taxonID	type	description	language	source
CC0887D4FFF6FFEEFF4CFE95820CFDAF.taxon	discussion	Notes. Although Zhao & Li (2017) consider this genus polyphyletic it is here used in the traditional sense as listed in the World Spider Catalog (2024), since no definite limits of the genus were shown by Zhao & Li (2017). We add in the text the abbreviation “ s. l. ” to indicate what we mean by using this genus name.	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF6FFEAFF4CFD4984AEFC85.taxon	description	Figs 1 – 5, 8, 10 – 11	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF6FFEAFF4CFD4984AEFC85.taxon	materials_examined	Type material. Holotype male: LAOS: Vientiane Province: west Vang Vieng, Tham Khan, 18 ° 55 ' 32.52 " N, 102 ° 25 ' 8.04 " E, 314 m a. s. l., inside cave, by day, by hand, 27 December 2023, Q. D. Hoang leg. (VNMN). Paratypes: 2 females, with same data as for holotype (VNMN), 1 male, west Vang Vieng, Tham Soksay, 18 ° 5523.30 " N, 102 ° 23 ' 8.71 " E, 273 m a. s. l., inside cave, by day, by hand, 7 Dec. 2023, C. Kennedy leg. (SMF); 1 female, with same data as previous specimen, except for: Tham Puna, 18 ° 55 ' 33.16 " N, 102 ° 22 ' 56.60 " E, 280 m a. s. l., 9 October 2024 (SMF); 2 females, north Vang Vieng, west Ban Phoxay, Tham Kieo, 19 ° 0 ' 48.00 " N, 102 ° 25 ' 54.12 " E, 370 m a. s. l., inside cave, by hand, 2 Dec. 2012, P. Jäger leg. (SMF). Additional material examined. 1 immature female, with same data as for holotype (VNMN); 1 juvenile female, with same data as for holotype, except for: P. Jäger & S. Packheiser leg. 24 Jul. 2018 (SMF).	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF6FFEAFF4CFD4984AEFC85.taxon	etymology	Etymology. The specific epithet is derived from the type locality; adjective.	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF6FFEAFF4CFD4984AEFC85.taxon	diagnosis	Diagnosis. Males of the new species (Figs 1 B – C, 2 A – C) resemble those of D. linzhiensis (Hu, 2001) (Li et al. 2019: fig. 3 A – C) in having a similar conformation of RTA and LTA, short cymbial furrow, conductor with jagged margin and embolus without coil, but it differs from the latter by the following characteristics: patellar apophysis thin and pointed tip (Fig. 1 B – C), relatively blunt and wide tip in D. linzhiensis (Li et al. 2019, fig. 3 B – C); RTA nearly as half as tibia in the new species and swollen medially (Fig. 1 C), more than half tibia length and slightly concave in D. linzhiensis (Li et al. 2019, fig. 3 C); conductor blunt (Fig. 1 B – C) vs. pointed tip in D. linzhiensis (Li et al. 2019, fig. 3 B, C), embolus base narrow at 5 o’clock (Fig. 1 B) while wide and at 6 o’clock in D. linzhiensis (Li et al. 2019, fig. 3 B). Females of the new species (Figs 3 A – D, 4 A – D) are similar to that of D. episomos Wang, 2003 (Wang 2003, fig. 30 B) in having a small atrium situated near epigastric furrow, similar shape and position of epigynal teeth, but can be distinguished from the latter by: copulatory ducts wide and long, running parallel anteriorly and connecting spermathecae posteriorly, extremely short in D. episomos (Wang 2003, fig. 30 B); spermathecal heads invisible in the new species, visible in D. episomos, (Wang 2003, fig. 30 B).	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF6FFEAFF4CFD4984AEFC85.taxon	description	Description. Male (holotype). Measurements: Total length 8.41. Prosoma length 4.23, width 2.81. Opisthosoma length 4.18, width 2.24. Prosoma dorsally yellowish brown covered by sparse short grey setae, with a conspicuous longitudinal fovea (Fig. 1 A). Sternum yellowish brown covered by some grey setae. Chelicerae brown, with 3 promarginal, 2 retromarginal teeth and 1 escort seta. Opisthosoma yellowish brown covered with dense long grey setae and with some lighter inverted chevrons, dorsally (Fig. 1 A). Eye sizes and interdistances: AME 0.09, ALE 0.16, PME 0.17, PLE 0.18; AME – AME 0.07, AME – ALE 0.04, AME – PME 0.06, ALE – PLE 0.01, PME – PME 0.06, PME – PLE 0.09. Leg measurements: I 22.94 (5.64, 7.29, 6.58, 3.43); II 19.41 (5.05, 5.88, 5.59, 2.89); III 17.39 (4.54, 5.21, 5.50, 2.14); IV 22.49 (5.75, 6.50, 7.43, 2.81). Leg formula I – IV – II – III. Palp (Figs 1 B, C): Yellowish brown. Patellar apophysis thin, pointed and ventrad (Figs 1 B, C), RTA nearly half of tibia length, LTA about ¼ of RTA length (Fig. 1 C), cymbial furrow nearly half cymbium length, conductor simple and wide, lamella with jagged margin (Figs 1 B, C), embolus stout and without any coils, slightly bent at tip, embolus arising in 6 o’clock position, turning retrolateral first, and then prolaterally and distally, median apophysis elongated (Figs 1 B, C). Female (paratype from Tham Khan). As in male, except for: Measurements: Total length 9.83. Prosoma length 4.28, width 2.90. Opisthosoma length 4.55, width 3.02. Eye sizes and interdistances: AME 0.08, ALE 0.17, PME 0.16, PLE 0.15; AME – AME 0.08, AME – ALE 0.06, AME – PME 0.07, ALE – PLE 0.01, PME – PME 0.07, PME – PLE 0.11. Leg measurements: I 18.11 (4.67, 6.04, 4.89, 2.51); II 15.30 (4.21, 4.95, 3.97, 2.17); III 14.30 (3.86, 4.47, 4.03, 1.94); IV 18.52 (4.83, 5.78, 5.65, 2.26). Leg formula IV – I – II – III. Epigyne (Figs 3 B – D): Epigyne slightly wider than long; epigynal teeth short, far away from each other; atrium present near epigastric furrow; copulatory ducts wide, long, running parallel anteriorly and connecting to spermathecae posteriorly, touching each other dorsally; spermathecal heads invisible; spermathecal stalks broad, expanded anteriorly; spermathecal bases small and separated. Variation. Males: Total length: 6.8, prosoma length 3.8, width 2.5. Opisthosoma length 3.1, width 1.9. Females: Total length: 8.1 – 9.2, prosoma length 3.8 – 4.3, width 2.5. Opisthosoma length 4.3 – 4.9, width 3.0. The male from Tham Soksay (Figs 2 A – C) exhibits a variation compared to the holotype (Figs 1 B – C) as patellar apophysis directed more distad (Fig. 2 C). Similarly, the female from Tham Kieo (Figs 4 A – D) also shows notable variation compared to specimens from Tham Khan (Figs 3 B – D) as epigynal teeth situated more posteriorly (Fig. 4 A). Furthermore, although the specimens in this study were collected from different caves, we consider all differences intraspecific variation.	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF6FFEAFF4CFD4984AEFC85.taxon	distribution	Distribution. Laos (Vientiane Province) (Figs 8, 10, 11). Natural history. The species has only been found in the caves from Vang Vieng karst, Vientiane Province, Laos, spiders residing on a funnel-webs (Figs 5 A – D).	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF3FFE4FF4CFF6D84F0FDD4.taxon	description	Figs 6 – 8, 10, 11	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF3FFE4FF4CFF6D84F0FDD4.taxon	materials_examined	Type material. Holotype male: LAOS: Vientiane Province: west Vang Vieng, Houay Leng, above Tham Khan, 18 ° 55 ' 32.08 " N, 102 ° 24 ' 53.99 " E, 303 m a. s. l., 27 December 2023, along cliff, in vegetation, pitfall trap, members of Arachnology Research Workshop leg. (SMF). Paratype: 1 male, with same data as for holotype, except for: 19 October 2024, by hand, by night, P. Jäger leg. (SMF).	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF3FFE4FF4CFF6D84F0FDD4.taxon	etymology	Etymology. The specific epithet is a Latin adjective, meaning “ outside / outward ” and refers to the life style of the new species, living outside of caves in contrast to the nearby recorded D. laosensis spec. nov.; adjective.	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF3FFE4FF4CFF6D84F0FDD4.taxon	diagnosis	Diagnosis. Males of the new species (Figs 6 A – C, 7 A – C) are similar to those of D. yigongensis Zhao & Li, 2019, D. postremus and D. songi in having a short patellar apophysis, apical embolus with loops, narrow and longitudinally oriented tegulum sclerite and a retrolaterad acuminate conductor tip, but are distinguished by LTA extending distally beyond tibia and reaching cymbium in retrolateral view (shorter and not reaching cymbium in all other species), and embolus arising from embolus base in a 5.30 o’clock position (in 6.30 - in D. yigongensis, in 5 - in D. postremus and in 3 - o’clock position in D. songi).	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF3FFE4FF4CFF6D84F0FDD4.taxon	description	Description. Male (holotype). Measurements: Total length 5.8. Prosoma length 3.0, width 2.1. Opisthosoma length 2.8, width 1.7. Prosoma dorsally brown with dark marbled pattern covered sparsely by setae, with a conspicuous longitudinal fovea and striae (Fig. 7 A). Sternum pale yellowish brown covered by some setae (Fig. 7 B). Chelicerae brown, with 3 promarginal, 5 (right) / 6 (left) retromarginal teeth and 1 escort seta. Opisthosoma dark brown, dorsally covered with pale dots in anterior half, in posterior half with some inverted light chevrons (Fig. 7 A). Eye sizes and interdistances: AME 0.07, ALE 0.15, PME 0.16, PLE 0.15; AME – AME 0.05, AME – ALE 0.02, AME – PME 0.07, ALE – PLE 0.08, PME – PME 0.09, PME – PLE 0.02. Leg measurements: I 11.8 (2.8, 3.55, 2.7, 1.75); II 9.25 (2.5, 2.95, 2.3, 1.5); III 8.35 (2.3, 2.6, 2.25, 1.2); IV 11.2 (3.0, 3.65, 3.15, 1.4). Leg formula I – IV – II – III. Palp (Figs 7 A – C): Light brown. Patellar apophysis short, slender, bent and pointed, claw-shaped in retrolateral view, distad. RTA longer than half of tibia length, slightly concave medially in ventral view; LTA located close to RTA. Conductor with U-shaped part in prolateral view, ventral lamella with median lobe and only slightly serrated margin, conductor dorsal apophysis curved distally. Embolus long and with 3 loops apically, embolus arising in 5.30 o’clock position. Median apophysis elongated, with same width over its entire length. Female. Unknown.	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFF3FFE4FF4CFF6D84F0FDD4.taxon	distribution	Distribution. Laos (Vientiane Prov.; Figs 8, 10,11). Natural history. The species has been found in the leaf litter of vegetation growing close to limestone cliffs in the Vang Vieng karst, Vientiane Province, Laos.	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFFCFFE5FF4CF8AE8328FA82.taxon	description	Figs 9 – 11	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
CC0887D4FFFCFFE5FF4CF8AE8328FA82.taxon	materials_examined	Type material. Holotype: female (WILD- 09 - ARA- 1305), INDIA: Uttarakhand: Joshimath, Chamoli District (NDBR), 30 ⁰ 32 ' 57.6 " N, 79 ⁰ 36 ' 02.4 " E, 2293 m a. s. l., 16 July 2009, S. Quasin leg. Paratypes: 1 male (WILD- 09 - ARA- 1306), same data as for holotype; 1 female (WILD- 09 - ARA- 1307), 30 ⁰ 33 ' 02.6 " N, 79 ⁰ 33 ' 14.0 " E, 20 July 2009, 2243 m a. s. l., S. Quasin leg. (all not examined). Material examined. 1 male, INDIA: Uttarakhand: Western Himalaya, Nanda Devi Biosphere Reserve, 30 ° 33 ' 1.60 " N, 79 ° 33 ' 57.47 " E, 2070 m a. s. l., 16 March 2011, along road, under logs and stones, by hand, P. Jäger & S. Quasin leg. (SMF). Diagnosis. See Quasin et al. (2017). Description. See Quasin et al. (2017). Distribution. India (Joshimath of Uttarakhand) (Figs 10, 11). Notes. This species was actually compared in the original diagnosis (Quasin et al. 2017) with two members of the venustus species group, but not formally included, which we do here according to the palpal morphology (Fig. 9).	en	Hoang, Quang Duy, Nophaseud, Liphone, Jäger, Peter (2025): Notes on the genus Draconarius Ovtchinnikov, 1999 (Araneae: Agelenidae) with description of two new species from Laos. Zootaxa 5590 (4): 555-567, DOI: 10.11646/zootaxa.5590.4.6, URL: https://doi.org/10.11646/zootaxa.5590.4.6
