taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
9A29DAAEEC315C889DC8F8B99290810C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401745	https://doi.org/10.3897/asp.83.e150460.figure3	Figure 3. Thulyphoribius melitense gen. nov. et sp. nov., head and details of the oral cavity armature (OCA) and cuticle ornamentation. A Head with protruded buccal cone, showing the papular peribuccal lobes. B Detail of the mouth opening, showing the continuous buccal ring and the second band of teeth of the OCA. C – D Details of the buccal opening, showing the continuous buccal ring, dorsal (C) and ventral (D) teeth of the third band of the OCA. E Ornamented cuticle covered with small tubercles approximately 0.2 microns in size. Arrowheads indicate the second band of teeth, indented arrowheads indicate the third band of teeth, and asterisks indicate the papular peribuccal lobes. Scale bars = A: 10 μm; B: 2 μm; C – E: 5 μm.	Figure 3. Thulyphoribius melitense gen. nov. et sp. nov., head and details of the oral cavity armature (OCA) and cuticle ornamentation. A Head with protruded buccal cone, showing the papular peribuccal lobes. B Detail of the mouth opening, showing the continuous buccal ring and the second band of teeth of the OCA. C – D Details of the buccal opening, showing the continuous buccal ring, dorsal (C) and ventral (D) teeth of the third band of the OCA. E Ornamented cuticle covered with small tubercles approximately 0.2 microns in size. Arrowheads indicate the second band of teeth, indented arrowheads indicate the third band of teeth, and asterisks indicate the papular peribuccal lobes. Scale bars = A: 10 μm; B: 2 μm; C – E: 5 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
9A29DAAEEC315C889DC8F8B99290810C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401746	https://doi.org/10.3897/asp.83.e150460.figure4	Figure 4. Thulyphoribius melitense gen. nov. et sp. nov. bucco-pharyngeal apparatus under PCM. A Bucco-pharyngeal apparatus in toto, lateral view. B Detail of the buccal opening, showing the apparent lamellae. C Stylet furcae. D Detail of the first portion of the buccal tube, highlighting the distinctive ventral lamina. E. Placoids. Arrowheads indicate the notch in the middle portion of the ventral lamina. Indented arrowheads indicate a septum that, under PCM, gives the impression of peribuccal lamellae surrounding the mouth opening. Scale bars = A, E: 20 μm; B, C: 10 μm.	Figure 4. Thulyphoribius melitense gen. nov. et sp. nov. bucco-pharyngeal apparatus under PCM. A Bucco-pharyngeal apparatus in toto, lateral view. B Detail of the buccal opening, showing the apparent lamellae. C Stylet furcae. D Detail of the first portion of the buccal tube, highlighting the distinctive ventral lamina. E. Placoids. Arrowheads indicate the notch in the middle portion of the ventral lamina. Indented arrowheads indicate a septum that, under PCM, gives the impression of peribuccal lamellae surrounding the mouth opening. Scale bars = A, E: 20 μm; B, C: 10 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
9A29DAAEEC315C889DC8F8B99290810C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401747	https://doi.org/10.3897/asp.83.e150460.figure5	Figure 5. Thulyphoribius melitense gen. nov. et sp. nov. – claws under phase contrast microscopy (PCM) and scanning electron microscopy (SEM). A, B Second pair of claws shown under PCM in Hoyer’s medium (A) and SEM (B). C Detail of the base of the external claw of the second pair of legs, shown under PCM, highlighting the internal septa. D, E Third pair of claws shown under PCM in Hoyer’s medium (D) and SEM (E). F, G Fourth pair of claws shown under PCM in Polyvinil Lactophenol medium (F) and SEM (G). Black asterisks indicate the base equipped with internal septa. White asterisks indicate the “ hump ” in the internal and anterior claws. Indented empty arrowheads indicate the cuticular bar at the base of the claws. Indented arrowheads indicate the small accessory points. Scale bars = A, B, D, E, G: 10 μm; C: 2 μm; F: 20 μm.	Figure 5. Thulyphoribius melitense gen. nov. et sp. nov. – claws under phase contrast microscopy (PCM) and scanning electron microscopy (SEM). A, B Second pair of claws shown under PCM in Hoyer’s medium (A) and SEM (B). C Detail of the base of the external claw of the second pair of legs, shown under PCM, highlighting the internal septa. D, E Third pair of claws shown under PCM in Hoyer’s medium (D) and SEM (E). F, G Fourth pair of claws shown under PCM in Polyvinil Lactophenol medium (F) and SEM (G). Black asterisks indicate the base equipped with internal septa. White asterisks indicate the “ hump ” in the internal and anterior claws. Indented empty arrowheads indicate the cuticular bar at the base of the claws. Indented arrowheads indicate the small accessory points. Scale bars = A, B, D, E, G: 10 μm; C: 2 μm; F: 20 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
9A29DAAEEC315C889DC8F8B99290810C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401743	https://doi.org/10.3897/asp.83.e150460.figure1	Figure 1. Mitogenome and phylogenetic position of Thulyphoribius melitense gen. nov. et sp. nov. A Mitogenome visualization: inner circle represents GC content. Arrows indicate direction of transcription. B Bayesian phylogenetic reconstruction of Isohypsibioidea: values above branches represent BI posterior probabilities (pp) / ML bootstrap (BS). Nodes with pp < 0.70 were collapsed. Scale bar indicates substitutions / site.	Figure 1. Mitogenome and phylogenetic position of Thulyphoribius melitense gen. nov. et sp. nov. A Mitogenome visualization: inner circle represents GC content. Arrows indicate direction of transcription. B Bayesian phylogenetic reconstruction of Isohypsibioidea: values above branches represent BI posterior probabilities (pp) / ML bootstrap (BS). Nodes with pp < 0.70 were collapsed. Scale bar indicates substitutions / site.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
9A29DAAEEC315C889DC8F8B99290810C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401748	https://doi.org/10.3897/asp.83.e150460.figure6	Figure 6. Thulyphoribius melitense gen. nov. et sp. nov. - hatchling under Phase Contrast Microscope (PCM) in Polyvinil Lactophenol medium. A Habitus. B Bucco-pharyngeal apparatus in toto, lateral view. C Third pair of claws with barely visible cuticular bar under their base. D Fourth pair of claws. Scale bars = A: 50 μm; B – D: 10 μm.	Figure 6. Thulyphoribius melitense gen. nov. et sp. nov. - hatchling under Phase Contrast Microscope (PCM) in Polyvinil Lactophenol medium. A Habitus. B Bucco-pharyngeal apparatus in toto, lateral view. C Third pair of claws with barely visible cuticular bar under their base. D Fourth pair of claws. Scale bars = A: 50 μm; B – D: 10 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
829F0FE6A46E50EF81AEFB5FE3734089.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401744	https://doi.org/10.3897/asp.83.e150460.figure2	Figure 2. Thulyphoribius melitense gen. nov. et sp. nov. – habitus seen (A) under Phase Contrast Microscopy in Hoyer’s medium (PCM) and (B) Scanning Electron Microscopy (SEM). Scale bars = A 100 μm; B: 50 μm.	Figure 2. Thulyphoribius melitense gen. nov. et sp. nov. – habitus seen (A) under Phase Contrast Microscopy in Hoyer’s medium (PCM) and (B) Scanning Electron Microscopy (SEM). Scale bars = A 100 μm; B: 50 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
829F0FE6A46E50EF81AEFB5FE3734089.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401745	https://doi.org/10.3897/asp.83.e150460.figure3	Figure 3. Thulyphoribius melitense gen. nov. et sp. nov., head and details of the oral cavity armature (OCA) and cuticle ornamentation. A Head with protruded buccal cone, showing the papular peribuccal lobes. B Detail of the mouth opening, showing the continuous buccal ring and the second band of teeth of the OCA. C – D Details of the buccal opening, showing the continuous buccal ring, dorsal (C) and ventral (D) teeth of the third band of the OCA. E Ornamented cuticle covered with small tubercles approximately 0.2 microns in size. Arrowheads indicate the second band of teeth, indented arrowheads indicate the third band of teeth, and asterisks indicate the papular peribuccal lobes. Scale bars = A: 10 μm; B: 2 μm; C – E: 5 μm.	Figure 3. Thulyphoribius melitense gen. nov. et sp. nov., head and details of the oral cavity armature (OCA) and cuticle ornamentation. A Head with protruded buccal cone, showing the papular peribuccal lobes. B Detail of the mouth opening, showing the continuous buccal ring and the second band of teeth of the OCA. C – D Details of the buccal opening, showing the continuous buccal ring, dorsal (C) and ventral (D) teeth of the third band of the OCA. E Ornamented cuticle covered with small tubercles approximately 0.2 microns in size. Arrowheads indicate the second band of teeth, indented arrowheads indicate the third band of teeth, and asterisks indicate the papular peribuccal lobes. Scale bars = A: 10 μm; B: 2 μm; C – E: 5 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
829F0FE6A46E50EF81AEFB5FE3734089.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401746	https://doi.org/10.3897/asp.83.e150460.figure4	Figure 4. Thulyphoribius melitense gen. nov. et sp. nov. bucco-pharyngeal apparatus under PCM. A Bucco-pharyngeal apparatus in toto, lateral view. B Detail of the buccal opening, showing the apparent lamellae. C Stylet furcae. D Detail of the first portion of the buccal tube, highlighting the distinctive ventral lamina. E. Placoids. Arrowheads indicate the notch in the middle portion of the ventral lamina. Indented arrowheads indicate a septum that, under PCM, gives the impression of peribuccal lamellae surrounding the mouth opening. Scale bars = A, E: 20 μm; B, C: 10 μm.	Figure 4. Thulyphoribius melitense gen. nov. et sp. nov. bucco-pharyngeal apparatus under PCM. A Bucco-pharyngeal apparatus in toto, lateral view. B Detail of the buccal opening, showing the apparent lamellae. C Stylet furcae. D Detail of the first portion of the buccal tube, highlighting the distinctive ventral lamina. E. Placoids. Arrowheads indicate the notch in the middle portion of the ventral lamina. Indented arrowheads indicate a septum that, under PCM, gives the impression of peribuccal lamellae surrounding the mouth opening. Scale bars = A, E: 20 μm; B, C: 10 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
829F0FE6A46E50EF81AEFB5FE3734089.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401747	https://doi.org/10.3897/asp.83.e150460.figure5	Figure 5. Thulyphoribius melitense gen. nov. et sp. nov. – claws under phase contrast microscopy (PCM) and scanning electron microscopy (SEM). A, B Second pair of claws shown under PCM in Hoyer’s medium (A) and SEM (B). C Detail of the base of the external claw of the second pair of legs, shown under PCM, highlighting the internal septa. D, E Third pair of claws shown under PCM in Hoyer’s medium (D) and SEM (E). F, G Fourth pair of claws shown under PCM in Polyvinil Lactophenol medium (F) and SEM (G). Black asterisks indicate the base equipped with internal septa. White asterisks indicate the “ hump ” in the internal and anterior claws. Indented empty arrowheads indicate the cuticular bar at the base of the claws. Indented arrowheads indicate the small accessory points. Scale bars = A, B, D, E, G: 10 μm; C: 2 μm; F: 20 μm.	Figure 5. Thulyphoribius melitense gen. nov. et sp. nov. – claws under phase contrast microscopy (PCM) and scanning electron microscopy (SEM). A, B Second pair of claws shown under PCM in Hoyer’s medium (A) and SEM (B). C Detail of the base of the external claw of the second pair of legs, shown under PCM, highlighting the internal septa. D, E Third pair of claws shown under PCM in Hoyer’s medium (D) and SEM (E). F, G Fourth pair of claws shown under PCM in Polyvinil Lactophenol medium (F) and SEM (G). Black asterisks indicate the base equipped with internal septa. White asterisks indicate the “ hump ” in the internal and anterior claws. Indented empty arrowheads indicate the cuticular bar at the base of the claws. Indented arrowheads indicate the small accessory points. Scale bars = A, B, D, E, G: 10 μm; C: 2 μm; F: 20 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
829F0FE6A46E50EF81AEFB5FE3734089.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401748	https://doi.org/10.3897/asp.83.e150460.figure6	Figure 6. Thulyphoribius melitense gen. nov. et sp. nov. - hatchling under Phase Contrast Microscope (PCM) in Polyvinil Lactophenol medium. A Habitus. B Bucco-pharyngeal apparatus in toto, lateral view. C Third pair of claws with barely visible cuticular bar under their base. D Fourth pair of claws. Scale bars = A: 50 μm; B – D: 10 μm.	Figure 6. Thulyphoribius melitense gen. nov. et sp. nov. - hatchling under Phase Contrast Microscope (PCM) in Polyvinil Lactophenol medium. A Habitus. B Bucco-pharyngeal apparatus in toto, lateral view. C Third pair of claws with barely visible cuticular bar under their base. D Fourth pair of claws. Scale bars = A: 50 μm; B – D: 10 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
829F0FE6A46E50EF81AEFB5FE3734089.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401749	https://doi.org/10.3897/asp.83.e150460.figure7	Figure 7. Thulyphoribius melitense gen. nov. et sp. nov. – habitus and details of the legs under SEM (Stub No. UNICT. 39). A habitus, ventral view. B First pair of legs. C Second pair of legs. D Third pair of legs. Arrowheads indicate the botton-like structures in the dorsal portion of the leg. Indented arrowhead indicates the small accessory points. Scale bars = A: 100 μm; B – D: 10 μm.	Figure 7. Thulyphoribius melitense gen. nov. et sp. nov. – habitus and details of the legs under SEM (Stub No. UNICT. 39). A habitus, ventral view. B First pair of legs. C Second pair of legs. D Third pair of legs. Arrowheads indicate the botton-like structures in the dorsal portion of the leg. Indented arrowhead indicates the small accessory points. Scale bars = A: 100 μm; B – D: 10 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
829F0FE6A46E50EF81AEFB5FE3734089.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1401750	https://doi.org/10.3897/asp.83.e150460.figure8	Figure 8. Thulyphoribius melitense gen. nov. et sp. nov. – cysts seen under Phase Contrast Microscope (PCM). A specimen outside its cyst. B Cyst with specimen inside, where buccal apparatus is barely visible at the centre of the cyst. Scale bars 100 μm.	Figure 8. Thulyphoribius melitense gen. nov. et sp. nov. – cysts seen under Phase Contrast Microscope (PCM). A specimen outside its cyst. B Cyst with specimen inside, where buccal apparatus is barely visible at the centre of the cyst. Scale bars 100 μm.	2025-08-27	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo		Zenodo	biologists	Camarda, Daniele;Lisi, Oscar;Stec, Daniel;Vecchi, Matteo			
