identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
CC0887D4FFF6FFEEFF4CFE95820CFDAF.text	CC0887D4FFF6FFEEFF4CFE95820CFDAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Draconarius Ovtchinnikov 1999	<div><p>Genus Draconarius Ovtchinnikov, 1999</p><p>Notes. Although Zhao &amp; Li (2017) consider this genus polyphyletic it is here used in the traditional sense as listed in the World Spider Catalog (2024), since no definite limits of the genus were shown by Zhao &amp; Li (2017). We add in the text the abbreviation “s.l.” to indicate what we mean by using this genus name.</p></div>	https://treatment.plazi.org/id/CC0887D4FFF6FFEEFF4CFE95820CFDAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hoang, Quang Duy;Nophaseud, Liphone;Jäger, Peter	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF6FFEAFF4CFD4984AEFC85.text	CC0887D4FFF6FFEAFF4CFD4984AEFC85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Draconarius laosensis Hoang & Nophaseud & Jäger 2025	<div><p>Draconarius laosensis spec. nov.</p><p>Figs 1–5, 8, 10–11</p><p>Type material. Holotype male: LAOS: Vientiane Province: west Vang Vieng, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.4189&amp;materialsCitation.latitude=18.9257" title="Search Plazi for locations around (long 102.4189/lat 18.9257)">Tham Khan</a>, 18°55'32.52"N, 102°25'8.04"E, 314 m a.s.l., inside cave, by day, by hand, 27 December 2023, Q.D. Hoang leg. (VNMN) . Paratypes: 2 females, with same data as for holotype (VNMN), 1 male, west Vang Vieng, Tham Soksay, 18°5523.30"N, 102°23'8.71"E, 273 m a.s.l., inside cave, by day, by hand, 7 Dec. 2023, C. Kennedy leg. (SMF) ; 1 female, with same data as previous specimen, except for: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.38239&amp;materialsCitation.latitude=18.925877" title="Search Plazi for locations around (long 102.38239/lat 18.925877)">Tham Puna</a>, 18°55'33.16"N, 102°22'56.60"E, 280 m a.s.l., 9 October 2024 (SMF) ; 2 females, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.431694&amp;materialsCitation.latitude=19.013334" title="Search Plazi for locations around (long 102.431694/lat 19.013334)">north Vang Vieng</a>, west Ban Phoxay, Tham Kieo, 19°0'48.00"N, 102°25'54.12"E, 370 m a.s.l., inside cave, by hand, 2 Dec. 2012, P. Jäger leg. (SMF) .</p><p>Additional material examined. 1 immature female, with same data as for holotype (VNMN); 1 juvenile female, with same data as for holotype, except for: P. Jäger &amp; S. Packheiser leg. 24 Jul. 2018 (SMF) .</p><p>Etymology. The specific epithet is derived from the type locality; adjective.</p><p>Diagnosis. Males of the new species (Figs 1B–C, 2A–C) resemble those of D. linzhiensis (Hu, 2001) (Li et al. 2019: fig. 3A–C) in having a similar conformation of RTA and LTA, short cymbial furrow, conductor with jagged margin and embolus without coil, but it differs from the latter by the following characteristics: patellar apophysis thin and pointed tip (Fig. 1B–C), relatively blunt and wide tip in D. linzhiensis (Li et al. 2019, fig. 3B–C); RTA nearly as half as tibia in the new species and swollen medially (Fig. 1C), more than half tibia length and slightly concave in D. linzhiensis (Li et al. 2019, fig. 3C); conductor blunt (Fig. 1B–C) vs. pointed tip in D. linzhiensis (Li et al. 2019, fig. 3B, C), embolus base narrow at 5 o’clock (Fig. 1B) while wide and at 6 o’clock in D. linzhiensis (Li et al. 2019, fig. 3B). Females of the new species (Figs 3A–D, 4A–D) are similar to that of D. episomos Wang, 2003 (Wang 2003, fig. 30B) in having a small atrium situated near epigastric furrow, similar shape and position of epigynal teeth, but can be distinguished from the latter by: copulatory ducts wide and long, running parallel anteriorly and connecting spermathecae posteriorly, extremely short in D. episomos (Wang 2003, fig. 30B); spermathecal heads invisible in the new species, visible in D. episomos, (Wang 2003, fig. 30B).</p><p>Description. Male (holotype). Measurements: Total length 8.41. Prosoma length 4.23, width 2.81. Opisthosoma length 4.18, width 2.24. Prosoma dorsally yellowish brown covered by sparse short grey setae, with a conspicuous longitudinal fovea (Fig. 1A). Sternum yellowish brown covered by some grey setae. Chelicerae brown, with 3 promarginal, 2 retromarginal teeth and 1 escort seta. Opisthosoma yellowish brown covered with dense long grey setae and with some lighter inverted chevrons, dorsally (Fig. 1A). Eye sizes and interdistances: AME 0.09, ALE 0.16, PME 0.17, PLE 0.18; AME–AME 0.07, AME–ALE 0.04, AME–PME 0.06, ALE–PLE 0.01, PME–PME 0.06, PME–PLE 0.09. Leg measurements: I 22.94 (5.64, 7.29, 6.58, 3.43); II 19.41 (5.05, 5.88, 5.59, 2.89); III 17.39 (4.54, 5.21, 5.50, 2.14); IV 22.49 (5.75, 6.50, 7.43, 2.81). Leg formula I–IV–II–III.</p><p>Palp (Figs 1B, C): Yellowish brown. Patellar apophysis thin, pointed and ventrad (Figs 1B, C), RTA nearly half of tibia length, LTA about ¼ of RTA length (Fig. 1C), cymbial furrow nearly half cymbium length, conductor simple and wide, lamella with jagged margin (Figs 1B, C), embolus stout and without any coils, slightly bent at tip, embolus arising in 6 o’clock position, turning retrolateral first, and then prolaterally and distally, median apophysis elongated (Figs 1B, C).</p><p>Female (paratype from Tham Khan). As in male, except for: Measurements: Total length 9.83. Prosoma length 4.28, width 2.90. Opisthosoma length 4.55, width 3.02. Eye sizes and interdistances: AME 0.08, ALE 0.17, PME 0.16, PLE 0.15; AME–AME 0.08, AME–ALE 0.06, AME–PME 0.07, ALE–PLE 0.01, PME–PME 0.07, PME–PLE 0.11. Leg measurements: I 18.11 (4.67, 6.04, 4.89, 2.51); II 15.30 (4.21, 4.95, 3.97, 2.17); III 14.30 (3.86, 4.47, 4.03, 1.94); IV 18.52 (4.83, 5.78, 5.65, 2.26). Leg formula IV–I–II–III.</p><p>Epigyne (Figs 3B–D): Epigyne slightly wider than long; epigynal teeth short, far away from each other; atrium present near epigastric furrow; copulatory ducts wide, long, running parallel anteriorly and connecting to spermathecae posteriorly, touching each other dorsally; spermathecal heads invisible; spermathecal stalks broad, expanded anteriorly; spermathecal bases small and separated.</p><p>Variation. Males: Total length: 6.8, prosoma length 3.8, width 2.5. Opisthosoma length 3.1, width 1.9. Females: Total length: 8.1–9.2, prosoma length 3.8–4.3, width 2.5. Opisthosoma length 4.3–4.9, width 3.0. The male from Tham Soksay (Figs 2A–C) exhibits a variation compared to the holotype (Figs 1B–C) as patellar apophysis directed more distad (Fig. 2C). Similarly, the female from Tham Kieo (Figs 4A–D) also shows notable variation compared to specimens from Tham Khan (Figs 3B–D) as epigynal teeth situated more posteriorly (Fig. 4A). Furthermore, although the specimens in this study were collected from different caves, we consider all differences intraspecific variation.</p><p>Distribution. Laos (Vientiane Province) (Figs 8, 10, 11).</p><p>Natural history. The species has only been found in the caves from Vang Vieng karst, Vientiane Province, Laos, spiders residing on a funnel-webs (Figs 5A–D).</p></div>	https://treatment.plazi.org/id/CC0887D4FFF6FFEAFF4CFD4984AEFC85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hoang, Quang Duy;Nophaseud, Liphone;Jäger, Peter	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF3FFE4FF4CFF6D84F0FDD4.text	CC0887D4FFF3FFE4FF4CFF6D84F0FDD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Draconarius externus Hoang & Nophaseud & Jäger 2025	<div><p>Draconarius externus spec. nov.</p><p>Figs 6–8, 10, 11</p><p>Type material. Holotype male: LAOS: Vientiane Province: west Vang Vieng, Houay Leng, above <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.415&amp;materialsCitation.latitude=18.925577" title="Search Plazi for locations around (long 102.415/lat 18.925577)">Tham Khan</a>, 18°55'32.08"N, 102°24'53.99"E, 303 m a.s.l., 27 December 2023, along cliff, in vegetation, pitfall trap, members of Arachnology Research Workshop leg. (SMF) . Paratype: 1 male, with same data as for holotype, except for: 19 October 2024, by hand, by night, P. Jäger leg. (SMF) .</p><p>Etymology. The specific epithet is a Latin adjective, meaning “outside/outward” and refers to the life style of the new species, living outside of caves in contrast to the nearby recorded D. laosensis spec. nov.; adjective.</p><p>Diagnosis. Males of the new species (Figs 6A–C, 7A–C) are similar to those of D. yigongensis Zhao &amp; Li, 2019, D. postremus and D. songi in having a short patellar apophysis, apical embolus with loops, narrow and longitudinally oriented tegulum sclerite and a retrolaterad acuminate conductor tip, but are distinguished by LTA extending distally beyond tibia and reaching cymbium in retrolateral view (shorter and not reaching cymbium in all other species), and embolus arising from embolus base in a 5.30 o’clock position (in 6.30- in D. yigongensis, in 5- in D. postremus and in 3-o’clock position in D. songi).</p><p>Description. Male (holotype). Measurements: Total length 5.8. Prosoma length 3.0, width 2.1. Opisthosoma length 2.8, width 1.7. Prosoma dorsally brown with dark marbled pattern covered sparsely by setae, with a conspicuous longitudinal fovea and striae (Fig. 7A). Sternum pale yellowish brown covered by some setae (Fig. 7B). Chelicerae brown, with 3 promarginal, 5(right)/6(left) retromarginal teeth and 1 escort seta. Opisthosoma dark brown, dorsally covered with pale dots in anterior half, in posterior half with some inverted light chevrons (Fig. 7A). Eye sizes and interdistances: AME 0.07, ALE 0.15, PME 0.16, PLE 0.15; AME–AME 0.05, AME–ALE 0.02, AME–PME 0.07, ALE–PLE 0.08, PME–PME 0.09, PME–PLE 0.02. Leg measurements: I 11.8 (2.8, 3.55, 2.7, 1.75); II 9.25 (2.5, 2.95, 2.3, 1.5); III 8.35 (2.3, 2.6, 2.25, 1.2); IV 11.2 (3.0, 3.65, 3.15, 1.4). Leg formula I–IV–II–III.</p><p>Palp (Figs 7A–C): Light brown. Patellar apophysis short, slender, bent and pointed, claw-shaped in retrolateral view, distad. RTA longer than half of tibia length, slightly concave medially in ventral view; LTA located close to RTA. Conductor with U-shaped part in prolateral view, ventral lamella with median lobe and only slightly serrated margin, conductor dorsal apophysis curved distally. Embolus long and with 3 loops apically, embolus arising in 5.30 o’clock position. Median apophysis elongated, with same width over its entire length.</p><p>Female. Unknown.</p><p>Distribution. Laos (Vientiane Prov.; Figs 8, 10,11).</p><p>Natural history. The species has been found in the leaf litter of vegetation growing close to limestone cliffs in the Vang Vieng karst, Vientiane Province, Laos.</p></div>	https://treatment.plazi.org/id/CC0887D4FFF3FFE4FF4CFF6D84F0FDD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hoang, Quang Duy;Nophaseud, Liphone;Jäger, Peter	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFFCFFE5FF4CF8AE8328FA82.text	CC0887D4FFFCFFE5FF4CF8AE8328FA82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Draconarius joshimath Quasin, Siliwal & Uniyal 2017	<div><p>Draconarius joshimath Quasin, Siliwal &amp; Uniyal, 2017</p><p>Figs 9–11</p><p>Type material. Holotype: female (WILD-09-ARA-1305), INDIA: Uttarakhand: Joshimath, Chamoli <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=0.6006667&amp;materialsCitation.latitude=0.54933333" title="Search Plazi for locations around (long 0.6006667/lat 0.54933333)">District</a> (NDBR), 30 ⁰32'57.6"N, 79 ⁰36'02.4"E, 2293 m a.s.l., 16 July 2009, S. Quasin leg. Paratypes: 1 male (WILD-09- ARA-1306), same data as for holotype; 1 female (WILD-09-ARA-1307), 30 ⁰33'02.6"N, 79 ⁰33'14.0"E, 20 July 2009, 2243 m a.s.l., S. Quasin leg. (all not examined) .</p><p>Material examined. 1 male, INDIA: Uttarakhand: Western Himalaya, Nanda Devi Biosphere Reserve, 30°33'1.60"N, 79°33'57.47"E, 2070 m a.s.l., 16 March 2011, along road, under logs and stones, by hand, P. Jäger &amp; S. Quasin leg. (SMF). Diagnosis. See Quasin et al. (2017). Description. See Quasin et al. (2017). Distribution. India (Joshimath of Uttarakhand) (Figs 10, 11). Notes. This species was actually compared in the original diagnosis (Quasin et al. 2017) with two members of the venustus species group, but not formally included, which we do here according to the palpal morphology (Fig. 9).</p></div>	https://treatment.plazi.org/id/CC0887D4FFFCFFE5FF4CF8AE8328FA82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hoang, Quang Duy;Nophaseud, Liphone;Jäger, Peter	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
