taxonID	type	description	language	source
CD54F960FFD2FFA55153A82E3E0D4337.taxon	materials_examined	Material. Andreevskii Cordon, 18 – 30.10. – 1 – 9.11.2023, + VO (every night) – 68 ♂♂, 13 ♀♀; ibid., plateau, in light trap, 23 – 24.10.2023 – 5 ♂; ibid., ridge, in bottle, 24 – 25.10.2023 – 1 ♂ (Zinchenko); Kaldernyi Cordon, at light, 30 – 31.10.2023 – 7 ♂; ibid., 31.10 – 1.11.2023 – 2 ♂ + VO.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD2FFA55153A82E3E0D4337.taxon	distribution	Distribution. Russia (European part, Crimea, Urals, S RFE: Amur Obl., Jewish A. O., S Khabarovsk Kr., Primorsky Kr., Sakhalin, S Kurils – Kunashir); Europe, North Africa, Turkey, Transcaucasia, Turkmenistan, Northeast China, South Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu), North America (introduced?).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD2FFA55153A82E3E0D4337.taxon	discussion	Remarks. This species is represented by the East Asian subspecies C. pennaria ussuriensis O. Bang-Haas, 1927. Flight period of this species on Kunashir Island extends from mid-October to early November, considering the previous data (Beljaev et al. 2023). In the continental part of the RFE, moths are typically observed from mid-September to the end of October. The larvae feed on a various leaved woody plants.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD5FFA45153A94D3B0A4446.taxon	materials_examined	Material. Andreevskii Cordon: 13 – 14.10.2023 – 1 ♀, 15 – 16.10.2023 – 1 ♂, 16 – 17.10.2023 – 2 ♂, 1 ♀, 19 – 20.10.2023 – 2 ♀ + VO, 20 – 21.10.2023 – 1 ♀.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD5FFA45153A94D3B0A4446.taxon	distribution	Distribution. Russia (S RFE: SW Sakhalin, S Kurils — Kunashir; Japan (Hokkaido, Honshu, Shikoku, Kyushu).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD5FFA45153A94D3B0A4446.taxon	discussion	Remarks. Flight period of G. flavolinearia on Kunashir Island extends from the second week of September to the end of October, considering the previous data (Rybalkin 2020; Beljaev et al. 2023). On Sakhalin Island, the species is observed during the first half of September (Beljaev, Titova 2023). In Japan, the larvae feed on woody Rosaceae species (Sato 2011).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD4FFA452D1AE1F38E7435C.taxon	materials_examined	Material. Andreevskii Cordon: 26 – 27.10. 2023 – 1 ♂, 1 – 23.11.2023 (every night) – 103 ♂; ibid., plateau: 5 – 6.11., 8 – 11.11., 13 – 15.11., 16 – 21.11. 2023, in various localities of mixed forest – 96 ♂♂. Manual collection with a net: right bank of the river Andreyevka, road in alder forest, 21.11.2023 – 1 ♂; ibid., plateau, birch forest, 21.11.2023 (Zinchenko) – 1 ♂; ibid., climb to plateau and plateau, evening catch, 22.11.2023 (Dubatolov, Zinchenko) – 3 ♂; Andreevskii Cordon, ibid., ridge and Sasa meadow on plateau, 16.11.2023 (Zinchenko) – 3 ♂; ibid., plateau, mixed forest, at night, 18.11.2023 (Zinchenko) – 2 ♂; ibid., plateau, birch forest, 19.11.2023 (Zinchenko) – 1 ♂; near Andreevskii Cordon, by net, 16.11.2023 (Zinchenko) – 3 ♂; ibid., plateau, mixed forest, at night, 13.11.2023 (Zinchenko) – 4 ♂.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD4FFA452D1AE1F38E7435C.taxon	distribution	Distribution. Russia (S Buryatia,? S Irkutskaya Obl., S Amur Obl., Jewish A. O., S Khabarovsk Kr., Primorsky Kr., Sakhalin, S Kurils – Kunashir); Japan (Hokkaido, Honshu, Shikoku, Kyushu, Okinawa Island), South Korea, China (Central, Taiwan Island).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD4FFA452D1AE1F38E7435C.taxon	discussion	Remarks. This species, new for the Kuril Islands, was observed on Kunashir from late October to mid-November. In Japan, moths appear from early November to late December (Sato 2011). In the continental part of the RFE, fly of the species is observed from late September to late October (Beljaev 1996). The larvae feed on various leaved woody plants.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD4FFA752D1A92438CB4020.taxon	materials_examined	Material. Kaldernyi Cordon, at light (near the morning), 31.10 – 1.11.2023 – 1 ♂.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD4FFA752D1A92438CB4020.taxon	distribution	Distribution. Russia (Altai, Tyva, Irkutskaya Obl., Buryatia, RFE: Amur Obl., Jewish A. O., Khabarovsk Kr., Primorsky Kr., Sakhalin, S Kurils – Kunashir); E Kazakhstan, Mongolia, Northeast China, North Korea, Japan (Hokkaido, Honshu, Shikoku).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD4FFA752D1A92438CB4020.taxon	discussion	Remarks. This species, newly recorded for the Kuril Islands, is represented on Kunashir by the subspecies E. jacobsoni giganthea Inoue, 1955, also found in Sakhalin and Japan. In Japan, the moths appear from late October to late November. In Hokkaido it is commonly found in large numbers in larch forests and is sometimes considered a pest (Sato 2011). In the continental part of the RFE, moths are observed from late September to late October (Beljaev 1996). The larvae feed on a variety of coniferous species (Pinaceae). The status of the taxon giganthea has been subject to debate. Initially Inoue described it as a valid species. In the description he noted that it is " very close related to jacobsoni Djakonov from the Central Altai …, but larger in size and it appears in autumn while jacobsoni … wing in spring " (Inoue 1955: 76). Later, gigantea was downgraded to a subspecies of Erannis defoliaria (Clerck, 1759), based on the similarities in the appearance of moths and in the shape of the valva in the male genitalia (Inoue 1982). This view has been widely accepted in Japanese literature and on the Internet sites (Nakajima 1998; Niwakagamania et al. 2024). In contrast, Beljaev (Beljaev 1996) proposed recognizing gigantea as a subspecies of Erannis jacobsoni (Djakonov, 1926), based on the almost complete similarity of the wing pattern and coloration with the Far Eastern subspecies Erannis jacobsoni sichotenaria Kurentzov, 1937. This approach has been adopted in Russian literature, including taxonomic catalogs (Mironov et al. 2008; Beljaev 2016; Beljaev, Mironov 2019). Nakajima (Nakajima 2010) restored gigantea to the species rank, based on a comparison of the male genitalia of Japanese specimens with the European E. defoliaria only, without comparing them to E. jacobsoni. He questioned the Beljaev's decisions, citing the fact that he had not examined the type specimen of E. jacobsoni, and that moths of the later emerge in spring. According to the original description, the type series of E. jacobsoni (described in ‡ Hibernia, a misspelling of Hybernia Berthold, 1827) consist of three males and one female. The holotype is not designated. The male genitalia of one syntype were depicted in the original description of the species (Djakonov 1926), and the photo of the other male syntype was published later (Djakonov 1929). These data confirm the identity of E. jacobsoni. However, the published collection date of the type specimens — 30 June 1898, which was one of the reasons for the misinterpretation of giganthea, — does not correspond to the real time of emerge of moths of E. jacobsoni, which occurs in autumn. Since The type series includes one male and one female, reared from larvae. Therefore, the date is likely to refer to the larvae collection date, which is quite real, and not the capture of adult moths. A comparison of the male genitalia of E. jacobsoni jacobsoni from Altai (Djakonov 1926) and Buryatia (Beljaev 1996; Makhov 2021), E. jacobsoni sichotenaria from Primorsky Kr. (Beljaev 1996) and North Korea (Tóth 2017), and gigantea from Japan (Nakajima 1998; Nakajima 2010) shows their structural similarities, making it impossible to distinguish morphologically the latter taxon as a separate species. Therefore, at this stage of research, it is most appropriate to retain the status of gigantea as a subspecies of E. jacobsoni. The Online Taxonomic Facility of Geometridae (Rajaei et al. 2022) recognizes the taxon E. jacobsoni gigantea, while the Global Biodiversity Information Facility lists it as both E. gigantea (Erannis gigantea … 2024), and as E. jacobsoni subsp. gigantea (Erannis jacobsoni … 2024).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA75153AD3839EA421C.taxon	materials_examined	Material. Kaldernyi Cordon, at light: 30 – 31.10.2023 – 1 ♂, 31.10 – 1.11.2023 – 1 ♂; Andreevskii Cordon: 3 – 4.11.2023 – 1 ♂, 7 – 8.11.2023 – 1 ♂, 13 – 14.11.2023 – 3 ♂, 14 – 15.11.2023 – 4 ♂, 16 – 17.011.2023 – 6 ♂, 18 – 19.11.2023 – 2 ♂; 2 ♂, 19 – 20.11.2023 – 2 ♂; ibid., plateau, different localities, in light trap: 8 – 9.11.2023 – 6 ♂, 13 – 14.11.2023 – 4 ♂, 16 – 17.11.2023 – 2 ♂, 17 – 18.11.2023 – 1 ♂, 20 – 21.11.2023 – 1 ♂; ibid., ridge, at night, 13.11.2023 (Zinchenko) – 1 ♂; ibid., plateau, in the nearby forest at night, by the mowing on the grass layer, 18.11.2023 (Zinchenko) – 1 ♀.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA75153AD3839EA421C.taxon	distribution	Distribution. Russia (European Part (Moskovskaya Obl., introduced), S RFE: S Amur Obl., Jewish A. O., S Khabarovsk Kr., Primorsky Kr., SW Sakhalin, S Kurils – Kunashir), Japan (Hokkaido, Honshu), South Korea,? North China.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA75153AD3839EA421C.taxon	discussion	Remarks. This species is reported for the first time from the Kuril Islands. On Kunashir, the moths were observed from late October to mid-November, while on Sakhalin, there was only one moth recorded in mid-October (Beljaev, Titova 2023). In Hokkaido, L. orthogrammaria flies during October (Sato 2011). In the continental part of the RFE, the moths are observed also throughout October (Beljaev 1996). The larvae feed on various leaved woody plants.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA752D1AA393F064720.taxon	materials_examined	Material. Kaldernyi Cordon, in the daytime: 30.10.2023 – 3 ♂, 31.10.2023 – 7 ♂, 1.11.2023 – 3 ♂ (Dubatolov, Zinchenko); ibid., at light, 30 – 31.10.2023 – 2 ♂, 31.10 – 1.11.2023 – 1 ♂; Andreevskii Cordon, at light: 1 – 2.11.2023 – 1 ♂, 3 – 4.11.2023 – 3 ♂, 4 – 5.11.2023 – 2 ♂, 5 – 6.11.2023 – 2 ♂, 6 – 7.11.2023 – 1 ♂ + VO, 7 – 8.11.2023 – 1 ♂; ibid., plateau, mixed forest, in the daytime, 18.11.2023 (Zinchenko) – 1 ♂.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA752D1AA393F064720.taxon	distribution	Distribution. Russia (S RFE: S Amur Obl., Jewish A. O., S Khabarovsk Kr., Primorsky Kr., SW Sakhalin, S Kurils – Kunashir); Japan (Hokkaido, Honshu, Shikoku, Kyushu), North Korea, South Korea.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA752D1AA393F064720.taxon	discussion	Remarks. The flight period of moths of P. obliquaria on Kunashir was observed from late October to mid-November, while on Sakhalin they were noted earlier, in the last ten days of October (Beljaev, Titova 2023). In Japan, the moths appear from late October to late December, depending on the region, and the species is usually numerous in various habitats (Sato 2011). In the continental part of the RFE, the moths are observed throughout October (Beljaev 1996). The larvae feed on various leaved woody plants, with a preference for species of Quercus (Fagaceae).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA951EBA86438E74381.taxon	materials_examined	Material. Andreevskii Cordon, 5 – 6.11.2023 – 1 ♂.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA951EBA86438E74381.taxon	distribution	Distribution. Russia (S RFE: S Kurils – Kunashir); Japan (Hokkaido, Honshu).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA951EBA86438E74381.taxon	discussion	Remarks. This species is newly record- ed for both the Kuril Islands and Russia as a whole. In Japan in Honshu, in the Kanto region, the species is distributed in the mountains between 700 and 1000 m a. s. l., where moths appear from late November to early December. The species coexists with Alsophila japonensis, but habitat of A. inouei is restricted to a narrow range of high altitudes, and its flight period is short (Nakajima 2011). Larvae of A. inouei has been observed feeding on Alnus hirsuta (Betulaceae) (Nakajima 2011), though it is likely to be polyphagous on various leaved woody plants. * Alsophila japonensis (Warren, 1894) (Figs 3 D, 4 C)	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA951EBA86438E74381.taxon	materials_examined	Material. Andreevskii Cordon, at light: 10 – 11.11.2023 – 3 ♂, 13 – 14.11.2023 – 9 ♂, 15 – 16.11.2023 – 4 ♂, 16 – 17.11.2023 – 1 ♂, 18 – 19.11.2023 – 11 ♂, 19 – 20.11.2023 – 4 ♂, 20 – 21.11.2023 – 1 ♂, 21 – 22.11.2023 – 2 ♂, 22 – 23.11.2023 – 1 ♂; ibid., road and nearby forest on the plateau, at night, 15.11.2023 – 1 ♂ (Dubatolov, Zinchenko); ibid., plateau, in light trap: 16 – 17.11.2023 – 5 ♂, 18 – 19.11.202 – 32 ♂; ibid., plateau, 15.11.2023 (Dubatolov, Zinchenko) – 1 ♂; ibid., road and nearby forest on the plateau, at night, 19.11.2023 (Zinchenko) – 1 ♂.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA951EBA86438E74381.taxon	distribution	Distribution. Russia (S RFE: S Khabarovsk Kr., Primorsky Kr., SW Sakhalin, S Kurils – Kunashir); Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima, Yakushima), South Korea, China (Northeast, Central: Jiangsu).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD7FFA951EBA86438E74381.taxon	discussion	Remarks. The species is new for the Kuril Islands. The collected specimens belong to the nominate Japanese subspecies A. japonensis japonensis, which is also found on Sakhalin. On Kunashir, the moths fly from the second week until the end of November, whereas on Sakhalin, the species was recorded from the mid- to late October (Beljaev, Titova 2023). In Japan, A. japonensis emerges in Hokkaido and in mountains of Honshu from late October to mid-November, and in plains of the latter island from December to early February (Nakajima 2011). In the continental part of the RFE, the moths are observed from mid-October to the first part of November (Beljaev 1996). The larvae feed on various leaved woody plants.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD9FFA952D1A9D73F2B43F4.taxon	materials_examined	Material. Surroundings of Andreevskii Cordon, road on a slope, on alder bark, 18.11.2023 – 1 ♂, 1 ♀ in copula; Andreevskii Cordon, at light, between 1 and 2 am, 19 – 20.11.2023 – 1 ♂; ibid., 21 – 22.11.2023 – 1 ♂; ibid., 22 – 23.11.2023 – 1 ♂; ibid., the beginning of the trailway near the cordon, alder forest, 20.11.2023 (Zinchenko) – 1 ♂; ibid., road on a slope, alder forest, 21.11.2023 – 6 ♂; ibid., plateau, nearby mixed forest, at night, 18.11.2023 – 1 ♂; ibid., 19.11.2023 (Zinchenko) – 2 ♂; ibid., 21.11.2023 (Zinchenko) – 3 ♂; ibid., ridge, mixed forest, 19.11.2023 – 2 ♂ (Zinchenko); ibid., plateau, at night, 22.11.2023 (Dubatolov, Zinchenko) – 91 ♂; ibid., plateau, in the daytime, 22.11.2023 (Dubatolov, Zinchenko) – 28 ♂. Photo of moths: Yuzhno-Kurilsk, 27 – 29.11.2023 (Linnik). Video of moths: 23 km NE of Yuzhno-Kurilsk, Filatovka river, 44 ° 11 ' 47 " N, 146 ° 01 ' 09 " E, 30.11.2023 (Linnik). Photos of moths: 6 km NW of Yuzhno-Kurilsk, Lake Lagunnoe, 44 ° 03 ′ 46 ″ N, 145 ° 45 ′ 36 ″ E, 2.12.2023 (Stefanov).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD9FFA952D1A9D73F2B43F4.taxon	distribution	Distribution. Russia (S RFE: SW Sakhalin, S Kurils – Kunashir), Japan (Hokkaido, Honshu, Shikoku, Kyushu).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD9FFA952D1A9D73F2B43F4.taxon	discussion	Remarks. The species is new for the Kuril Islands. On Kunashir, the moths flied from the second half of November to early December and were most numerous in forests in late November. The female (Fig. 4 E) was found together with a male on the trunk of an alder tree on 18 November. On Sakhalin, moths of this species were recorded earlier, during the first half of November (Beljaev, Titova 2023). In Japan, I. asahinai inhabits the plains on Hokkaido and mountainous areas on Honshu to Kyushu. In the Kanto region, it appears from mid-November to December, with the peak flight period occurring around mid-November (Nakajima 2011). The larvae are believed to be feeding on various leaved woody plants, similar to other species of Inurois. In 2024, in the Prozrachny Stream valley (9 km north of Yuzhno-Kurilsk, 44 ° 11 ' 47 " N, 146 ° 01 ' 09 " E), Mikhail Ragimov observed a mass flight of I. asahinai (“ thousands of moths ”) on 20 November shortly after the first snowfall.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD9FFA85153A98C38E74159.taxon	materials_examined	Material. Andreevskii Cordon, at light, 21.11.2023 – 1 ♂; ibid., 22 – 23.11.2023 – 3 ♂; ibid., road on a slope, alder forest, at evening by net, 21.11.2023 – 2 ♂; ibid., plateau, birch forest, at night, 21.11.2023 (Zinchenko) – 6 ♂; ibid., at night, 22.11.2023 – 25 ♂ (Dubatolov, Zinchenko). Photos of moths: 6 km NW of Yuzhno-Kurilsk, Lagunnoe Lake, 44 ° 03 ′ 46 ″ N, 145 ° 45 ′ 36 ″ E, 2.12.2023 (Stefanov).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD9FFA85153A98C38E74159.taxon	distribution	Distribution. Russia (S RFE: S Amur Obl., Jewish A. O., S Khabarovsk Kr., Primorsky Kr., S Kurils – Kunashir); South Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu), China (Shaanxi,? Sichuan, Taiwan Island) (for the details see Beljaev 2022).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD9FFA85153A98C38E74159.taxon	discussion	Remarks. The species is new for the Kuril Islands. On Kunashir, the moths were observed in the last third of November and early December, appearing later than other winter geometrids, with the exception of Inurois punctigera. In Japan, in the plains and low mountain areas of the Kanto region (Honshu Island) I. fumosa appears from late December to late January, with a peak flight in mid-January, whereas in mountains it flies from mid-November to early December (Nakajima 2011). In the continental part of the RFE, the moths are observed from early October (in the north of its range and in mountains) to mid-November (in south) (Beljaev 1996). The larvae feed on various leaved woody plants.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD8FFA852D1AB1F3EE34460.taxon	materials_examined	Material. Plateau, birch and broad-leaved forest, at night, 21.11.2023 (Zinchenko) – 8 ♂; ibid., plateau, birch and broad-leaved forest, evening catch, 22.11.2023 (Zinchenko) – 10 ♂; plateau, broad-leaved forest, at late evening, 23.11.2023 – 3 ♂.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD8FFA852D1AB1F3EE34460.taxon	distribution	Distribution. Russia (S RFE: S Kurils – Kunashir); Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima), South Korea.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD8FFA852D1AB1F3EE34460.taxon	discussion	Remarks. It is a new species for both the Kuril Islands and Russia as a whole. On Kunashir, the moths were observed in the last third of November, alongside I. fumosa. In Japan, I. punctigera is vertically distributed from plains to the lower subalpine zones. Around the Kanto region, moths of this species appear from late December to March in the plains. In mountainous areas, it has two flight periods: first, from mid-November to early December, and second, from late April to early May, separated by the snow season (Nakajima 1998, 2011 [as Inurois membranaria]; Yamamoto, Sota 2009). The larvae feed on various leaved woody plants. It should be noted that most Japanese authors, starting with Inoue (Inoue 1986), have erroneously identified Inurois punctigera as Inurois membranaria (nec Christoph, 1881). For further details, see Beljaev (Beljaev 2022).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD8FFA85153AECB3E3A4008.taxon	materials_examined	Material. Andreevskii Cordon, 7 – 8.11.2023 – 1 ♀; ibid., alder forest, at bait traps, 7.11.2023 – 1 ♂, 1 ♀; ibid., plateau, mixed forest, on birch, 7 – 8.11.2023 (Zinchenko) – 1 ♂.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD8FFA85153AECB3E3A4008.taxon	distribution	Distribution. Russia (European part, N Caucasus, Urals, South Siberia, S RFE: Amur Obl., Khabarovsk Kr., Primorsky Kr., Sakhalin, S Kurils — Kunashir); almost cosmopolitan, except most northern and most southern territories, and Australia. Migrant.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD8FFA85153AECB3E3A4008.taxon	discussion	Remarks. This is the first record of O. obstipata in November, later than its previous record in Kunashir in October (Beljaev et al. 2023), and also the latest for the entire Russian Far East. In Japan, it is distributed almost throughout the country, being common in various habitats from plains to mountains. It appears around the Kanto region from late March to early November, and is thought to be polyvoltine (Nakajima, Yazaki 2011). The larvae are polyphagous, feeding on herbs, shrubs, and trees.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD8FFAB5153AA503B9E4446.taxon	materials_examined	Material. Andreevskii Cordon, at bait traps, 18.10.2023 – 1 ♀; ibid., at light, 22 – 23.10.2023 – 1 ♀; ibid., plateau, birch forest, in bottle, 24 – 25.10.2023 (Zinchenko) – 2 ♀.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD8FFAB5153AA503B9E4446.taxon	distribution	Distribution. Distribution. Russia (S RFE: SW Primorsky Kr., S Kurils — Kunashir); Northeast China (Jilin), South Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu, Yakushima).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFD8FFAB5153AA503B9E4446.taxon	discussion	Remarks. This species was first recorded for the Kuril Islands by Beljaev et al. (Beljaev et al. 2023) based on a single female collect- ed at the same location at approximately the same time, i. e., 18 – 19 October 2022. In most of Japan Ph. lucicolens develops in two generations, moths fly in June and July, and in October, and very common in many places (Nakajima, Yazaki 2011). It cannot be ruled out that the collected females migrated from Hokkaido Island, as moths of the first generation have never been recorded on the Kuril Islands. The larvae feed on various species of Quercus (Fagaceae).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAB52D1A9D739AE4720.taxon	materials_examined	Material. Surroundings of Andreevskii Cordon, at night in the forest, 17.10.2023 – 1 ♀; Andreevskii Cordon, 19 – 20.10.2023 – 1 ♀; ibid., 28 – 29.10.2023 – 1 ind. VO.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAB52D1A9D739AE4720.taxon	distribution	Distribution. Russia (European part, N Caucasus, Urals, West Siberia, South Siberia, Jakutia, RFE: Kamchatka Pen., Commander Islands, Amur Obl., S Khabarovsk Kr., Primorsky Kr., Sakhalin, Kurils – Shumshu, Paramushir, Simushir, Urup, Iturup, Kunashir, Shikotan); Europe, Turkey, Transcaucasia, N Kazakhstan, Mongolia, China, North Korea, South Korea, Japan, N India, North America.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAB52D1A9D739AE4720.taxon	discussion	Remarks. The taxonomic status of East Asian populations of Dysstroma, currently associated with D. citrata, needs to be revised. For more information see Beljaev (Beljaev 2016), and Beljaev, Titova (Beljaev, Titova 2023).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAB52D1AE1F3AC94381.taxon	materials_examined	Material. Andreevskii Cordon, mixed forest, on birch, 19.11.2023 (Zinchenko) – 1 ♀.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAB52D1AE1F3AC94381.taxon	distribution	Distribution. Russia (European part, N Caucasus, Urals, West Siberia, South Siberia, S Jakutia, RFE: Magadanskaya Obl., Kamchatka Pen., Amur Obl., Jewish A. O., Khabarovsk Kr., Primorsky Kr., Sakhalin, S Kurils – Kunashir); Japan (Hokkaido, Honshu, Shikoku, Kyushu), North Korea, South Korea, China, N Mongolia, Kazakhstan,? Kyrgyzstan, Turkey, Europe.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAB52D1AE1F3AC94381.taxon	discussion	Remarks. In East Asia, this species is represented by the subspecies H. impluviata extremata Bryk, 1942 (= H. impluviata insulata Inoue, 1953), described from Kunashir Island (Bryk 1942). The specimen collected on Kunashir was discovered at a time that is atypical for the flight of moths of this species. In the continental territories of the Far East at the same latitudes, this species typically flies in one generation from the end of May to mid-July. In Japan, H. impluviata inhabits mountainous areas up to the alpine zone and is usually seen from July to August, supposedly having a univoltine life cycle. In some regions, however, it can appear from May to June or July to August, exhibiting bivoltine life cycles (Nakajima, Yazaki 2011). Therefore, the specimen from Kunashir is likely an accidentally hatched individual from the wintering generation. The larvae feed on various leaved woody plants, preferring Betulaceae and Salicaceae.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAB5153AD3939C44072.taxon	materials_examined	Material. Andreevskii Cordon, at light, 16 – 17.10.2023 – 1 ♀.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAB5153AD3939C44072.taxon	distribution	Distribution. Russia (S RFE: S Khabarovsk Kr., Primorsky Kr., S Sakhalin, S Kurils – Kunashir);? South-West China (Sichuan, Yunnan), South Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAB5153AD3939C44072.taxon	discussion	Remarks. The moth was collected within the period of its previous registration on Kunashir in 2022 — from 20 September to 22 October (Beljaev et al. 2023). The flight period of P. comis on Kunashir is similar to that on Sakhalin (Beljaev, Titova 2023). In Japan, the species is relatively common from low mountain areas to alpine zones. It appears in low mountain areas from October to November and in mountain and subalpine zones from late August to September (Nakajima, Yazaki 2011). The larvae feed on various species of Abies (Pinaceae).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAA5153AA0A39894246.taxon	materials_examined	Material. Andreevskii Cordon, at light, 16 – 30.10. – 1 – 20.11.2023 (every night) – 96 ♂, 25 ♀, + numerous VO; ibid., plateau, in light trap: 23 – 24.10.2023 – 1 ♂, 23 – 24.10.2023 – 11 ♂; Andreevskii Cordon, mixed forest on the high right bank of the Andreevka river, in light trap, 26 – 27.10.2023 – 1 ♂; Lesnaya river, coniferous forest, 22.10.2023 – 1 ♂; Andreevskii Cordon, at bait traps, 23.10.2023 – 2 ♂ + VO; ibid., at bait traps, 5.11.2023 – 1 ♀, 10.11.2023 – 1 ♂; ibid., plateau, alder forest, at bait traps, 10.11.2023 – 7 ♂; ibid., plateau, birch forest, in bottle: 21 – 22.10.2023 – 2 ♂, 22 – 23.10.2023 – 2 ♂, 23 – 24.10.2023 – 2 ♂, 24 – 25.10.2023 – 5 ♂, 28 – 29.10.2023 – 1 ♂, 29 – 30.10.2023 – 1 ♂, 3 – 4.11.2023 – 1 ♂ (Zinchenko); ibid., plateau, spruce forest, in bottle, 7 – 8.11.2023 – 1 ♂ (Zinchenko); mixed forest on the high right bank of the Andreevka river, in bottle, 7 – 8.11.2023 – 2 ♂ (Zinchenko); Kaldernyi Cordon: at light – 30 – 31.10.2023 – 4 ♂, in light trap – 30 ♂, 35 ♀; ibid., 31.10 – 1.11.2023 –> 50 ind.; surroundings of Kaldernyi Cordon, road, in light trap, 30 – 31.10.2023 – 4 ♂; ibid., at bait traps, 31.10 – 1.11.2023 – 3 ♂ (Zinchenko); Kaldernyi Cordon, at bait traps, 30 – 31.10.2023 – 1 ♂.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAA5153AA0A39894246.taxon	distribution	Distribution. Russia (European part, N Caucasus, Urals, West Siberia, South Siberia, Jakutia, RFE: Magadan Obl., Kamchatka Pen., Amur Obl., Jewish A. O., Khabarovsk Kr., Primorsky Kr., Sakhalin, S Kurils — Urup, Iturup and Kunashir); Europe, Turkey, Transcaucasia, N Kazakhstan, Mongolia, China (E Inner Mongolia), South Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu),? North America.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAA5153AA0A39894246.taxon	discussion	Remarks. In East Asia, the subspecies E. autumnata autumnus (Bryk, 1942), described from Iturup (Bryk 1942), is distributed. The investigation of this species in 2023 reveals its long flight period on Kunashir, from mid-October up to the second half of November. In Japan, it appears in October (Nakajima, Yazaki 2011). In the same-latitude continental part of the RFE, the moths are observed from the mid-September to mid-October, in mountains from early September (Beljaev 1996). The larvae feed on various trees and shrubs, with a preference for species of Betulaceae and Salicaceae. * Operophtera brunnea Nakajima, 1991 (= Operophtera variabilis Nakajima, 1991) (Figs 3 H, 4 G, 4 H)	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAA5153AA0A39894246.taxon	materials_examined	Material. Kaldernyi Cordon, at light, 30 – 31.10.2023 – 1 ♂, 31.10 – 1.11.2023 – 1 ♂; surroundings of Kaldernyi Cordon, road, in light trap, 30 – 31.10.2023 – 1 ♂; Andreevskii Cordon, 3 – 23.11.2023 – 270 ♂; ibid., road on a slope, alder forest, 5.11.2023 – 1 ♂, 7.11.2023 – 1 ♂, 10.11.2023 – 2 ♂, 13.11.2023 – 1 ♂; 21.11.2023 – 4 ♂; ibid., plateau, distant forest, in light trap:, 8 – 9.11.2023 – 7 ♂, 12 – 13.11.2023 – 7 ♂, 16 – 17.11.2023 – 6 ♂; 18 – 19.11.2023 – 2 ♂; ibid., plateau, nearby mixed forest, at night: 13.11.2023 – 24 ♂, 18.11.2023 – 14 ♂, 19.11.2023 – 3 ♂, 21.11.2023 – 7 ♂ (Zinchenko); ibid., 22.11.2023 – 21 ♂ (Dubatolov, Zinchenko); ibid., plateau, nearby birch forest, in light trap, 20 – 21.11.2023 – 6 ♂; mixed forest on the high right bank of the Andreevka river, in light trap, 5 – 6.11.2023 – 1 ♂; ibid., at night, manual collection: 13.11.2023 – 15 ♂ (Zinchenko), 14.11.2023 – 1 ♂ (Dubatolov), 15.11.2023 – 22 ♂ (Dubatolov, Zinchenko); Andreevskii Cordon, in the mixed forest, on bark near bait traps, 10.11.2023 (Dubatolov) – 2 ♂; Sernovodka river, 8.11.2023 (Zinchenko) – 1 ♂; Yuzhno-Kurilsk, office of the Kurilsky Nature Reserve, in a spider's web, 24.11.2023 – 1 ♂.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAA5153AA0A39894246.taxon	distribution	Distribution. Russia (S Amur Obl., Jewish A. O., S Khabarovsk Kr., Primorsky Kr., Sakhalin, S Kurils – Kunashir); Japan (Hokkaido, Honshu, Izu Oshima, Shikoku, Kyushu), South Korea, China (Northeast, Central and Taiwan Island).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDBFFAA5153AA0A39894246.taxon	discussion	Remarks. The species is newly recorded for the Kuril Islands. The Kunashir population belongs to the nominative Japanese subspecies O. brunnea brunnea, also found on Sakhalin. On Kunashir, the moths were observed from the end of October up to the end of November. In Japan, the species is widespread across the country, from the plains to the lower subalpine zone, with moths appearing from late November to mid-February depending on the locality (Nakajima, Yazaki 2011). On the continental part of the RFE, the moths are observed from the beginning of October to the beginning of November (Beljaev 1996). The larvae feed on various leaved woody plants. It should be noted that while the synonymy of O, brunnea and O, variabilis was reasonably proposed by Nakajima back in 2010, it has not yet been adopted by major biodiversity and taxonomy databases, where the species are still listed as separate (Rajaei et al. 2022; Operophtera variabilis … 2024; Taxonomy browser … 2024; Niwakagamania et al. 2024).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDAFFAD512CA81F3B904699.taxon	materials_examined	Material. Andreevskii Cordon: 6 – 7.11.2023 – 2 ♂, 7 – 8.11.2023 – 10 ♂, 8 – 9.11.2023 – 3 ♂, 10 – 11.11.2023 – 3 ♂, 11 – 12.11.2023 – 3 ♂, 13 – 14.11.2023 – 7 ♂, 14 – 15.011.2023 – 1 ♂, 18 – 19.11.2023 – 2 ♂.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDAFFAD512CA81F3B904699.taxon	distribution	Distribution. Russia (S Amur Obl., Jewish A. O., S Khabarovsk Kr., Primorsky Kr., S Sakhalin, S Kurils – Kunashir); Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima), South Korea, North-East and Central China.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDAFFAD512CA81F3B904699.taxon	discussion	Remarks. The species is newly recorded for the Kuril Islands. On Kunashir, the moths were observed from the first to the third week of November. In Japan, O. relegata is widely distributed from plains to subalpine areas, being common in many places. Depending on the locality, moths are found from early November to mid-January (Nakajima, Yazaki 2011). In the continental part of the RFE, the moths are observed throughout October (Beljaev 1996). The larvae feeding on various leaved woody plants.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDDFFAD5153AC06399D4367.taxon	materials_examined	Material. Andreevskii Cordon, 1 – 2.11.2023 – 1 ♀.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDDFFAD5153AC06399D4367.taxon	distribution	Distribution. Russia (Amur Obl., Khabarovsk Kr., Primorsky Kr., Sakhalin, S Kurils – Kunashir,), Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima), South Korea, China (Northeast, Taiwan Island).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDDFFAD5153AC06399D4367.taxon	discussion	Remarks. Previously, E. daemionata was found on the Kuril Islands during the usual flight period of moths from late May to early June (Rybalkin, Beljaev 2023). In Japan, the moths are numerous in March and April in various areas, from plains to mountains (Nakajima, Yazaki 2011). The present specimen collected in November is likely an accidentally hatched individual of the wintering generation. The host plants of the larvae remain unknown.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDDFFAD52D1A83E3E52467F.taxon	materials_examined	Material. Andreevskii Cordon, 15 – 16, 19 – 20, 26 – 27.10.2023 – 1 ♂, 2 ♀; ibid.: 1 – 2.11.2023 – 1 ♀, 3 – 4.11.2023 – 2 ♀, 6 – 7.11.2023 – 1 ♂, 1 ♀.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDDFFAD52D1A83E3E52467F.taxon	distribution	Distribution. Russia (S RFE: Khabarovsk Kr., Primorsky Kr., Sakhalin, S Kurils – Iturup, Kunashir, Shikotan); Japan (Hokkaido, Honshu – north of Chubu region), South Korea.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDDFFAD52D1A83E3E52467F.taxon	discussion	Remarks. Previously, E. scribai was observed on Kunashir only in July and August (Vasilenko 1992; Eupithecia scribai … 2024). In the Chubu region of Honshu, the species occurs in the mountainous areas above 1500 m a. s. l. and its moths appear in July and August (Nakajima, Yazaki 2011). The moths collected on Kunashir in late October and early November are likely from an accidentally hatched wintering generation. V. G. Mironov of the Zoological Institute (Saint Petersburg, Russia) suggests that these specimens represent an anomalous ‘ deadlock’ generation (personal communication). The host plant of the larvae remains unknown.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDDFFAD52D1ACDF3BDF4227.taxon	materials_examined	Material. Andreevskii Cordon, 22 – 23.10.2023 – 2 ♂, 26 – 27.10.2023 – VO, 27 – 28.10.2023 – VO, ibid., alder forest, 23 – 24.10.2023 – 1 ♂, 1 ♀; ibid., plateau, mixed forest, on grass, 28 – 29.10.2023 (Zinchenko) – 1 ♂; Andreevskii Cordon, 1 – 2.11.2023 – 1 ♂; ibid., ridge, birch forest, in bottle, 23 – 24.10.2023 – 1 ind.	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDDFFAD52D1ACDF3BDF4227.taxon	distribution	Distribution. Russia (S RFE: SE Khabarovsk Kr., Primorsky Kr., S Sakhalin, S Kurils — Ku - nashir); South Korea, Japan (Hokkaido, Honshu, Izu Islands, Shikoku, Kyushu, Tsushima, Yakushima).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
CD54F960FFDDFFAD52D1ACDF3BDF4227.taxon	discussion	Remarks. On Kunashir, P. excisa is occasionally observed from the second half of May (Rybalkin, Beljaev 2023) to early November. In Japan, the species is common in various regions from May to October, and develops in two or three generations per year. The larvae are known to develop on flowers of various Ericaceae, as well as on Quercus glauca (Fagaceae) and Euonymus japonicus (Celastraceae) (Nakajima, Yazaki 2011).	en	Beljaev, Evgeniy A., Vasilenko, Sergey V., Dubatolov, Vladimir V., Zinchenko, Vadim K. (2024): First report on late autumn geometer moths (Lepidoptera: Geometridae) from Kunashir Island, Russia (Southern Kurils). Amurian Zoological Journal XVI (4): 979-995, DOI: 10.33910/2686-9519-2024-16-4-979-995, URL: https://doi.org/10.33910/2686-9519-2024-16-4-979-995
