taxonID	type	description	language	source
D766C41BFFC8FF90F4DC69C4B205F9DA.taxon	type_taxon	Type species. Brevitubus ellipticus sp. nov.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC8FF90F4DC69C4B205F9DA.taxon	etymology	Etymology. The name is a combination of the Latin prefix “ brevi ” (meaning “ short ”) and the Latin noun “ tubus ” (meaning “ tube ”), referring to the short connecting tube. The gender is masculine.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC8FF90F4DC69C4B205F9DA.taxon	diagnosis	Diagnosis. This new genus can be separated from all known phrurolithid genera by the combination of the following characteristics: 1) carapace with a prominent chrysanthemum-petal-shaped pattern, centered around the longitudinal fovea (Figs 2 A, 3 A, 4 A, 5 A, 6 A, 7 A); 2) abdomen with a short, white transverse stripe dorsally, located nearly at its posterior end (Figs 2 A, 3 A, 4 A, 5 A, 6 A, 7 A), and ventrally with a longitudinal line on each side and a black patch anterior to the spinnerets (Figs 2 B, 3 B, 4 B, 5 B, 6 B, 7 B); 3) embolus short and small, sperm duct visible through the cuticle and forming a three-quarters circle in ventral view (Figs 2 E, 4 E, 7 E), and RTA modified, forming a half-circle-shaped bend in dorsal view (Figs 2 C, 4 C, 7 C); 4) spermathecae bean-shaped, more or less transversely elongated (Figs 3 D, 5 D, 6 D).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC8FF90F4DC69C4B205F9DA.taxon	description	Description. Total length 2.5 – 5.0 mm. Carapace oval, with a chrysanthemum-petal-shaped pattern, centered around the longitudinal fovea (Figs 2 A, 3 A, 4 A, 5 A, 6 A, 7 A). Cervical groove inconspicuous. Eight eyes in two rows, AER and PER slightly recurved in dorsal view. Chelicerae yellow, with 3 promarginal and 4 – 6 retromarginal teeth, and with 2 spines (long and short) on base of each chelicera. Endite yellow, longer than wide. Labium tongue-shaped, longer than wide. Sternum yellowish brown, unpatterned. Legs yellow, without annuli. Abdomen oval, without dorsal scutum in female (Figs 3 A, 5 A, 6 A), with dorsal scutum in male (Figs 2 A, 4 A, 7 A); dorsum with short, white transverse dorsal line close to posterior end; venter with pair of slender black longitudinal streaks extending continuously from genital groove to spinnerets, and with black patch anterior to spinnerets (Figs 2 B, 3 B, 4 B, 5 B, 6 B, 7 B). Palp. Femur with moderately developed ventral apophysis roughly located at the middle (Figs 2 D, 4 D, 7 D); tibia with conspicuous retrolateral apophysis (RTA) strongly bent, forming a half-circle hook in dorsal view, with or without basal finger-shaped branches (with finger-shaped branch in Figs 2 C, 2 D, 4 C, 4 D and fig. 8 D – F in Liu et al. 2023; without finger-shaped branches in Fig. 7 C, 7 D and fig. 17 A – G in Mu & Zhang 2023). Tegular apophysis located anteriorly, with small triangular apex. Embolus short and small. Conductor membranous, longitudinal palm-shaped (Figs 2 E, 4 E, 7 E). Epigyne. Small atrium anteriorly (Figs 5 C, 6 C), or absent (Fig. 3 C); copulatory openings inconspicuous; median septum of various shapes (elongated in Fig. 5 C, goblet-shaped in Fig. 6 C, rectangular) or absent (Fig. 3 C); connecting tubes very short, contiguous; spermathecae bean-shaped, more-or-less transversely elongated (Figs 3 D, 5 D, 6 D).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC8FF90F4DC69C4B205F9DA.taxon	discussion	Remarks. Although all members of the new genus have two lateral lines in the ventrum of the abdomen as well as a black patch anterior to the spinnerets, the patterns in the region between two lateral lines differs among different species: no additional pattern present (Figs 3 B, 7 B); another black patch present, and more anterior from the spinnerets (Figs 4 B, 6 B); another black patch present, forming a large patch by connecting the black patch closer to the spinnerets (Figs 2 B, 5 B). Composition. Brevitubus ellipticus sp. nov., B. lunatus sp. nov., B. subellipticus sp. nov., B. subpinglong sp. nov., B. ailan (Liu, Xu, Xiao, Yin & Peng, 2019) comb. nov., B. haitun (Liu, Xu & Yin, 2023) comb. nov., B. jiulong (Liu & Li, 2022) comb. nov., and B. pinglong (Liang, Li, Yin, Li & Xu, 2021) comb. nov.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC8FF90F4DC69C4B205F9DA.taxon	distribution	Distribution. China (Hunan, Jiangxi and Guangxi) (Fig. 8)	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC9FF9DF4DC6800B578FC52.taxon	description	Figs 2 – 3, 8	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC9FF9DF4DC6800B578FC52.taxon	materials_examined	Type material. China • ♂ holotype; Guangxi, Laibin City, Jinxiu Yao Autonomous County, Shengtangshan Scenic Area; 23 ° 58 ' 03 " N, 110 ° 06 ' 31 " E; 1523 m a. s. l.; 11 Oct. 2021; A. L. He, J. X. Liu, Z. G. Huang, R. R. Liao, Q. Li, X. M. Yang, Y. L. Wen, Z. Ye, and Y. Liu leg.; HNU 599. Paratypes. China • 1 ♀; same data as for holotype; HNU 598. • 2 ♂♂; same data as for holotype; Pingzhu Nature Reserve Station; 23 ° 57 ' 07 " N, 110 ° 16 ' 21 " E; 1148 m a. s. l.; 13 Oct. 2021; collected by the same collectors as for the holotype; HNU 600 to 601.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC9FF9DF4DC6800B578FC52.taxon	etymology	Etymology. The specific epithet is derived from the Latin word “ ellipticus ” meaning “ elliptical ”, referring to the shape of the spermathecae; adjective.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC9FF9DF4DC6800B578FC52.taxon	diagnosis	Diagnosis. This new species is similar to B. haitun (Liu, Xu & Yin, 2023) comb. nov. in having similar spermathecae, copulatory ducts and femoral apophysis, but can be distinguished by the presence of a distinct Vshaped anterior margin of epigyne (versus epigyne without a V-shaped anterior margin in B. haitun) in female (compare Fig. 3 C – D with fig. 9 C – D in Liu et al. 2023), and by RTA forming an about three-quarters circle, with a long and sharp distal end (in dorsal view) (versus RTA forming a half-circle, with a short and flat distal end, also with an additional triangular apophysis at the base in B. haitun) in male (compare Fig. 2 C – F with fig. 8 C – F in Liu et al. 2023).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC9FF9DF4DC6800B578FC52.taxon	description	Description. Male (holotype): total length 3.61, carapace 1.57 long, 1.22 wide; abdomen 1.82 long, 0.98 wide. Carapace nearly round, with black marginal bands. Cervical groove and radial furrow indistinct. Fovea dark red, longitudinal. AER and PER recurved. Eye sizes and interdistances: AME 0.08, ALE 0.09, PME 0.08, PLE 0.09, AME−AME 0.05, AME−ALE 0.02, PME−PME 0.09, PME−PLE 0.05, ALE−PLE 0.07, AME−PME 0.07, ALE−ALE 0.44, PLE−PLE 0.54. CH 0.10. Chelicera with 3 promarginal and 2 retromarginal teeth. MOA 0.20 long, anterior width 0.23, posterior width 0.29. Leg measurements: I 6.33 (1.57, 0.56, 2.02, 1.48, 0.70); II 5.22 (1.07, 0.51, 1.58, 1.25, 0.81); III 4.53 (1.05, 0.45, 1.04, 1.15, 0.84); IV 6.80 (1.67, 0.52, 1.59, 2.04, 0.98); Leg formula: 4123. Leg spination: Femur I pv 1 - 1 - 1 - 1, II pv 1 - 1 - 1; Tibia I v 2 - 2 - 2 - 2 - 2 - 2 - 2 - 2 - 2, II v 2 - 2 - 2 - 2 - 2 - 2 - 2 - 2; Metatarsus I v 2 - 2 - 2 - 2, II pv 1 - 1 - 1 - 1, rv 1 - 1 - 1. Abdomen oval, dorsally with short transverse white strip close to posterior end, ventrally with 2 longitudinally light brown stripes, 1 black patch anterior to spinnerets, and 1 additional patch centrally (Figs 2 A, 2 B). Palp. Femur with a distinct apophysis (Fig. 2 D). RTA strong, modified, forming a half-circle-shaped bend in dorsal view, and with sharp distal end and finger-shaped basal apophysis (Fig. 2 C); in prolateral view, both sharp distal end and finger-shaped basal apophysis red brown, highly sclerotized (Fig. 2 D). Tegular apophysis triangular. Sperm duct distinct, U-shaped (about a three-quarter circle), occupying ~ 1 / 3 length of tegulum. Embolus small and short. Conductor membranous, longitudinally elongated and supporting base of embolus (Fig. 2 E). Female (paratype HUN 600): total length 3.80; carapace 1.77 long, 1.49 wide; abdomen 1.93 long, 1.12 wide. Carapace yellowish-brown, with a chrysanthemum-petal-shaped pattern (but not very conspicuous) (Figs 3 A, 3 B). Cervical groove and radial furrow integrating into the pattern of carapace. Fovea dark red, longitudinal. AER and PER slightly recurved. Eye sizes and interdistances: AME 0.08, ALE 0.09, PME 0.09, PLE 0.10, AME−AME 0.02, AME−ALE 0.01, PME−PME 0.06, PME−PLE 0.03, ALE−PLE 0.07, AME−PME 0.07, ALE−ALE 0.45, PLE−PLE 0.56. CH 0.08. Chelicera with 3 promarginal and 2 retromarginal teeth. MOA 0.26 long, anterior width 0.23, posterior width 0.31. Leg measurements: I 6.59 (1.60, 0.58, 2.02, 1.54, 0.85); II 5.58 (1.33, 0.55, 1.58, 1.30, 0.82); III 4.68 (1.13, 0.54, 1.02, 1.21, 0.78); IV 6.99 (1.72, 0.54, 1.69, 2.08, 0.96). Leg formula: 4123. Leg spination: Femur I pv 1 - 1 - 1 - 1, II pv 1 - 1 - 1; Tibia I v 2 - 2 - 2 - 2 - 2 - 2 - 2, II pv 1 - 1 - 1 - 1 - 1 - 1 - 1, rv 1 - 1 - 1 - 1 - 1 - 1; Metatarsus I v 2 - 2 - 2 - 2, II pv 1 - 1 - 1 - 1, rv 1 - 1 - 1. Characteristics of abdomen resemble those of male, except for the absence of a ventral central patch in female. Epigyne. Copulatory openings indistinct (Fig. 3 C); copulatory ducts expanded sac-shaped (Fig. 3 D); connecting tubes short, upright, parallel and close to each other (Fig. 3 C). Spermathecae bean-shaped, obliquely and slightly elongating (Fig. 3 D). Fertilization ducts almost upright, originating from anterior margin of spermathecae, roughly equal in length to connecting tubes (Fig. 3 D).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC9FF9DF4DC6800B578FC52.taxon	distribution	Distribution. Known only from the type locality in Guangxi, China (Fig. 8).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC4FF9CF4DC6D88B2D4F949.taxon	description	Figs 4, 8	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC4FF9CF4DC6D88B2D4F949.taxon	materials_examined	Type material. China • ♂ holotype; Guangxi, Laibin City, Jinxiu Yao Autonomous County, Shengtangshan Scenic Area; 23 ° 58 ' 03 " N, 110 ° 06 ' 31 " E; 1523 m a. s. l.; 11 Oct. 2021; A. L. He, J. X. Liu, Z. G. Huang, R. R. Liao, Q. Li, X. M. Yang, Y. L. Wen, Z. Ye, and Y. Liu leg.; HNU 605.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC4FF9CF4DC6D88B2D4F949.taxon	etymology	Etymology. The species epithet is the Latin word, “ lunatus ”, meaning “ crescent-shaped ”, in reference to the shape of the retrolateral tibial apophysis; adjective.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC4FF9CF4DC6D88B2D4F949.taxon	diagnosis	Diagnosis. Male of the new species is similar to that of B. subpinglong sp. nov. in having a U-shaped sperm duct, palm-shaped membranous conductor, and short and small embolus (compare Fig. 4 C−F with Fig. 7 C−F), but it can be distinguished by the shape of RTA with an additional branch besides a relatively short distal end in the new species, while RTA with a long distal end and without any additional branch in B. subpinglong sp. nov. (compare Fig. 4 C−F with Fig. 7 C−F).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC4FF9CF4DC6D88B2D4F949.taxon	description	Description. Male (holotype): total length 3.49, carapace 1.57 long, 1.36 wide; abdomen 1.83 long, 1.08 wide. Carapace brown, with a conspicuous chrysanthemum-petal-shaped pattern (Fig. 4 A). Cervical groove and radial furrow integrating into the pattern of carapace. Fovea dark red, longitudinal. AER and PER recurved. Eye sizes and interdistances: AME 0.08, ALE 0.09, PME 0.10, PLE 0.10, AME−AME 0.03, AME−ALE 0.01, PME−PME 0.08, PME−PLE 0.04, ALE−PLE 0.09, AME−PME 0.07, ALE−ALE 0.40, PLE−PLE 0.57. CH 0.06. Chelicera with 3 promarginal and 2 retromarginal teeth. MOA 0.22 long, anterior width 0.22, posterior width 0.28. Leg measurements: I 6.25 (1.48, 0.52, 1.90, 1.48, 0.87); II 5.00 (1.10, 0.47, 1.45, 1.15, 0.83); III 4.39 (1.07, 0.35, 1.04, 1.19, 0.74); IV 6.63 (1.70, 0.47, 1.68, 1.82, 0.96); Leg formula: 4123. Leg spination: Femur I pv 1 - 1 - 1 - 1, II pv 1 - 1 - 1; Tibia I v 2 - 2 - 2 - 2 - 2 - 2 - 2, II v 2 - 2 - 2 - 2 - 2 - 2; Metatarsus I v 2 - 2 - 2 - 2, II pv 1 - 1 - 1 - 1, rv 1 - 1 - 1. Abdomen oval. Dorsum almost black except for a short white stripe located at the posterior end (Fig. 4 A). Venter light-yellow, with 2 black longitudinal stripes and 1 black patch anterior to spinnerets, and with 1 additional light brown patch centrally (Fig. 4 B). Palp. Femur with a distinct protrusion (Fig. 4 D). RTA very strong in retrolateral view (Fig. 4 F), with two apices in prolateral view (Fig. 4 D), and with a sharp distal end and a basal apophysis with two tiny apices in dorsal view (Fig. 4 C). Tegular apophysis triangular. Sperm duct distinct, U-shaped (forming about a three-quarter circle), occupying ~ 1 / 3 length of tegulum. Embolus small and short. Conductor membranous, longitudinally elongated and supporting the base of embolus (Fig. 4 E).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC4FF9CF4DC6D88B2D4F949.taxon	distribution	Distribution. Known only from the type locality in Guangxi, China (Fig. 8). Note. Although DNA sequences could not be successfully obtained for this new species, its morphological characteristics strongly support its placement within the new genus.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC5FF9EF4DC68B1B578FC76.taxon	description	Figs 5, 8	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC5FF9EF4DC68B1B578FC76.taxon	materials_examined	Type material. China • ♀ holotype; Guangxi, Fangchenggang City, Shangsi County, Imperial Robe Mountain, Songbai Station, Shiwandashan National Nature Reserve; 21 ° 57 ' 15 " N, 108 ° 02 ' 25 " E; 234 m a. s. l.; 1 May 2021; A. L. He, J. X. Liu, Y. Liang, R. R. Liao, Q. Li, H. Q. Zhang, S. X. Zhu, Z. Q. Zhao, and Y. Liu leg.; HUN 602. Paratypes. China • 2 ♀♀; same data as for holotype; HUN 603 to 604.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC5FF9EF4DC68B1B578FC76.taxon	etymology	Etymology. This specific epithet is the combination of the Latin prefix “ sub ” and “ ellipticus ”, meaning that this new species is similar to Brevitubus ellipticus sp. nov.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC5FF9EF4DC68B1B578FC76.taxon	diagnosis	Diagnosis. Female of the new species is similar to that of B. ellipticus sp. nov. in having similar bursae and spermathecae, but can be distinguished by the nonparallel CT and hole-like atria in this new species (versus parallel CT and slit-like atria in B. ellipticus sp. nov.) (compare Fig. 5 C – D with Fig. 3 C – D).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC5FF9EF4DC68B1B578FC76.taxon	description	Description. Female (holotype): total length 3.79; carapace 1.62 long, 1.44 wide; abdomen 1.94 long, 1.17 wide. Carapace yellowish-brown, with a conspicuous chrysanthemum-petal-shaped pattern (Fig. 5 A). Cervical groove radial furrow integrated into carapace pattern still recognizable. Fovea dark red, longitudinal. AER and PER recurved. Eye sizes and interdistances: AME 0.09, ALE 0.10, PME 0.09, PLE 0.10, AME−AME 0.03, AME−ALE 0.01, PME−PME 0.11, PME−PLE 0.04, ALE−PLE 0.07, AME−PME 0.06, ALE−ALE 0.46, PLE−PLE 0.56. CH 0.06. Chelicera with 3 promarginal and 4 retromarginal teeth. MOA 0.24 long, anterior width 0.25, posterior width 0.32. Leg measurements: I 6.79 (1.85, 0.51, 2.08, 1.47, 0.88); II 5.49 (1.36, 0.42, 1.69, 1.15, 0.87); III 4.89 (1.23, 0.39, 1.20, 1.28, 0.79); IV 7.05 (1.91, 0.48, 1.82, 1.88, 0.96); Leg formula: 4123. Leg spination: Femur I pv 1 - 1 - 1 - 1, II pv 1 - 1 - 1; Tibia I pv 1 - 1 - 1 - 1 - 1 - 1 - 1 - 1, rv 1 - 1 - 1 - 1 - 1 - 1 - 1 - 1 - 1, II pv 1 - 1 - 1 - 1 - 1 - 1 - 1 - 1, rv 1 - 1 - 1 - 1 - 1 - 1 - 1; Metatarsus I v 2 - 2 - 2 - 2, II pv 1 - 1 - 1 - 1, rv 1 - 1 - 1. Abdomen oval. Dorsum brown, with a sightly wide transverse band almost close to the posterior end (Fig. 5 A). Venter light yellow, with three black longitudinal stripes (the middle one wider than the other two) joining the black patch which surrounds the spinnerets (Fig. 5 B). Epigyne. Two atria like two holes, with highly sclerotized margins (Fig. 5 C). Copulatory ducts highly sclerotized, sac-shaped (Fig. 5 C); Connecting tubes short, with the parts connecting the bursae separated from each other and the parts connecting the spermathecae close to each other (Fig. 5 D). Spermathecae bean-shaped. Fertilization ducts originating the anterior margins of spermathecae (Fig. 5 D).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC5FF9EF4DC68B1B578FC76.taxon	distribution	Distribution. Known only from the type locality in Guangxi, China (Fig. 8).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC7FF9BF4DC6DB1B578FDEE.taxon	description	Figs 6 – 8	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC7FF9BF4DC6DB1B578FDEE.taxon	materials_examined	Type material. China • ♀ holotype; Guangxi, Laibin City, Jinxiu Yao Autonomous County, Yinshan Park; 24 ° 24 ' 00 " N, 110 ° 14 ' 37 " E; 1148 m a. s. l.; 8 Oct. 2021; A. L. He, J. X. Liu, Z. G. Huang, R. R. Liao, Q. Li, X. M. Yang, Y. L. Wen, Z. Ye, and Y. Liu leg.; HUN 592. Paratypes. China • 5 ♂♂; same data as for holotype; HUN 593 to 597.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC7FF9BF4DC6DB1B578FDEE.taxon	etymology	Etymology. This specific epithet is the combination of the Latin prefix “ sub ” and “ pinglong ”, meaning that this new species is similar to B. pinglong comb. nov.	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC7FF9BF4DC6DB1B578FDEE.taxon	diagnosis	Diagnosis. This new species is similar to Brevitubus pinglong (Liang, Li, Yin, Li & Xu, 2021) comb. nov. in having similar spermathecae and copulatory ducts in female but can be distinguished by the globet-shaped median septum in the new species (versus rectangular-shaped in B. pinglong) (compare Fig. 6 C, 6 D with fig. 2 C, 2 D in Liang et al. 2021). This new species is similar to Brevitubus haitun (Liu, Xu & Yin, 2023) comb. nov. in having similar femoral apophysis in male, but can be distinguished by the RTA modified forming a about three-quarter circle, with a long and sharp distal end (in dorsal view) in the new species (versus RTA modified to form a half circle, with a short and flat distal end and an additional triangular apophysis at the base in B. haitun) (compare Fig. 7 C with fig. 8 D in Liu et al. 2023).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC7FF9BF4DC6DB1B578FDEE.taxon	description	Description. Female (holotype): total length 3.84; carapace 1.68 long, 1.49 wide; abdomen 2.04 long, 1.18 wide. Carapace yellowish-brown, with a very conspicuous chrysanthemum-petal-shaped pattern (Fig. 6 A, 7 A). Cervical groove and radial furrow integrating into the pattern of carapace. Fovea dark red, longitudinal. AER and PER slightly recurved. Eye sizes and interdistances: AME 0.08, ALE 0.09, PME 0.08, PLE 0.09, AME−AME 0.04, AME−ALE 0.02, PME−PME 0.10, PME−PLE 0.05, ALE−PLE 0.09, AME−PME 0.07, ALE−ALE 0.44, PLE−PLE 0.56. CH 0.06. Chelicera with 3 promarginal and 2 retromarginal teeth. MOA 0.24 long, anterior width 0.23, posterior width 0.31. Leg measurements: I 6.92 (1.56, 0.42, 2.07, 1.82, 1.05); II 5.67 (1.32, 0.48, 1.63, 1.34, 0.90); III 4.78 (1.15, 0.40, 1.30, 1.23, 0.70); IV 7.29 (1.83, 0.41, 1.87, 2.12, 1.06); Leg formula: 4123. Leg spination: Femur I pv 1 - 1 - 1 - 1, II pv 1 - 1 - 1; Tibia I v 2 - 2 - 2 - 2 - 2 - 2 - 2 - 2, II v 2 - 2 - 2 - 2 - 2 - 2 - 2; Metatarsus I v 2 - 2 - 2 - 2, II pv 1 - 1 - 1 - 1, rv 1 - 1 - 1. Abdomen oval, dorsally with 1 short transverse white strip close to the posterior end (Figs 6 A, 7 A), ventrally with 2 longitudinally light brown stripes and 1 brown patch anterior to spinnerets (Figs 6 B, 7 B). Epigyne. Two atria sac-like, with highly sclerotized margins (Fig. 6 C). Copulatory ducts highly sclerotized and short (Fig. 6 D); Connecting tubes short and close to the bean-shaped spermathecae (Fig. 6 C). Spermathecae separated from each other by approximately the width of one spermatheca. Fertilization ducts originating from the anterior margins of the spermathecae, with the distal end not sharp, giving a false impression of being cut off (Fig. 6 D). Male (paratype): total length 3.58, carapace 1.5 long, 1.31 wide; abdomen 1.84 long, 1.15 wide. Carapace with a conspicuous chrysanthemum-petal-shaped pattern (Fig. 7 A). Cervical groove and radial furrow not evident. Fovea dark red, longitudinal. AER and PER recurved. Eye sizes and interdistances: AME 0.09 ALE 0.07, PME 0.08, PLE 0.07, AME−AME 0.04, AME−ALE 0.02, PME−PME 0.10, PME−PLE 0.04, ALE−PLE 0.08, AME−PME 0.07, ALE−ALE 0.41, PLE−PLE 0.51. CH 0.06. Chelicera with 3 promarginal and 2 retromarginal teeth. MOA 0.20 long, anterior width 0.24, posterior width 0.31. Leg measurements: I 6.87 (1.72, 0.40, 2.11, 1.71, 0.93); II 5.18 (1.22, 0.36, 1.65, 1.19, 0.76); III 4.36 (1.21, 0.37, 0.98, 1.12, 0.68); IV 7.27 (1.94, 0.50, 1.78, 2.09, 0.96); Leg formula: 4123. Leg spination: Femur I pv 1 - 1 - 1 - 1, II pv 1 - 1 - 1; Tibia I v 2 - 2 - 2 - 2 - 2 - 2 - 2 - 2 - 2, II v 2 - 2 - 2 - 2 - 2 - 2 - 2 - 2; Metatarsus I v 2 - 2 - 2 - 2, II pv 1 - 1 - 1 - 1, rv 1 - 1 - 1. Other characteristics resemble those of female. Palp. Femur with a distinct apophysis (Fig. 7 D). RTA present, strongly modified into a large hook (about a three quarters circle) and with a reddish-brown sharp distal end in dorsal view (Fig. 7 C). Tegular apophysis triangular, located on the tegular anteriorly. Sperm duct distinct, U-shaped (forming almost a three-quarter circle), occupying ~ 1 / 3 length of tegulum. Embolus short, slightly twisting, with a needle-like end. Conductor membranous, longitudinally elongated and supporting the base of the embolus (Fig. 7 E).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC7FF9BF4DC6DB1B578FDEE.taxon	distribution	Distribution. Known only from the type locality in Guangxi, China (Fig. 8).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC2FF9BF4DC6CD4B36FFBE0.taxon	materials_examined	Material examined. Holotype. China • ♀; Hunan Province, Chenzhou City, Yizhang County, Mangshan National Forest Park, Jiangjunzhai Scenic Spot; 24 ° 56 ' 35 " N, 112 ° 56 ' 54 " E; 1240 m a. s. l.; 6 Dec. 2017; H. Q. Yin, A. L. He, J. X. Liu, Y. H. Xie, Y. Yang, and P. Dong leg. (examined).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC2FF9BF4DC6CD4B36FFBE0.taxon	distribution	Distribution. Hunan, China (Fig. 8).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC2FF9BF4DC6ADFB310FA3E.taxon	materials_examined	Material examined. Holotype. China • ♂; Guangxi, Nanning City, Shanglin County, Yongjun Station; 23 ° 27 ' 19 " N, 108 ° 29 ' 37 " E; 518 m a. s. l.; 8 Nov. 2018 (examined).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC2FF9BF4DC6ADFB310FA3E.taxon	distribution	Distribution. Guangxi, China (Fig. 8).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC2FF9BF4DC6840B361F854.taxon	materials_examined	Material examined. Holotype. China • ♀; Jiangxi Province, Yichun City, Wanzai County, Luocheng Town, Zuojiashan Village, Jiulong Virgin Forest Scenic Spot; 28 ° 21 ' 08 " N, 114 ° 30 ' 28 " E; 164 m a. s. l.; 6 Feb. 2021 (examined).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFC2FF9BF4DC6840B361F854.taxon	distribution	Distribution. Jiangxi, China (Fig. 8).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFDCFF85F4DC6ECDB310FE33.taxon	materials_examined	Material examined. Holotype. China • ♀; Guangxi, Fangchenggang City, Shangsi County, Shiwandashan Mountain, Pinglong Protection Station; 21 ° 50 ' 32 " N, 107 ° 53 ' 00 " E; 619 m a. s. l.; 28 Apr. 2021 (examined).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
D766C41BFFDCFF85F4DC6ECDB310FE33.taxon	distribution	Distribution. Guangxi, China (Fig. 8).	en	Zhu, Shixiong, Liao, Rongrong, Yin, Haiqiang, Xu, Xiang (2025): Brevitubus gen. nov. (Araneae, Phrurolithidae), a new guardstone spider genus from southern China, with descriptions of four new species and proposal of four new combinations. Zootaxa 5637 (3): 450-468, DOI: 10.11646/zootaxa.5637.3.2, URL: https://doi.org/10.11646/zootaxa.5637.3.2
