K-, C-, UV-, algal cells 5–12 μm. Apothecia yellow to orange-yellow, 0.3–0.5 mm, slightly convex to strongly convex, 3 / 4 - immersed to superficial; ca. 20–80 apothecia / cm 2. Epihymenium orange-yellow, 12–17 μm thick, K + violet. Hymenium 50–70 μm thick. Hypothecium pale yellow (some orange-yellow in squash), ca. 40–100 μm thick. Paraphyses ca. 2–2.5 μm thick, apex not thickened to slightly thickened, branching. Ascospores 0 - septate, (5.4 –) 7.1–8.4 – 9.7 (– 11.4) × (3.8 –) 4.3–4.8 – 5.4 (– 6.1) μm (n = 47).</p><p>Habitat and distribution.</p><p>The species has only been found on the calcareous Saana fell in NW Finland. The specimens are from a dolomite stone and dolomite pebbles in sun-exposed to rather shady habitats. Companion species include Acarospora glaucocarpa .</p><p>Etymology.</p><p>The name refers to the Saana fell where both known specimens were collected.</p><p>Notes.</p><p>The species is most closely related to P. timdalii (ca. 5–6 % difference in ITS) from which it is difficult to distinguish by morphology. It may have slightly smaller apothecia and spores. It also resembles P. minuta and the closely-related P. rikkinenii in having similar small spores. The only known specimen of P. minuta has apothecia which leave shallow to deep pits. Protoblastenia rikkinenii has sparsely occurring apothecia. Protoblastenia fennoarctica resembles P. saanaensis, but usually has a thicker hypothecium, slightly larger spores and sometimes plane apothecia.</p><p>Other specimens examined.</p><p>Finland • Enontekiön Lappi, Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.833334&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.833334/lat 69.03333)">Saana Nature Reserve</a>, fell, steep SW-slope, dolomite rock outcrop, under overhanging SW-facing wall, on dolomite stone, 835 m a. s. l., 69°02'N, 20°50'E, 14. 8. 2011, J. Pykälä 44438 (H) .</p></div>	https://treatment.plazi.org/id/3817FD07D2215FE9B64B2ABC5DC32B13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
9F8D532582E8510C819564D63E148805.text	9F8D532582E8510C819564D63E148805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia terricola (Anzi) Lynge 1921	<div><p>Protoblastenia terricola (Anzi) Lynge, Rep. Sci. Res. Norweg. Exped. Novaya Zemlya, 1921 43: 216 (1928)</p><p>Biatora rupestris var. terricola Anzi, Cat. Lich. Sondr.: 78 (1860).</p><p>Description.</p><p>Thallus white to whitish-grey, rimose to areolate, areoles 0.2–0.8 mm, ca. 0.05–0.2 mm thick, K-, C-, UV + bluish-white. Apothecia dirty orange to brown-orange (some with an olive tinge), 0.4–0.9 mm, strongly convex, superficial, K + violet; ca. 70–120 apothecia / cm 2. Epihymenium dark orange-brown, 8–15 μm thick, K + violet. Hymenium 60–100 μm thick. Hypothecium violet to violet-brown, 70–180 μm thick. Paraphyses ca. 2–2.5 μm thick, apex not thickened to slightly thickened, sparingly branched and anastomosing. Ascospores 0 - septate, (9.4 –) 9.7–10.8 – 11.9 (– 12.4) × (4.7 –) 4.9–5.5 – 6.2 (– 7.3) μm (n = 16).</p><p>Habitat and distribution.</p><p>The species occurs only in northern Finland (biogeographical Provinces Ks and EnL) with one separate population in the biogeographical Province PK (not sequenced) in eastern Finland. It grows on calcareous rock outcrops and on calcareous soil.</p><p>Notes.</p><p>The specimens on calcareous soil (not sequenced) have a squamulose thallus, thus differing from the other species of Protoblastenia (Kainz and Rambold 2004) . However, on dolomite rock outcrops, the thallus is more weakly developed, rimose to areolate. The apothecia of P. terricola tend to be dirty orange to brown. Protoblastenia terricola may be most difficult to distinguish from P. pseudoterricola . The only known specimen of P. pseudoterricola has a 60–100 μm thick hypothecium and a 0.05–0.1 mm thick thallus.</p><p>Specimen sequenced.</p><p>Finland • Koillismaa, Salla, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.183332&amp;materialsCitation.latitude=66.433334" title="Search Plazi for locations around (long 29.183332/lat 66.433334)">Oulanka National Park, Savilampi 1.2 km NE</a>, bank of Savinajoki River, dolomite rock outcrop, on SE-facing wall, 66°26'N, 29°11'E, 23. 8. 2011, J. Pykälä 44876 (H) .</p></div>	https://treatment.plazi.org/id/9F8D532582E8510C819564D63E148805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
9DCFD2E05AC45FABB581918161BBE60C.text	9DCFD2E05AC45FABB581918161BBE60C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia timdalii Pykala & Myllys 2025	<div><p>Protoblastenia timdalii Pykälä &amp; Myllys sp. nov.</p><p>Fig. 5 B</p><p>Diagnosis.</p><p>Differs from P. saanaensis in having slightly larger apothecia and spores.</p><p>Type.</p><p>Finland • Enontekiön Lappi, Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.85&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.85/lat 69.03333)">Saana</a>, fell, steep NE-slope, dolomite rock outcrop, on NE-facing wall, 820 m a. s. l., 69°02'N, 20°51'E, 11 August 2011, J. Pykälä 44177 (H 9250946 – holotype, O – isotype, GenBank accession number: PV 766711) .</p><p>Description.</p><p>Thallus white, grey or pale brown, endolithic, rimose to areolate, areoles 0.1–0.4 mm, ca. 0–0.15 mm thick, K-, C-, UV-. Apothecia yellow-orange, 0.3–0.8 mm, convex to strongly convex, semi-immersed to superficial; ca. 40–60 apothecia / cm 2. Epihymenium brown-yellow to dirty yellow, K + violet. Hymenium 60–70 μm thick, pale, partly yellow. Hypothecium colourless to yellow, ca. 60–100 μm thick. Paraphyses ca. 2–2.5 μm thick, apex not thickened to slightly thickened. Ascospores 0 - septate, (6.8 –) 7.9–9.2 – 10.5 (– 12.6) × (3.8 –) 4.2–4.8 – 5.5 (– 7.0) μm (n = 46).</p><p>Habitat and distribution.</p><p>Two specimens are known from the calcareous fells of Enontekiö, one from a NE-facing wall of a dolomite rock outcrop and one from dolomite stones on Dryas octopetala heath. This suggests that the species may have considerable microhabitat variation on the fells. Companion species include Halecania alpivaga, Polyblastia spp. and Thelidium auruntii .</p><p>Etymology.</p><p>The name is in honour of Prof Einar Timdal (Oslo) for his numerous important contributions to the taxonomy, biogeography and conservation of Fennoscandian lichens, including studies on the related genus Psora and an important technical edition of the Nordic lichen journal Graphis Scripta.</p><p>Notes.</p><p>The species is most closely related to P. pseudoincrustans ined. This undescribed species differs in having plane to convex apothecia, a thick hymenium (80–115 μm thick) and larger spores (mean 12 × 7 μm) (Kainz and Rambold 2004). However, in our phylogeny, P. pseudoincrustans is divided into two species. The closely-related P. saanaensis (differing ca. 5 % in ITS) is also morphologically very similar to P. timdalii . Protoblastenia saanaensis may have slightly smaller apothecia and spores, but more material is needed to confirm this.</p><p>Other specimens examined.</p><p>Finland • Enontekiön Lappi, Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.466667&amp;materialsCitation.latitude=69.2" title="Search Plazi for locations around (long 21.466667/lat 69.2)">Toskalharji, 1.2 km NE of Toskaljärvi</a>, fell, SW-slope, gentle E-slope, Dryas heath, on dolomite stones, 875 m a. s. l., 69°12'N, 21°28'E, 5. 8. 2011 J. Pykälä 43624 (H) .</p></div>	https://treatment.plazi.org/id/9DCFD2E05AC45FABB581918161BBE60C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
33C4E5932077503CB3F19712B1F22336.text	33C4E5932077503CB3F19712B1F22336.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia violacea Pykala & Myllys 2025	<div><p>Protoblastenia violacea Pykälä &amp; Myllys sp. nov.</p><p>Fig. 5 C</p><p>Diagnosis.</p><p>Differs from most Protoblastenia species with violet hypothecium in having a thick hypothecium and from P. westbergii in having less densely occurring apothecia and smaller spores.</p><p>Type.</p><p>Finland • Enontekiön Lappi, Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.5&amp;materialsCitation.latitude=69.183334" title="Search Plazi for locations around (long 21.5/lat 69.183334)">Toskalharji, Toskalpahta</a>, fell, dolomite rock outcrop, beneath SW-facing wall, on dolomite pebbles, 800 m a. s. l., 69°11'N, 21°30'E, 4 August 2011, J. Pykälä 43524 (H 9250947 – holotype, GenBank accession number: PV 766713) .</p><p>Description.</p><p>Thallus white, grey or pale brown, rimose to areolate, areoles 0.15–0.4 mm, ca. 0.01–0.15 mm thick, K-, C-, UV-, algal cells 6–10 μm. Apothecia orange, 0.3–1.0 mm, convex to mainly strongly convex, superficial, high; ca. 20–40 apothecia / cm 2, sometimes forming clusters of smaller apothecia on surface of old apothecia. Epihymenium orange-brown, 12–14 μm thick, K + violet. Hymenium 60–80 μm thick. Hypothecium purple, orange-red to red-violet, ca. 100–280 μm thick. Paraphyses ca. 2–2.5 (– 3) μm thick, apex not thickened to slightly thickened, often branching. Ascospores 0 - septate, (6.3 –) 7.9–9.2 – 10.5 (– 12.7) × (3.7 –) 4.2–4.6 – 5.1 (– 5.6) μm (n = 52).</p><p>Habitat and distribution.</p><p>The three specimens are known from the calcareous Toskalharji fell (NW Finland), all from dolomite pebbles. Companion species include Farnoldia jurana and Lecidella stigmatea .</p><p>Etymology.</p><p>The name refers to the violet hypothecium of the species.</p><p>Notes.</p><p>The species is related to P. pseudoterricola and P. terricola, which have mainly brown apothecia and broader spores. Protoblastenia violacea differs from P. siebenhaariana in having more densely occurring apothecia (only ca. 1–10 apothecia / cm 2 in P. siebenhaariana; Kainz and Rambold (2004)), slightly smaller apothecia and a thinner hymenium. Protoblastenia westbergii has more densely occurring apothecia and larger spores.</p><p>Other specimens examined.</p><p>Finland • Enontekiön Lappi, Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.483334&amp;materialsCitation.latitude=69.183334" title="Search Plazi for locations around (long 21.483334/lat 69.183334)">Toskalharji, Toskalpahta</a>, fell, SW-slope, on dolomite pebbles, 765 m a. s. l., 69°11'N, 21°29'E, 1. 8. 2011, J. Pykälä 43015 (H) ; • Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.5&amp;materialsCitation.latitude=69.183334" title="Search Plazi for locations around (long 21.5/lat 69.183334)">Toskalharji, 1.2 km NE of Toskaljärvi</a>, fell, SW-slope, gentle E-slope, Dryas heath, on dolomite pebbles, 875 m a. s. l., 69°11'N, 21°30'E, 5. 8. 2011, J. Pykälä 43605 (H) .</p></div>	https://treatment.plazi.org/id/33C4E5932077503CB3F19712B1F22336	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
A170C9C68A7653FB98BA14A165D0344F.text	A170C9C68A7653FB98BA14A165D0344F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoblastenia westbergii Pykala & Myllys 2025	<div><p>Protoblastenia westbergii Pykälä &amp; Myllys sp. nov.</p><p>Fig. 5 D</p><p>Diagnosis.</p><p>Differs from P. dolomitica in having slightly broader spores, darker orange apothecia and a thicker hypothecium.</p><p>Type.</p><p>Finland • Enontekiön Lappi, Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.85&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.85/lat 69.03333)">Saana Nature Reserve, E-part</a>, fell, steep SW-slope, on dolomite boulder, 910 m a. s. l., 69°02'N, 20°51'E, 13 August 2011, J. Pykälä 44316 (H 9229821 – holotype, GenBank accession number: PV 766718) .</p><p>Description.</p><p>Thallus white to whitish-grey, rimose to areolate, areoles 0.15–0.8 mm, ca. 0.05–0.3 mm thick, K-, C-, UV- to UV + bluish-white, algal cells 4–8 μm, thallus often strongly invaded by cyanobacteria. Apothecia orange to dark brownish-orange, 0.2–1.1 mm, slightly convex to convex, 3 / 4 - immersed to superficial, in one specimen with pale margin ca. 30–50 μm thick; ca. 40–200 apothecia / cm 2. Epihymenium dirty yellow to orange-brown, 10–15 μm thick, K + violet. Hymenium 40–70 μm thick, often with oil droplets. Hypothecium red-brown to red-violet, ca. 60–150 μm thick. Paraphyses ca. 2–2.5 μm thick, apex not thickened to slightly thickened to 3 μm thick, some branched and anastomosing. Ascospores 0 - septate, (8.4 –) 10.5–12.0 – 13.6 (– 15.0) × (5.0 –) 5.4–6.1 – 6.9 (– 7.8) μm (n = 39).</p><p>Habitat and distribution.</p><p>Four specimens are known from the calcareous Saana and Toskalharji fells in NW Finland. On Saana, both specimens occurred close to each other on a steep SW-slope on a dolomite boulder. On Toskalharji, the species was found on a SE-facing slope on dolomite pebbles. The species may prefer dry, sun-exposed habitats. Companion species include Farnoldia jurana, Parmeliella thriptophylla (Ach.) Müll. Arg., Thelidium auruntii and Verrucaria sp.</p><p>Etymology.</p><p>The name is in honour of Dr Martin Westberg (Uppsala) for his important work on the taxonomy and biogeography of northern European lichens.</p><p>Notes.</p><p>The species is closely related to P. dolomitica (4 % difference in ITS). Protoblastenia dolomitica has, on average, smaller and paler, sometimes olive-yellow apothecia, narrower spores and a thinner hypothecium. However, P. westbergii often also has a thin hypothecium. Protoblastenia arupii has slightly broader spores, a thin hypothecium and possibly thinner thallus.</p><p>Other specimens examined.</p><p>Finland • Enontekiön Lappi, Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.433332&amp;materialsCitation.latitude=69.183334" title="Search Plazi for locations around (long 21.433332/lat 69.183334)">Toskalharji, Toskaljärvi N</a>, fell, dolomite scree, gentle SE-slope, on dolomite pebbles, 715 m a. s. l., 69°11'N, 21°26'E, 2. 8. 2011, J. Pykälä 43196 (H) ; • Enontekiö, Porojärvet, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.433332&amp;materialsCitation.latitude=69.2" title="Search Plazi for locations around (long 21.433332/lat 69.2)">Toskalharji, Toskaljärvi N</a>, fell, dolomite rock outcrop, SE-slope, on dolomite pebbles, 720 m a. s. l., 69°12'N, 21°26'E, 2. 8. 2011, J. Pykälä 43279 (H) ; • Enontekiö, Kilpisjärvi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.85&amp;materialsCitation.latitude=69.03333" title="Search Plazi for locations around (long 20.85/lat 69.03333)">Saana Nature Reserve, E-part</a>, fell, steep SW-slope, on dolomite boulder, 910 m a. s. l., 69°02'N, 20°51'E, 13. 8. 2011, J. Pykälä 44314 (H) .</p></div>	https://treatment.plazi.org/id/A170C9C68A7653FB98BA14A165D0344F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pykälä, Juha;Myllys, Leena	Pykälä, Juha, Myllys, Leena (2025): Unexpected species richness of the lichen genus Protoblastenia (Lecanorales, Psoraceae) in Finland. MycoKeys 124: 193-226, DOI: 10.3897/mycokeys.124.162802
