taxonID	type	description	language	source
D4614D30FFE6FFBEE6CF7FCB53ADFC67.taxon	type_taxon	Type species Benlomondia benlomondensis Shear & Marek, new species	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE6FFBEE6CF7FCB53ADFC67.taxon	diagnosis	Diagnosis. The minute size of its species (ca 5 mm) separates Benlomondia from other Caseyidae, which are larger than 8 mm in length, except for Martenseya minutocaeca Shear, 2021, but that species is eyeless while Benlomondia species have black, pigmented ommatidia. From members of the subfamily Ochrogrammatinae, Benlomondia is distinct in lacking broad lamellae projecting from the male mandibular stipes, and from species of Caseya Cook & Collins, 1895, in the male seventh coxae without modifications or processes. The status of the genus Speoseya Causey, 1963, is presently unclear, but that single species is twice the size of either of the two species of Benlomondia. Additionally, the new genus is distinct from the related genus Opiona Chamberlin, 1952, in several characters of the gonopods. In Benlomondia the sterna, coxae and angiocoxites of the gonopods are fused into single units on each side, and the angiocoxites, while separated left from right, are so tightly appressed in the midline that they are functionally one structure. While some species of Opiona have the distal elements of the gonopods arising from a coxosternum, in no case are these distal elements also part of the gonopod complex, and no species are known in which the angiocoxites are closely appressed. Benlomondia species gonopods also lack flagellocoxites, present in nearly all Opiona species, nor have vestigial articulated telopodites, a feature of more than half the species of Opiona. The ninth legs of Benlomondia have reduced telopodites which are laterally flattened, unlike the inflated button-shaped telopodites of Opiona species. The long coxal lobes, strongly reduced and modified telopodites of the third male legs characteristic of species of Opiona, are not seen in Benlomondia.	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE6FFBEE6CF7FCB53ADFC67.taxon	etymology	Etymology. The name of the new genus is feminine in gender and refers to the community of Ben Lomond, California, type locality of the type species.	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE6FFBEE6CF7FCB53ADFC67.taxon	description	Description. Tiny caseyid millipedes (<5 mm long) with 30 postcephalic rings. Trunk tapering evenly anteriorly and posteriorly. Rings without shoulders or paranota, smooth and shining. Unpigmented except for faint grayish mottling on anterior end of some individuals. Head (Fig. 1) with three or four black ommatidia on each side. Metazonites with few faint ventrolateral striae. Segmental setae in a straight posterior row, acute. Male mandibles not modified. First legpair of males incrassate, with one or two swordlike, slightly twisted macrosetae on femora and postfemora; tarsus with ventral comb of modified setae (Fig. 2). Second legpair of males reduced, coxae with short, curved gonapophyses densely set with long, curled setae (Figs 2, 3). Coxae of third legpair of males with short distal lobes, densely set with recumbent, curved setae and single long, acute terminal seta; telopodites slightly reduced, prefemora not enlarged and flattened (Figs 4, 10). Legpairs four to seven unmodified, hardly enlarged. Gonopod complex (Figs 5, 6, 9) consisting of sternum, coxa and angiocoxite fused on each side with angiocoxites tightly appressed to each other in midline; distally angiocoxites with complex branches and fine, unsocketed cuticular filaments posteriorly. Colpocoxites elongate, poorly sclerotized. Ninth legpair (Figs 7, 9) with telopodites and coxal processes roughly equal in size, coxal processes broad, curved, with pore at the base. Telopodites of a single podomere, laterally flattened, setose. Tenth legpair with enlarged coxae having anteriorly opening glands, coxae without posteriorly directed processes. Notes. While the pregonopodal leg modifications of the two species of Benlomondia resemble those of species of Opiona, they are much less obvious. The gonapophyses of the male second legs are shorter than the femora; the telopodites of almost normal size, with the typical five postcoxal podomeres. The male third legs of Opiona species have very elongate projecting coxal lobes, with a strongly reduced telopodite attached about in the midlength of the lobes or even more distally. The femora of the third leg telopodites are typically expanded and flattened. Benlomondia species do not have such elongate coxal lobes and the attachment of an unreduced telopodite is more basal; the telopodite femora are cylindrical, not expanded or flattened. The tenth legpair coxae of Opiona species have hooked processes, which are absent in Benlomondia species. The lesser modifications seen in Benlomondia species may be associated with the unusually small size of the animals.	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE5FFBFE6CF78BA535CFEA7.taxon	description	Figs 1 – 8, 11	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE5FFBFE6CF78BA535CFEA7.taxon	materials_examined	Types: Male holotype, male paratype and two female paratypes from Ben Lomond, Santa Cruz Co., California, 37.089 ° N, - 122.089 ° W, collected 22 January 1955 by D. Burdick & M. Wasbauer. Parts of the holotype and paratype males are mounted on SEM stubs WS 38 - 4. Types deposited in Essig Museum of Entomology, University of California at Berkeley, Berkeley, California (EMEC). Additional specimens: CALIFORNIA: Santa Cruz Co.: UCSC campus, ravine beside Earth Sciences Building, 36.998 ° N, - 122.059 ° W, 15 February 2004, W. Leonard, m ff (VMNH); UCSC campus, Cave Gulch, near large cave opening, lat / long as above, 14 February 2004, W. Leonard, ff (VMNH).	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE5FFBFE6CF78BA535CFEA7.taxon	etymology	Etymology. The species name refers to the type locality.	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE5FFBFE6CF78BA535CFEA7.taxon	diagnosis	Diagnosis. Distinct from Benlomondia mateo, sp. nov., in details of the gonopods and ninth legpair. In B. benlomondensis, the stem formed from the appressed angiocoxites is narrower than in B. mateo and the terminal branches are different (cf. Figs 6 and 9). The coxal processes of the ninth legpair (cp, Fig. 7) are apically bifid in B. benlomondensis, while those of B. mateo are entire, broader, and sickle-shaped (cp, Fig. 9).	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE5FFBFE6CF78BA535CFEA7.taxon	description	Description. Holotype male. About 4.5 mm long, 0.52 mm in diameter. Eyepatch linear, consisting of four small, black ommatidia (Fig. 1). With the characters of the genus. Male first legpair incrassate, femora each with two twisted blade-like macrosetae, single similar macrosetae on postfemora, tarsi with ventral combs of 5 – 8 short setae (t 1, Fig. 2). Male second legs with gonapophyses (ga, Figs 2, 3) shorter than femora, anteriorly hooked, densely set with long, thin curling setae. Opening of vas deferens or possible accessory pore not observed; telopodites not greatly reduced, without specialized setae (t 2, Fig. 2). Male third leg coxae (cx 3, Fig. 4) with low, short distal lobe with many recumbent, curled setae (cl, Fig. 4); telopodites not reduced, femora slightly depressed ventrally, otherwise unmodified (t 3, Fig. 4). Legpairs 4 – 7 not incrassate. Gonopods in anterior view (Fig. 5) with coxae, sternum and angiocoxites of each side fused (cxs, Fig. 5), angiocoxites narrow. In posterior view (Figs 6, 11), angiocoxites distally broadly expanded, with three divisions: the posteriormost membranous and set with many fine cuticular filaments; the next anterior extending at right angles, with an acute basal branch and hooked apex; the most anterior rounded lobes with cuticular filaments (ac, Figs 5, 6, 11). Colpocoxites long, narrow, poorly sclerotized (cc, Figs 5, 6, 11). Ninth leg coxae (cx 9, Fig. 7) with large distal processes (cp, Fig. 7), slightly twisted and bifid apically, gland openings at bases of processes (g, Fig. 7); telopodites single-articled, not exceeding coxal processes, laterally flattened, setose (t 9, Fig. 7). Tenth coxae enlarged (cx 10, Fig 8), bearing gland openings (g, Fig. 8). Telopodites (t 10, Fig. 8) somewhat reduced. Paratype female. As male in nonsexual characters, vulvae without distinguishing features.	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE5FFBFE6CF78BA535CFEA7.taxon	discussion	Notes. The label for the holotype and accompanying paratypes provide no more detailed locality than “ Ben Lomond, ” a small community north of Santa Cruz. Our coordinates are therefore for the center of town, though it is likely the specimens were collected elsewhere in the vicinity.	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE3FFB8E6CF7F2C54F5F841.taxon	description	Figs 9, 10, 12	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE3FFB8E6CF7F2C54F5F841.taxon	materials_examined	Type: Male holotype from La Honda, San Mateo Co., California, 37.320 ° N, - 122.274 ° W, elev. ca. 400 ft asl, collected 18 April 1981 by D. Chandler. Parts of the holotype are mounted on SEM stub WS 39 - 2. Holotype deposited in Field Museum of Natural History, Chicago, Illinois (FMNH).	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE3FFB8E6CF7F2C54F5F841.taxon	etymology	Etymology. The species name, a noun in apposition, refers to the type locality.	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE3FFB8E6CF7F2C54F5F841.taxon	diagnosis	Diagnosis. Distinct from Benlomondia benlomondensis, sp. nov., in the characters given in the description of that species. In addition, the colpocoxites of B. mateo are narrower, and because of the curvature of the coxal processes of legpair 9, the rather more pedunculate telopodites exceed the coxal processes (see Fig. 9).	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE3FFB8E6CF7F2C54F5F841.taxon	description	Description. Male holotype. About 4.0 mm long, 0.49 mm wide. Linear eyepatch with three (four?) black ommatidia. Unpigmented. With the characters of the genus; pregonopodal leg modifications and coxae of legpair 10 essentially as described and illustrated for B. benlomondensis, sp. nov., above (legpair 3 in Fig. 10). Gonopods in anterior view (Fig. 12) with broad base, margins of angiocoxite (ac, Fig. 12) stem irregularly serrate. In posterior view (Fig. 9), tips of angiocoxites flaring, with bifid lateral process, small acute process at base of lateral process. Flaring portion of angiocoxite tips with few fine, unsocketed cuticular filaments (ac, Fig. 9). Colpocoxites (cc, Fig. 9) weakly sclerotized, long, narrow, pointed. Legpair nine (Fig. 9) with broad coxae (cx 9, Fig. 9) bearing bladelike, laterally curved processes (cp, Fig. 9); gland openings (g, Fig. 9) at bases of coxal processes. Telopodites laterally flattened, with distinct peduncle (t 9, Fig. 9). Females unknown.	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
D4614D30FFE3FFB8E6CF7F2C54F5F841.taxon	discussion	Notes. As with the preceding species, the label simply gives the town of La Honda as the locality, so the coordinates we provide are for the city center, though the specimen was likely collected elsewhere.	en	Shear, William A., Marek, Paul E. (2025): Additions to the family Caseyidae Verhoeff, 1909. V. The new genus Benlomondia and two new species (Diplopoda, Chordeumatida, Striariidea, Caseyoidea). Zootaxa 5659 (3): 400-406, DOI: 10.11646/zootaxa.5659.3.5, URL: https://doi.org/10.11646/zootaxa.5659.3.5
