taxonID	type	description	language	source
E14D87AEFFB16E0B2A8DFCD56B3C2E79.taxon	description	Historically, there have been only two genera in Hemipeplinae viz. Hemipeplus Latreille, 1829 and Holopeplus Arrow, 1930. The type species for Hemipeplus is H. hemipterus by subsequent monotypy while the type species for Holopeplus is H. cubensis by original monotypy. Hemipeplus is the most speciose genus and is cosmopolitan, but Holopeplus is monotypic and only documented from the Greater Antilles. In this study, a third genus Eurypeplus KC & Pollock, gen. nov., is described from Africa (Madagascar), which is monotypic, with E. quadrimaculatus KC & Pollock, sp. nov., being its type species.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB16E0C2A8DF9B26CD82AD7.taxon	diagnosis	Adult Diagnosis: Head wide; eyes located posterior of midlength of head (Pollock 1997, 1999); pronotum wide and cordate often with a distinct, anterior setose pronotal pad; abdominal process narrow; last tergite of abdomen not angulate or pointed in males (Pollock 1997, 1999); body elongate; elytra flat and subparallel, bearing apical patches.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB66E0C2A8DFCE16AB92FE9.taxon	description	Types species: Eurypeplus quadrimaculatus KC & Pollock, gen. nov., sp. nov., by present designation. Derivation of generic name: The name Eurypeplus (masculine) is based on the superficial resemblance of the type species of this genus to members of the mycterid subfamily Eurypinae. Adult Description: Head wide; eyes located posterior of midlength of head, as in Hemipeplus; cranial neck present, as in the species of Hemipeplinae; pronotum elongate and not wide, cordate, campanulate, transverse, or subquadriform; in the type specimen, which is a female, a visible fovea present medially on the fourth abdominal ventrite; body elongate, but more dorsoventrally convex than in Hemipeplus or Holopeplus, somewhat resembling Eurypinae; elytra widened at midlength, no visible apical patches; prosternal process short, as in most Hemipeplus; hypomeral edges broadly separated; mesanepisterna broad and flat, as in Hemipeplinae; mesosternum narrower and more medially raised than in the other two genera; abdominal process not as narrow and pointed as in other Hemipeplinae, rather wide, but equally long; scutellar shield raised, as in Mycterinae, but other characters do not line up with Mycterinae. Taxonomically, it appears rather to be somewhere in between Eurypinae and Hemipeplinae, but we prefer not to erect a separate subfamily, Eurypeplinae. Based on its ventral characters and rather elongate body, we classify it within Hemipeplinae as the third genus in this subfamily.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB66E0C2A8DF98D6B992D29.taxon	description	There are 17 described species of Hemipeplinae from the New World, included in the genera Hemipeplus and Holopeplus. These species were described or re-described exhaustively by Pollock (1999). In this study, five new species are described expanding the number of New World species to 22. Four species were found in the Nearctic (USA) while 21 species were found in Central and South America, and the Greater Antilles. Most of the species in this region have distinct temples and pronotal pads.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB66E0C2A8DFEDF6C772906.taxon	description	Derivation of generic name: The name Holopeplus is a derivation of the name Hemipeplus. Arrow (1930) described Holopeplus based on the feature that its elytra cover the entire abdomen unlike in Hemipeplus, hence holo, meaning ‘ full’, and peplus meaning ‘ garment’.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB66E0C2A8DFEDF6C772906.taxon	diagnosis	Adult Diagnosis: Head elongate; eyes located at about mid-length of head (Arrow 1930; Pollock 1999); pronotum elongate with convex anterior margin; anterior pronotal pad distinct, heavily setose, raised shelf-like; abdominal process narrow; the last abdominal tergite angulate and pointed in males (Arrow 1930); most of the other characters consistent with the adults of Hemipeplus; body elongate; elytra flat and subparallel, bearing apical patches.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB46E0F2A8DFF486D682E67.taxon	description	FIGURE 2. Holopeplus gundlachi (Grouvelle, 1878). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB46E0F2A8DFF486D682E67.taxon	materials_examined	Type Material. Hemipeplus gundlachi, Holotype, sex unknown, in MNHN (Grouvelle Collection); not studied, see Grouvelle (1878 a, b) for details. Holopeplus cubensis Arrow: 2 specimens (1 specimen, parts only), sex unknown, on same card, labeled: ‘ [round label with red margin] Type / Holquin, Cuba, Sharp Coll. 1905 - 313 / Holopeplus cubensis, type arrow / identical with type of Hemipeplus gundlachi Grouvelle 1878 — Types compared March 1934 — H. S. ’, in BMNH. Other Material Examined. CUBA. Cienfuegos. Cayamas, 3.6, E. A. Schwarz, (DAPC, 1 ♂). Guantanamo. El Yunque, in / around campismo popular, 20 ° 20.4 ’ N, 74 ° 32.9 ’ W ca. 40 masl, 10 – 13. vi. 2012, M. Fikáček leg., at light in the campismo popular, (NMP 6 V, 1 ♂) (Fig. 2).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB46E0F2A8DFF486D682E67.taxon	diagnosis	Diagnosis. Holopeplus gundlachi is distinguished by its elongate head and pronotum, characterized by a convex anterior pronotal margin. Holopeplus gundlachi has the following diagnostic features: temples absent to moderately present in some specimens; antennomeres mostly subfiliform; eyes small (slightly shorter than scape), moderately wide and moderately convex; head elongate; pronotum elongate; anterior half slightly wider than posterior half; anterolateral angles indistinct, rounded; pronotal pad distinct and shelf-like; posterolateral angles obtuse, rounded; pronotal lobe short with median notch; scutellar shield transverse, subpentagonal, about twice as wide as long; color rufotestaceous, head and pronotum darker; distribution: Bahamas, Cuba. Male Genitalia (Figs 2 D – F). (Tegmen length = 0.48 mm (n = 2 )) Lobe of basale short and wide; shoulders of basale obtusely angled without much slant; suture between apicale and basale prominent and raised; parameres elongate with inner margins arcuate; penis broadly widened basally with notch near the base.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB46E0F2A8DFF486D682E67.taxon	description	Variation. The examined specimen from Guantánamo (Cuba) has slight temples but no differences in genitalia were found.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB46E0F2A8DFF486D682E67.taxon	discussion	Remarks. This is the only known species of Holopeplus. Geographical Distribution (Fig. 61). Greater Antilles (Bahamas and Cuba).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB56E132A8DFB0F68132F2D.taxon	description	FIGURE 3. 1. Hemipeplus hemipterus Lacordaire, 1854 (Neotype). (Form 1). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax. FIGURE 3. 2. Hemipeplus hemipterus Lacordaire, 1854. (Form 2). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB56E132A8DFB0F68132F2D.taxon	materials_examined	Type Material. Lost / destroyed, see Arrow (1930) for explanation. Neotype (here designated following Article 75.3 of ICZN (1999 )), sex unknown, labeled: ‘ FLORIDA: DADE COUNTY EVERGLADES, NAT. PK. Long Pine Key 19 - IV- 1983 M. C. Thomas / Hemipeplus marginipennis (LeConte) det. D. A. Pollock / [red label] NEOTYPE ♂ Hemipeplus hemipterus Lacordaire des. KC & Pollock 2023 ’, in FSCA (Fig. 3.1); Nemicelus marginipennis LeConte, Holotype, (Georgia, USA); not studied, see Thomas (1985) for details. Ochrosanis dohrnii Pascoe: Holotype, sex unknown, labeled: ‘ [round label with red margin] Type / [oval pink label] West Indies / Ochrosanis dohrnii Type Pasc / [label upside down] Pascoe Coll. 93 - 60 / Hemipeplus marginipennis (LeConte) det. D. A. Pollock / [red label] HOLOTYPE Ochrosanis dohrnii Pascoe exam. DA Pollock ‘ 95 / [with a barcode] NHMUK 014663488 ’, in BMNH. Left maxilla and mentum separately mounted, labeled: ‘ Ochrosanis dohrnii Pasc. / [label upside down] Pascoe Coll. 93 - 60 ’, in BMNH. Hemipeplus dejeanii Waterhouse: Holotype, sex unknown, labeled: ‘ [round label with red margin] Type / [upper side of round label] Florida [underside of round label] 44. 31. / Hemipeplus dejeanii, (Type.) C. Waterh. / Hemipeplus marginipennis (LeConte) det. D. A. Pollock [duplicate] / [red label] HOLOTYPE Hemipeplus dejeani; Waterh. exam. D. A. Pollock ‘ 95 / [with a barcode] NHMUK 014663486 ’, in BMNH. Other Material Examined. UNITED STATES OF AMERICA. Florida. Alachua Co. Gainesville Airport, 25. vi. 1987, (FSCA, 1); Gainesville (SE), Kincaid Rd., 10 – 17. iv. 1999, B. D. Sutton leg., in edge old field & dry oak hamm., 6 m malaise trap, (FSCA, 1). Brevard Co. Titusville, 19. vii. 1949, W. F. Chamberlain leg., at light, (TAMU, 1). Broward Co. Hollywood, 19. vi. 1989, D. Crane leg., on Sabal palmetto, (DAPC, 2). Collier Co. Big Cypress Nat. Pk. Bear Island, 5. iii. 1988, D. L. Matthews & T. A. Lott leg., mercury vapor light, (DAPC, 1); Immokalee, SW FL REC 27. x. 2011 – 3. xi. 2011, Susan Halbert leg., in suction trap – Short / North, (FSCA, 1); same locality, 3. xi. 2011 – 10. xi. 2011, (FSCA, 1); same locality, 15. xii. 2011 – 22. xii. 2011, (FSCA, 1); same locality, 17 – 24. vii. 2014, (FSCA, 2); same locality, 27. xii. 2012 – 3. i. 2012, (FSCA, 1); same locality, 24 – 31. x. 2016, (FSCA, 1); same locality, 16 – 23. x. 2014, (FSCA, 1); same locality, 15 – 22. x. 2015, (FSCA, 1); same locality, 15 – 22. x. 2015, (FSCA, 1); same locality, 24 – 31. x. 2013, (FSCA, 1); same locality, 3 – 10. v. 2012, (FSCA, 1); same locality, 19. vii. 2012 – 26. vii. 2012, (FSCA, 1); same locality, 2 – 9. iv. 2015, (FSCA, 1); same locality, 2 – 9. iv. 2015, (FSCA, 1); same locality, 18 – 25. ix. 2014, (FSCA, 1); same locality, 22 – 29. ix. 2011, (FSCA, 1). Duval Co. Jacksonville Fla, vii. 1958, D. W. Ribble leg., J. F. Lawrence collection, (MCZC, 1). Flagler Co. 4 mi. S Washington Oaks Gardens St. Pk., 20. vii. 1991, P. E. Skelley & L. Heyer leg., light trap nr. Beach, (DAPC, 1 ♂, 1 sex unknown). Franklin Co. St. George Island, 20, 23. v. 1986, H. & A. Howden leg., (DAPC, 1). Gulf Co. 3 mi. W. jct 30 – C & 30 – E, Cape San Blas, 4. vii. 1982, E. G. Riley leg., (EGRC, 1). Hamilton Co. Hamilton, Fredrick Blanchard collection, (MCZC, 2). Hendry Co. nr. La Bele, 20 – 27. viii. 2001, N. Terrell leg., suction trap, (FSCA, 2). Highlands Co. Archbold Biol. Sta. 8 ml. S. Lake Placid, 29. vi. 1988, P. Skelley leg., blacklight trap, (DAPC, 1); Archbold Biol. Sta. 8. iii. 1983, UVL, L. L. Lampert Jr. leg., (FSCA, 1); Sebring. Xii. C. T. Parsons leg., (MCZC, 2); same locality, 1 – 16. ix., (MCZC, 1); same locality, 1 – 11. ix., (MCZC, 1); same locality, 1 – 18. ix., (MCZC, 1). Hillsborough Co. Ruskin, 17. ix. 2008, S. Halbert & D. Ziesk leg., on Sabal palmetto, (FSCA, 4). Indian River Co. 8 / 4 / [8. iv?] W. G. Dietz leg., (MCZC, 1). Jefferson Co. Aucilla Wldlf. Mgt. Area jct. hwys. 59 & 98, 11. vi. 1988, R. Turnbow leg., (TAMU, 2). Miami-Dade Co. Everglades Nat. Pk. Long Pine Key, 19. iv. 1983, M. C. Thomas leg., (DAPC, 1); Everglades Nat. Pk. Long Pine Key 16. v. 1991, E. Morris leg., in halogen light, (FSCA, 1); Camp Mahachee Hammock,?. v. 1991, J. Gleason leg., blacklight trap, (FSCA, 1); Coral Gables, Girl Scout Camp Mahachee 9950 Old Cutler Road, 27 – 28. ii. 1998, R. M. Baranowski leg., blacklight trap, (FSCA, 1). Monroe Co. Big Pine Key Alligator Point nr. 1 – 2. vii. 1978., L. Stange leg., blacklight trap, (DAPC, 1); Everglades Nat. Pk. Pineland Trail, 14. v. 1977, R. Turnbow leg., (TAMU, 1); Everglades Nat. Pk. 13. v. 1977, R. Turnbow leg., (FSCA, 1). Pinellas Co. W. S. Blatchley leg., (BMNH, 19). Polk Co. Winter Haven, 30. vi. – 7. vii. 2006., Susan Halbert leg., short suction trap, (FSCA, 1). Putnam Co. Georgetown Fla, iv. 1948, C. T. Bruce leg., (MCZC, 2). Seminole Co. Econ Wild Area, 8. iv. 2000, Trevor Smith leg., in scrub oak / saw palmetto (burned), malaise trap, (FSCA, 1). St. John’s Co. Farer-Dykes State Park, 21. iv. 1996, R. A. Androw leg., UV, (DAPC, 1 ♂). Volusia Co. Enterprise, 15. v.? G. Lewis leg., (BMNH, 1 ♂, 1 ♀, 1 sex unknown); Enterprise, 10. x.?, on Sabal palmetto, (MCZC, 1); same locality, 3. vi.?, Liebeck collection, (MCZC, 8); same locality, H. G. Hubbard & Roland Hayward leg., (MCZC, 4); same locality, F. C. Bowditch collection, (MCZC, 1); Florida Westgate, 20. xi. 1963, J. F. Brimley leg., (DAPC, 2). Unknown locality. (BMNH, 3); G. Lewis leg., (BMNH, 1); (MCZC, 2); collection of Fredrick Allen Eddy, collected on S. palmetto leaves, (MCZC, 4); 2613,?. v. 1980 [3 collected on palm] collection of Fredrick Allen Eddy, (MCZC, 4); 1980, collection of Fredrick Allen Eddy, (MCZC, 7). Georgia. Tybee Is, 1. vi.?, Liebeck collection, (MCZC, 2); 27. vi.?, (MCZC, 1). Illinois. Cook Co. 29. iii. 1980, leg., R. El-Mal, on palm frond, (DAPC, 1). COMMONWEALTH OF THE BAHAMAS. South Bimini Isl. B. W. I.,?. vii. 1951., C. & P. Vaurie leg., (DAPC, 1 ♂); 5. vii. 1951, C. & P. Vaurie leg., (DAPC, 1) (Fig. 3.2); same locality,?. vi. 1951, (DAPC, 1 ♂, 1 sex unknown); same locality, B. W. I. v. 1951, Cazier & Gertsch leg., (DAPC, 1 ♀). CANADA. Québec Montreal?. iv. 1933, ex. Palms from Jacksonville, Fla, (DAPC, 1). CUBA. Cayamas, 5. i.?, (DAPC, 2); same locality, 3. vi.?, (DAPC, 1). UNKNOWN LOCALITY. (BMNH, 3).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB56E132A8DFB0F68132F2D.taxon	diagnosis	Diagnosis. Hemipeplus hemipterus is distinguished by three key characters: the absence of temples, truncate elytral apices, and pale coloration. Hemipeplus hemipterus has the following diagnostic features: temples absent (in some specimens appear as short and rounded extensions behind eyes); scape elongate; eyes large (as long as the scape), wide (convex) to moderately wide; pronotum elongate with a deep anterior emargination throughout the width of anterior margin; anterior half gradually wider than posterior half; anterolateral angles indistinct, rounded; pronotal pad slightly raised on lateral view; posterolateral angles obtuse, roundish; pronotal lobe short with median notch; scutellar shield subpentagonal, transverse, less than twice as wide as long; elytra very flat, truncate at the apices exposing most of the last visible abdominal tergite; color uniformly pale brown; distribution: southeast USA, Bahamas, Canada (accidental introduction on palm), Cuba, Mexico (Pollock 1999). Male Genitalia (Figs 3. 2 D – E). (Tegmen length = 1.05 – 1.18 mm (n = 6 )) Lobe of basale long and blunt; shoulders very slanted and not angulate; apicale and basale fused without a distinct suture; parameres elongate, slightly arcuate apically; penis shorter than tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB56E132A8DFB0F68132F2D.taxon	description	Variation. Specimens remarkably vary in size and width of the pronotum; some specimens have slightly produced temples which appear rounded but not angulate; elytra of some specimens appear slightly sinuate and not truncate at apices.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFB56E132A8DFB0F68132F2D.taxon	discussion	Remarks. At least Hemipeplus hemipterus, H. abditus (Mexico), and H. rodericensis (Africa) most likely deserve a separate placement, perhaps a genus, for their unique male genitalic structure (elongate tegmen with apicale and basale fused), and external morphology (very long subquadriform pronotum, and a long and narrow prosternal process that extends to the posterior of procoxae). Apart from the long subquadriform pronotum, H. egregius and H. gounellei should also belong to this group. These species are also truly flat while most other species have a slightly convex elytral disc apically. The apicale and basale of the tegmen are separated with distinct suture in almost all other Hemipeplus whose male specimens were studied, none of them have pronotum that long and subquadriform, and the prosternal process is short and limited to the anterior of procoxae. A more reliable separation of these species into a distinct genus may be achievable following molecular analysis in the future. Geographical Distribution (Fig. 61). Southeast USA, Bahamas, Canada (stray), Cuba, Mexico (Pollock 1999). Natural History. Hemipeplus hemipterus is associated with Sabal palmetto (Arecaceae); however, upon inspection of larval gut contents, fungal components (Alternaria Nees ex Wallroth sp., Curvularia Boedijn sp., and Ulocladium Preuss sp.) were found and no damage to the palm fronds was seen (Thomas 1985). Adults have been collected from unopened fronds and decaying stumps of the palmetto; they have also been collected from Acoelorraphe wrightii (Griseb. & H. Wendl.) H. Wendl. ex Becc. (Arecaceae) (Thomas 1985; Thomas & Woodruff 1986; Pollock 1999). According to specimen label data, adults have been collected from light traps, malaise traps, and suction traps.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA96E152A8DFAC96D562BF1.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ MEXICO, Tam., Hwy 101 11 mi. SW. Cd. Victoria, 4000 ’ 22 July 1982 C. W. & L. O’Brien & G. Wibmer / UNAM / [red label] HOLOTYPE Hemipeplus abditus Pollock D. A. Pollock 1997 ’, in UNAM. Two paratypes, labeled: ‘ MEX. Qto. 14 mi. E. Landa de Matamoros, V- 26 - 1974, C. W. & L. O’Brien & GB Marshall / [blue label] PARATYPE Hemipeplus abditus Pollock’, in DAPC (Fig. 4). Other Material Examined. MEXICO. Tamaulipas. Hwy 101 11 mi. SW. Cd. Victoria, 4000 ft., 22. vii. 1982, C. W. & L. O’Brien & G. Wibmer leg., (DAPC, 1 ♂).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA96E152A8DFAC96D562BF1.taxon	diagnosis	Diagnosis. Hemipeplus abditus can be distinguished from the similar-appearing H. hemipterus (Fig. 3.1) by the presence of short, angulate temples and a darker coloration. Hemipeplus abditus has the following diagnostic features: temples short and angulate but distinct; scape elongate; eyes large (as long as the scape), moderately wide, and moderately convex; pronotum elongate with anterior emargination slightly shallower than in H. hemipterus; anterior half only slightly wider than posterior half; anterolateral angles indistinct, rounded; pronotal pad more distinctly raised than in H. hemipterus; posterolateral angles obtuse, slightly angulate; pronotal lobe short with median notch; scutellar shield subpentagonal, transverse, not twice as wide as long; elytra very flat, truncate at the apices exposing most of the last visible abdominal tergite; color brown with darker head and pronotum; distribution: Mexico (Queretaro and Tamaulipas). FIGURE 4. Hemipeplus abditus Pollock, 1999 (Paratype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Tegmen dorsal view (with scale bar); E. Tegmen lateral view with one paramere removed. Male Genitalia (Figs 4 D – E). (Tegmen length = 0.96 – 0.98 mm (n = 2 )) Very similar to those of H. hemipterus, except that the shoulders of the basale are more angulate, and the parameres are straighter in lateral view, whereas those of H. hemipterus are more sinuate.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA96E152A8DFAC96D562BF1.taxon	discussion	Remarks. Since the genitalia are not very distinct from those of H. hemipterus, H. abditus might be a Mexican subspecies of H. hemipterus; signs of temples also appear in some specimens of H. hemipterus, although they are not angulate. However, Pollock (1999) suggested that these two species are sympatric. Analysis of DNA of these two species could produce a more accurate result. For now, we treat H. abditus as a valid species, given the consistent morphological differences in all examined specimens. Geographical Distribution (Fig. 61). Mexico (Queretaro and Tamaulipas).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFAF6E152A8DFDB56A8C2F27.taxon	materials_examined	Type Material. Holotype, sex unknown, in NMNH (Cuba, Cayamas); not studied, see Pollock (1999) for details. One paratype, male, labeled: ‘ Cayamas 6. [1?] Cuba / EA Schwarz Collector / NMNH / [blue label] PARATYPE Hemipeplus glabratus D. A. Pollock / [pink label] DAPC’, in DAPC (Fig. 5).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFAF6E152A8DFDB56A8C2F27.taxon	diagnosis	Diagnosis. Hemipeplus glabratus can be distinguished by its lack of temples, moniliform scape, glabrous body, and rufotestaceous color. Hemipeplus glabratus has the following diagnostic features: temples absent; scape moniliform; eyes large (longer than scape + pedicel), wide and convex; pronotum subcordiform with a deep anterior emargination; anterolateral angles produced, broadly rounded; pronotal pad indistinct; posterolateral angles obtusely rounded; pronotal lobe indistinct with faint median notch; elytra more or less flat; scutellar shield subpentagonal, transverse, less than twice as wide as long; color uniformly rufotestaceous and shiny without much vestiture or punctures; distribution: Cuba. Male Genitalia (Figs 5 D – F). (Tegmen length = 0.37 mm (n = 1 )) Lobe of basale short and blunt; shoulders very slanted and obtuse, rounded; apicale and basale with distinct suture; parameres weakly sinuate, arcuate at the apices; inner margin of apices axe-shaped; penis longer than tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFAF6E152A8DFDB56A8C2F27.taxon	discussion	Remarks. Hemipeplus glabratus is the only known species in the New World without temples and with a moniliform scape and glabrous body. Geographical Distribution (Fig. 61). Cuba (Cayería las Cayamas).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFAF6E182A8DFACE6A50281D.taxon	description	(Figs 6, 61)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFAF6E182A8DFACE6A50281D.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ BELIZE: Cayo; Las Cuevas Research Station; 550 m 16 ° 43.99 N, 88 ° 59.20 W V / 20 - 21 / 2000 M. Caterino / BMNH {E} 2000 - 124 M. Caterino / [blue label] ♀ / [red label] HOLOTYPE ♀ Hemipeplus anishae KC & Pollock’, in BMNH. One paratype, female, labeled same as holotype, in BMNH. Derivation of Specific Epithet. The specific name (noun in the genitive case) is given in allusion to Anisha Sapkota, wife of the first author.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFAF6E182A8DFACE6A50281D.taxon	diagnosis	Diagnosis. Hemipeplus anishae sp. nov. is the only dark and shiny New World species with elongate scape and without temples or pronotal pad. Hemipeplus anishae sp. nov. has the following diagnostic features: temples absent; scape elongate; eyes large (as long as scape + pedicel), wide and strongly convex; pronotum subcordiform with a deep anterior emargination; anterolateral angles produced, broadly rounded; pronotal pad indistinct; posterolateral angles obtusely rounded; pronotal lobe present with median notch; elytra more or less flat; color unevenly rufopiceous, shiny; distribution: Belize (Cayo).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFAF6E182A8DFACE6A50281D.taxon	description	Description. Measurements (in mm) — GHW: 0.85 – 0.91; HL: 0.80 – 0.86; AL: 1.21 – 1.27; GPW: 0.90 – 0.96; PL: 0.76 – 0.83; GEW: 1.26 – 1.33; EL: 4.10 – 4.50; TL: 5.66 – 6.19. FIGURE 5. Hemipeplus glabratus Pollock, 1999 (Paratype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. FIGURE 6. Hemipeplus anishae sp. nov. (Holotype) A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax. Head (Fig. 6 B) wide, (GHW / HL: 1.05 – 1.06); eyes large (as long as scape + pedicel), wide and strongly convex; temples absent; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape elongate (as long as antennomeres II + III) and widened apically; pedicel small, moniliform; antennomeres III – IV elongate and apically dilated; V – X subtriangular, each wider than the next; XI subpyriform with apex narrowed after median notch; lateral mandibular tooth absent. Pronotum (Fig. 6 B) wide (GPW / PL: 1.15 – 1.18); anterior margin with deep emargination; anterolateral angles produced and broadly rounded; lateral margins sinuate; posterior half narrowed; posterolateral angles obtusely rounded, only slightly produced; pronotal lobe significant with a deep median notch; pronotal pad (Fig. 6 C) indistinct; pronotal pits deep with prominent oval grooves. Scutellar shield broadly convex, transverse, not twice as wide as long. Elytra (Fig. 6 A) elongate (EL / GEW: 3.25 – 3.38), more or less flat; vestiture inconspicuous; apical patch not visible. Ventral surface. Prosternal process short, not extending to the posterior of procoxae; hypomeral edges widely separated; mesosternal process narrow and linear. Color shiny, unevenly rufopiceous; ventrally rufous with hypomera piceous and margins of abdominal ventrites rufopiceous; legs same color as body with tarsi lighter.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFAF6E182A8DFACE6A50281D.taxon	discussion	Remarks. The specimens of Hemipeplus anishae sp. nov. were collected from the same location as H. belizensis sp. nov. in Belize. Geographical Distribution (Fig. 61). Belize (Cayo).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA26E182A8DFD196A992C07.taxon	materials_examined	Type Material. Holotype, male, top specimen of three specimens on same pin, labeled: ‘ Republic Dominican Bani 19.1.1976 leg. J. S. Klapperich [underside of label] 4.11.1973 / SMNS / [green label] ♂ top / [red label] HOLOTYPE (top spec.) & paratypes Hemipeplus dominicensis, n. sp. D. A. Pollock’, in SMNS. Two paratypes, mounted on the same pin as the holotype, in SMNS.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA26E182A8DFD196A992C07.taxon	diagnosis	Diagnosis. Hemipeplus dominicensis can be distinguished by its elongate scape, lack of temples, and uniformly rufotestaceous color. Hemipeplus dominicensis has the following diagnostic features: temples absent; scape elongate, widened apically; eyes large (as long as scape + pedicel), moderately wide and moderately convex; pronotum subcordiform with an indistinct but broad anterior emargination; anterolateral angles slightly produced, broadly rounded; pronotal pad present but less distinct; posterolateral angles obtusely rounded; pronotal lobe brief with indistinct median notch; scutellar shield subhexagonal, transverse, about twice as long as wide; elytra flat; color uniformly rufotestaceous, dull; distribution: Dominican Republic. Male Genitalia (Figs 7 D – F). (Tegmen length = 0.42 mm (n = 1 )) Lobe of basale short, sharp, and thin; shoulders acutely pointed with an upward slant; apicale and basale without distinct suture; parameres sinuate with pointed apices; inner margin of parameres with a notch near halfway; penis subequal in length to tegmen; parameres laterally surrounded by sheath-like structures (hereafter referred to as lateral sheaths).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA26E182A8DFD196A992C07.taxon	discussion	Remarks. Hemipeplus dominicensis is closely related to H. chaos (USA, Mexico) and H. insularis (Dominican Republic) owing to the presence of lateral sheaths surrounding the parameres. Geographical Distribution (Fig. 61). Southern Dominican Republic.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA26E1B2A8DF9EE6A462DD3.taxon	materials_examined	Type Material. Holotype, sex unknown, in NMNH (Florida, Enterprise); not studied, see Thomas (1985) for details. FIGURE 7. Hemipeplus dominicensis Pollock, 1999 (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. Other Material Examined. UNITED STATES OF AMERICA. Alabama. Baldwin Co. Bon Secour NWR 30 ° 14 ’ 31 ’’ N 87 ° 49 ’ 49 ’’ W, 14. vi. 1994. T. L. Schiefer leg., blacklight trap, (MUIC, 2). Florida. Brevard Co. Cocoa Beach, 12. v. 2003., J. Brambila leg., in light, (FSCA, 1). Clay Co. Camp Blanding Training Site, ‘ Insect Survey Site 12 Cutover Sand Pine Scrub 29 ° 55.895 ’ N 81 ° 59.940 ’ W, 24. ix. 1999, Marc & Maria Minno leg., light trap, (FSCA, 2); Camp Blanding Training Site, ‘ Insect Survey Site 11 Sand Pine Scrub 29 ° 55.599 ’ N 81 ° 59.914 ’ W, 24. ix. 1999, FIGURE 8. Hemipeplus microphthalmus (Schwarz, 1878) (female). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Genitalia dorsal view without spiculum gastrale (from a male); E. Ditto, tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. Marc & Maria Minno leg., light trap, (FSCA, 3). Collier Co. Immokalee, SW FL REC, 14 / 22. vi. 2007, S. Halbert leg., suction trap (FSCA, 1); same locality, 7 – 14. vi 2.2012, Susan Halbert leg., suction trap – short / north, (FSCA, 1); same locality, 24 – 31. v. 2012, (FSCA, 1); Big Cypress Nat’l Park, Bear Island 10. vi. 1988, Matthews & Lott leg., at M. V light, (DAPC, 3); Ochopee, 26. v. 1976, C. W. O’Brien & Marshall leg., beating base cut sawgrass, (DAPC, 1). Flagler Co. Palm Coast, 22. ix. 1990, P. E. Skelley leg., porch light, (DAPC, 3). Franklin Co. East Point, 29. v. 2002, J. Brambila leg., at light, (FSCA, 1 ♀). Highlands Co. Archbold Biol. Sta., 8 mi. S. Lake Placid; 1 – 26. v. 1997, G. Steck & B. Sutton leg., flight intercept trap, (FSCA, 1); Archbold Biol. Sta. Lk. Placid, 27. ii. 1985, M. Deyrup leg., on culm of Andropogon brachystachyus, (DAPC, 1); Archbold Biol. Stat. 14 – 18. iv. 1989, Chen Wen Young leg., (DAPC, 1). Hillsborough Co. Ruskin 13. vii. 1997, B. Stinson leg., Jackson trap, (FSCA, 1). Lake Co. 4. ix. 1954., H. A. Denmark leg., at light C. A. Frost collection 1962, (MCZC, 1). Levy Co. 4.0 mi. SW Archer 16. v. 2001, P. Skelley leg., blacklight, (FSCA, 2). Miami-Dade Co. Key Biscayne, Bill Baggs Cape Florida St. Ok. 25 ° 40 ’ 08 ’’ N 80 ° 09 ’ 17 ’’ W, 18. xii. 2005, Tremove Smith leg., sifting sand dune leaf litter, (FSCA, 2); Everglades Natl. Pk., nr. Royal Palm Hammock, 10 – 11. v. 1990. E. Riley leg., in sawgrass prairie, (TAMU, 2); Coopertown, 29. ii. 1968, A. Smetana leg., (DAPC, 1). Okaloosa Co. Ft. Walton Bch., 3. iii. 1973, E. J. Kiteley, (DAPC, 1 ♀, 1 sex unknown); same locality, 29. i. 1980, (DAPC, 1 ♂); same locality, 10. ii. 1975, (DAPC, 1 ♀) (Fig. 8). Osceola Co. Kissimmee, Disney Wilderness Pres., 19. xi. 1998., A. Hamon et al. leg., on Andropogon glomeratus var. glaucopsis, (FSCA, 1). Polk Co. Winter Haven, 3207 Lake Alfred Rd., 13 – 20. ix. 2012, Susan Halbert leg., short suction trap, (FSCA, 1). Putnam Co. 3 mi. E. Melrose, K. Ordway Pres. 25. iii – 1. vi. 1999, P. E. Skelley, barrier pitfall, (FSCA, 1); Crescent City, vi. 1988, C. T. Brues leg., (MCZC, 1). St. Lucie Co. Ft. Pierce, 30. i. 2003, K. Hibbard leg., on grass, (FSCA, 1). Sarasota Co. Myakka 16 – 18. i. 1984, R. Wharton leg., (TAMU, 2). Georgia. Clinch Co. 15 km. w. Fargo Okefenokee Swamp, xi. 1979, D. Williams leg., (DAPC, 2); Fargo 28. xii. 1976, W. E. Steiner leg., (DAPC, 1). Fa [rgo?], 14. ii. 1984, R. Beshear leg., on Andropogon, (FSCA, 1). Clinton Co. Berlese of sphagnum mat 25. iii. 1973, J. S. Ashe leg., (DAPC, 2). Texas. Cameron Co. 1 mi. W Boca Chica Beach., 19. x. 1990, T. Carlow & E. Riley, sweeping Salicornia etc. at night, (TAMU, 1). Galveston Co. 13. v. 1993, P. Ireland leg., (DAPC, 2); 3.5 mi. SW Jamaica Beach, 29. iii. 1997, E. G. Riley leg., night sweeping behind sand dunes, (EGRC, 11); same locality, date and collector, (TAMU, 10). Kennedy Co. 13.5 mi. S. Santa, 11. x. 1994, E. G. Riley- 17 leg., (EGRC, 8). Kleberg Co. ca. 1 mi S. Nueces County line, sweep near beach 15. v. 1999, E. G. Riley leg., (TAMU, 6).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA26E1B2A8DF9EE6A462DD3.taxon	diagnosis	Diagnosis. Hemipeplus microphthalmus can be distinguished among US species by its small eyes, elongate temples, and pale color. Hemipeplus microphthalmus has the following diagnostic features: temples prominent and very elongate (more than 1 / 3 length of eye); scape submoniliform; eyes small (slightly longer than the scape), shallow to moderately wide and less convex to moderately convex; pronotum subcordiform, narrow and elongate with a slight and narrow anterior emargination; anterolateral angles produced, rounded; pronotal pad distinct; posterolateral angles obtusely rounded; pronotal lobe indistinct with median notch; scutellar shield subhexagonal to subpentagonal, transverse, less than twice to about twice as wide as long; elytra slightly convex; color uniformly testaceous to rufotestaceous; distribution: southeast USA extending west to Texas. Male Genitalia (Figs 8 D – F). (Tegmen length = 0.38 mm (n = 1 )) Lobe of basale short and blunt; shoulders slanted and obtusely angled; apicale and basale with distinct suture; parameres almost straight with slightly arcuate apices; apices of parameres very pointed and thin; inner margin of parameres with separate sharp thin lines that arise from basale; penis longer than tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA26E1B2A8DF9EE6A462DD3.taxon	description	Variation. Specimens exhibit variability in size and morphology, with some having smaller bodies, less convex eyes, and longer temples, while others are larger, with more convex eyes and shorter temples (Pollock, 1999). Geographical Distribution (Fig. 61). Southeast USA (Alabama, Florida, Georgia; South Carolina, Mississippi (Pollock 1999); Virginia (Hoffman 2006 )), extending west to Texas. Natural History. Unlike the other congeners found in the USA, Hemipeplus microphthalmus has been found associated with various types of grasses, such as Andropogon brachystachyus Chapm. (Poaceae), A. virginicus L., Cladium P. Browne spp. (Cyperaceae), and Typha L. spp. (Typhaceae), instead of palms (Arecaceae); they have also been collected by sweeping Salicornia L. (Amaranthaceae) at night (Blatchley 1925; Thomas 1985; Thomas & Woodruff 1986; Pollock 1999). According to specimen label data, they were also collected from Andropogon glomeratus (Walter) Britton, Sterns & Poggenb. Thomas (1985) mentioned that larvae of H. microphthalmus are found in the compressed leaf bases and stalks of various kinds of grasses. The larvae collected from Andropogon showed mostly fungal hyphae in their guts suggesting these beetles are primarily mycophagous (Thomas 1985). Adults have been collected using various traps, including flight intercept traps, blacklight traps, and malaise traps, as indicated by specimen label data and Pollock (1999).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA66E1D2A8DFF006DF02985.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ MEXICO: Guerrero microondas 10.75 km. N. Iguala, 3800 - 4300 ’ 21 Sept. 1989 R. Turnbow / [green label] FSCA / [red label] HOLOTYPE Hemipeplus tuberculatus Pollock D. A. Pollock 1997 ’, in UNAM. Two paratypes: 1 specimen, male, labeled: ‘ 8 mi. S. Izuoar de Matamaros, Puebla Mex, XII- 10 - 48 / H. B. Leech Collector / [blue label] PARATYPE Hemipeplus tuberculatus Pollock D. A. Pollock’, in DAPC (Fig. 9); 1 specimen, female, labeled: ‘ 9 mi. N. Juajuapan de Leon, 7000 ’, Oax. Mex., VI. 8, 1971, H. F. Howden / [blue label] PARATYPE Hemipeplus tuberculatus Pollock D. A. Pollock’, in DAPC. Other Material Examined. MEXICO. Oaxaca. MX 125 36 km NE Huajuapan de Leon, 18. x. 2001, F. Skillman & J. Davidson leg., (FSCA, 1 ♀).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA66E1D2A8DFF006DF02985.taxon	diagnosis	Diagnosis. Hemipeplus tuberculatus can be distinguished by its small eyes, elongate temples, wide pronotal pad, and rufous, dark color. Hemipeplus tuberculatus has the following diagnostic features: temples prominent and elongate (about 1 / 3 length of eye); scape slightly elongate; eyes small (slightly longer than scape), shallow and less convex; pronotum subcordiform, relatively narrow and elongate without distinct anterior emargination; anterolateral angles moderately produced, broadly rounded; pronotal pad distinct; posterolateral angles obtuse, rounded; pronotal lobe short with median notch; scutellar shield subhexagonal, transverse, twice as wide as long; body elongate; elytra more or less flat; color rufous, elytra slightly darker; distribution: southwest Mexico (Guerrero, Oaxaca, and Puebla). Male Genitalia (Figs 9 D – F). (Tegmen length = 0.52 mm (n = 1 )) Lobe of basale short, thin, and apically pointed; shoulders obtusely angled, not much slanted; apicale and basale with distinct suture; apicale narrowed at the base, parameres slightly arcuate near the apices; apices of parameres very pointed and thin; penis subequal to tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA66E1D2A8DFF006DF02985.taxon	description	Variation. Females have a median tubercle on the last abdominal ventrite.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA66E1D2A8DFF006DF02985.taxon	discussion	Remarks. To our knowledge, this is the only known species of Hemipeplinae in which males and females exhibit distinct external sexual dimorphism. Geographical Distribution (Fig. 61). Southwest Mexico (Guerrero, Oaxaca, and Puebla).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA76E1F2A8DFC616C3A281D.taxon	materials_examined	Type Material. Holotype, sex unknown, labeled: ‘ BRAZIL: Rondonia, 62 km. SW Ariquemes, nr. Fzda. Rancho Grande 12 – IX – 1992 U. Schmitz blacklight trap / [pink label] FSCA / [red label] HOLOTYPE Hemipeplus quadricollis Pollock D. A. Pollock 1997 ’, in MZSP. Three paratypes: 2 specimens, 1 male, 1 female, labeled: ‘ BRAZIL: Rondonia, 62 km. SW Ariquemes, nr Fzda. Rancho Grande, 12 – IX – 1992 U. Schmitz, blacklight trap [‘ BL Trap’ in the other] / quadri / [pink label] DAPC / [red label] FSCA / [blue label] PARATYPE Hemipeplus quadricollis D. A. Pollock’, in DAPC; 1 specimen, male, labeled: ‘ PERU Madre de Dios Rio Tambopata Res. 30 km (air) SW Pto. Maldonato, 290 m 12 ° 50 ’ S 069 ° 20 ’ W / Smithsonian Institution Canopy Fogging Project T. L. Erwin et al. colls. 02 May 84 02 / 02 / 082 / FOGGING 0050185 / HEMIPEPLUS SP. Det. J. F. Lawrence / [blue label] PARATYPE Hemipeplus quadricollis D. A. Pollock’, in NMNH (this is H. pseudoquadricollis). Other Material Examined. BRAZIL. Rondônia. 62 km SW Ariquemes, nr Fzda Rancho Grande, 25. xi. 1993, U. Schmitz leg., Black Light Trap, (FSCA, 1 ♀); same locality, 5 – 17. x. 1993, J. E. Eger leg., Black Light Trap, (FSCA, 1 ♂).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA76E1F2A8DFC616C3A281D.taxon	diagnosis	Diagnosis. Hemipeplus quadricollis can be distinguished from the similar-appearing H. pseudoquadricollis sp. nov. (Fig. 14) by its prominent temples and a narrower emargination on the anterior margin of the pronotum, which does not extend across the width of the neck. Hemipeplus quadricollis has the following diagnostic features: temples prominent and obtuse, rounded (about 1 / 3 length of eye); scape short, moniliform; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum subquadriform, wide and flat with indistinct emargination narrower than the neck; anterolateral angles slightly produced, rounded; pronotal pad not very distinct; posterolateral angles squarish, angulate; pronotal lobe slight without median notch; scutellar shield heptagonal, transverse, less than twice as wide as long; body short and subquadriform; elytra more or less flat; color uniformly testaceous to rufotestaceous; distribution: Brazil (Rondônia). FIGURE 10. Hemipeplus quadricollis Pollock, 1999 (Paratype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. Male Genitalia (Figs 10 D – F). (Tegmen length = 0.27 mm (n = 2 )) Lobe of basale short and slender; shoulders obtuse, rounded, not much slanted; apicale and basale with distinct suture; parameres straight, slightly sinuate near the apical half; apices of parameres very pointed and thin, inner margin with a notch near the apical half; penis subequal to tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA76E1F2A8DFC616C3A281D.taxon	description	Variation. Some specimens have slightly deeper anterior emargination on pronotum while some have fairly straight anterior margin of pronotum.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA76E1F2A8DFC616C3A281D.taxon	discussion	Remarks. A similar species found in this study, H. pseudoquadricollis, superficially resembles H. quadricollis and is sympatric; however, the male genital morphology is different. Externally as well, there are noticeable differences such as those given in the diagnosis. Geographical Distribution (Fig. 62). Brazil (Rondônia). Natural History. According to specimen label data, adults of H. quadricollis have been collected from blacklight traps; Pollock’s (1999) specimen, collected by canopy fogging in a rainforest, is H. pseudoquadricollis sp. nov.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA56E1F2A8DFD196A112FDD.taxon	materials_examined	Type Material. Holotype, sex unknown, in MNHN (Grouvelle Collection); not studied, see Pollock (1999) for details. Other Material Examined. MEXICO. Grouvelle Coll. 1903 – 123, (BMNH, 1 ♀); 1917, A. Grouvelle leg., Museum Paris, (DAPC 2 ♀). Guerrero km. 24, Carr. Taxco-Iguala 3 km. pointe de Mexcaltepec 1100 - 1250 msnm 30 – 31. viii. 2001, H. Brailovsky & E. Barrera leg., (FSCA, 1 ♂) (Fig. 11). Tabasco Grouvelle Coll., (BMNH, 1 ♀). Veracruz Fortín de las Flores 30. x. 1982, J. Huber & A. Gonzalez leg., (DAPC, 1 ♀). GUATEMALA. Vera Paz, Chiacam, Champion, B. C. A leg., (BMNH, 1 ♀).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA56E1F2A8DFD196A112FDD.taxon	diagnosis	Diagnosis. Hemipeplus mexicanus can be distinguished by its large, rounded temples, deeply emarginate pronotum anteriorly, and rufotestaceous to rufopiceous color. Hemipeplus mexicanus has the following diagnostic features: temples prominent and rounded (about 1 / 3 length of eye); scape submoniliform; eyes moderately large (as long as scape + pedicel), moderately wide and moderately convex; pronotum subcordiform, wide with deep anterior emargination; anterolateral angles produced, broadly rounded; pronotal pad very distinct, setose; posterolateral angles obtuse, rounded; pronotal lobe indistinct without median notch; scutellar shield transverse, subquadrate, about twice as wide as long; body elongate; elytra more or less flat; color rufotestaceous to rufopiceous; distribution: Guatemala, Mexico. Male Genitalia (Figs 11 D – F). (Tegmen length = 0.50 mm (n = 1 )) Lobe of basale long, moderately thick, blunt at the apex; shoulders obtuse, rounded, slanted; apicale and basale with distinct suture; parameres depressed near the bases; apices of parameres very pointed and thin; penis slightly longer than tegmen. Geographical Distribution (Fig. 61). Guatemala, Mexico.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA56E222A8DF9D86DFD2919.taxon	description	(Figs 12, 62)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA56E222A8DF9D86DFD2919.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ [underlined in green] BRAZIL: Am. [/ underlined in green] Reserva Ducke 26 km NE Manaus Barbosa, M. G. V / Plot A, Malaise 4 July 1995 / BMNH {E} 2003 - 84 / 84.1 / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus brasiliensis KC & Pollock’, in BMNH. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the type locality of the holotype, i. e., Brazil.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA56E222A8DF9D86DFD2919.taxon	diagnosis	Diagnosis. Hemipeplus brasiliensis sp. nov. can be distinguished from the similar-appearing H. angustipennis (Fig. 13) by its larger, more convex eyes, less produced anterolateral pronotal angles, and paler head and pronotum. Hemipeplus brasiliensis sp. nov. has the following diagnostic features: temples prominent, posteriorly rounded; scape short, moniliform; eyes large (longer than scape + pedicel), shallow and less convex; pronotum subcordiform, slightly widened anteriorly with a shallow anterior emargination; anterolateral angles moderately produced, broadly rounded; pronotal pad present; posterolateral angles square, rounded; pronotal lobe slight but visible with median notch; elytra more or less flat; color rufotestaceous with portions of elytra slightly piceous; distribution: Brazil (Amazonas). FIGURE 11. Hemipeplus mexicanus Grouvelle, 1896. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFA56E222A8DF9D86DFD2919.taxon	description	Description. Measurements (in mm) — GHW: 0.58; HL: 0.58; AL: 0.94; GPW: 0.66; PL: 0.52; GEW: 0.75; EL: 3.00; TL: 4.10. FIGURE 12. Hemipeplus brasiliensis sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. Head (Fig. 12 B) as long as wide, (GHW / HL: 1.00); eyes large (as long as scape + pedicel), shallow and less convex; temples wide and posteriorly rounded, about 1 / 3 length of eye, bulged outward; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape short, moniliform and produced externally on one side; pedicel small moniliform; antennomere III more elongate and apically dilated; IV short and moniliform; V slightly larger, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after median notch; lateral mandibular tooth absent. Pronotum (Fig. 12 B) wide (GPW / PL: 1.27); anterior margin with moderate emargination; anterolateral angles broadly rounded, slightly produced; lateral margins sinuate; posterior half slightly contracted than apical half; posterolateral angles square and rounded, very slightly produced; pronotal lobe slight with median notch; pronotal pad (Fig. 12 C) present but narrow; pronotal pits deep with oval grooves. Scutellar shield subhexagonal, transverse, not twice as wide as long. Elytra (Fig. 12 A) elongate (EL / GEW: 4.00) and flat; vestiture very short but dense; apical patch long and piceous. Ventral surface. Prosternal process short, not extending to the posterior of procoxae; hypomeral edges broadly separated; mesosternal process narrow and linear. Color rufotestaceous head and pronotum, with portions of elytra slightly piceous. Male Genitalia (Figs 12 D – F). (Tegmen length = 0.35 mm (n = 1 )) Lobe of basale short, slender with rounded apex; shoulders obtusely angled, fairly slanted; parameres almost straight, slightly narrowed near the bases of apicale; apices of parameres thinner than the bases with sharp thin tips; penis shorter than tegmen. Geographical Distribution (Fig. 62). Brazil (Amazonas). Natural History. The holotype of H. brasiliensis sp. nov. was collected in a malaise trap.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF986E222A8DFC1D6BBC2C6C.taxon	materials_examined	Type Material. Holotype, male, only images sent from NMNH, labeled: ‘ VENEZUELA, T. F. Amaz. Cerro de la Neblina Camp XII, 1950. m. near Pico Phelps 26 February 1985 / [second label mentioned in Pollock (1999) not seen in the photo sent] / [red label] HOLOTYPE [in Pollock’s handwriting] Hemipeplus angustipennis Pollock [printed] D. A. Pollock 1997 / [with a barcode] USNMENT, 01595475 ’, in NMNH.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF986E222A8DFC1D6BBC2C6C.taxon	diagnosis	Diagnosis. Hemipeplus angustipennis can be distinguished by its smaller, shallower eyes compared to those of the similar-appearing H. brasiliensis sp. nov. (Fig. 12), as well as by its darker head and pronotum, and more distinctly produced anterolateral margins of the pronotum. Hemipeplus angustipennis has the following diagnostic features: temples prominent and somewhat angulate, rounded posteriorly; scape short, moniliform; eyes small (shorter than scape + pedicel), very shallow and slightly convex; pronotum subcordiform, widely expanded anteriorly with a shallow wide anterior emargination; pronotal pad distinct; pronotal lobe truncate with median notch; elytra more or less flat; color head and pronotum piceous, elytra dark rufous, venter piceous; distribution: S. Venezuela (Amazonas). Geographical Distribution (Fig. 62). Venezuela (Amazonas) Natural History. Specimens of H. angustipennis examined by Pollock (1999) were collected from Bonnetia Mart. & Zucc. (Bonnetiaceae) scrub in a malaise trap.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF986E252A8DF90D6AF92F75.taxon	description	(Figs 14, 62)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF986E252A8DF90D6AF92F75.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ BRAZIL: Rondonia. 62 km SW Ariquemes, nr Fzda. Rancho Grande 20 - VIII- 1993 U. Schmitz, BL trap / MYCTERIDAE. Hemipeplus det. D. A. Pollock 2016 / [dark green label] ♂ / [light green label] FSCA / [red label] HOLOTYPE ♂ Hemipeplus pseudoquadricollis KC & Pollock’, in FSCA. Two paratypes: 1 specimen, male, labeled: ‘ PERU Madre de Dios Rio Tambopata Res. 30 km (air) SW Pto. Maldonato, 290 m 12 ° 50 ’ S 069 ° 20 ’ W / Smithsonian Institution Canopy Fogging Project T. L. Erwin et al. colls. 02 May 84 02 / 02 / 082 / FOGGING 0050185 / HEMIPEPLUS SP. Det. J. F. Lawrence / [blue label] PARATYPE Hemipeplus quadricollis FIGURE 13. Hemipeplus angustipennis Pollock, 1999 (Holotype). A. Dorsal habitus; B. Dorsal head and prothorax; C. Lateral view of head and prothorax. Image Credit. NMNH / Lourdes Chamorro. FIGURE 14. Hemipeplus pseudoquadricollis sp. nov. (Holotype, fig. D from paratype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. D. A. Pollock’, in NMNH (this was initially H. quadricollis); 1 specimen, male, labeled: ‘ BRAZIL: Mato Grosso, 12 ° 50 ’ S, 51 ° 47 ’ W 30. iii. 1968, O. W. Richards R. S. & R. G. S. Exped, B. M. 1968 - 260 / At light / MYCTERIDAE Hemipeplus sp. det. R. A. Crowson’, in BMNH. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the resemblance of this new species to H. quadricollis.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF986E252A8DF90D6AF92F75.taxon	diagnosis	Diagnosis. Hemipeplus pseudoquadricollis can be distinguished from the similar-appearing H. quadricollis (Fig. 10) by its shorter temples and pronotum with wider anterior emargination which extends across the width of the neck. Hemipeplus pseudoquadricollis has the following diagnostic features: temples short and angulate; scape short, moniliform; eyes large (longer than scape + pedicel); pronotum subquadriform, wide and flat with a distinct anterior emargination which extends across the width of the neck; anterolateral angles produced, rounded; pronotal pad distinct but narrow; posterolateral angles obtuse; pronotal lobe indistinct without median notch; body short and subquadriform; elytra more or less flat; color uniformly testaceous to rufotestaceous; distribution: Brazil (Rondônia), Peru (Madre de Dios).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF986E252A8DF90D6AF92F75.taxon	description	Description. Measurements (in mm) — GHW: 0.60 – 0.66; HL: 0.52 – 0.58; AL: 0.78 – 0.87; GPW: 0.66 – 0.68; PL: 0.48 – 0.53; GEW: 0.93 – 1.00, EL: 1.90 – 2.00; TL: 2.90 – 3.11. Head (Fig. 14 B) wide (GHW / HL: 1.14 – 1.15); eyes large (longer than scape + pedicel), wide and convex; temples short and angulate; occiput without raised areas; antennae shorter than head and pronotum length combined; antennal scape short and moniliform; pedicel small moniliform; antennomere III more elongate and apically dilated; IV short and moniliform; V slightly larger, apically dilated; VI – X subtriangular or bowl-shaped, each wider than the next; XI subpyriform with apex narrowed after notch; lateral mandibular tooth absent. Pronotum (Fig. 14 B) wide (GPW / PL: 1.28 – 1.38); anterior margin with distinct emargination as wide as neck; anterolateral angles rounded, fairly produced; lateral margins weakly sinuate; posterior half slightly contracted than anterior half; posterolateral angles obtuse; pronotal lobe truncate without median notch; pronotal pad (Fig. 14 C) slightly raised; pronotal pits moderately deep with moderate grooves. Scutellar shield elliptical (not heptagonal as in H. quadricollis), transverse, about twice as wide as long. Elytra (Fig. 14 A) short (EL / GEW: 2.00 – 2.04) and flat; vestiture almost absent, very scarce; apical patch not visible. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges well separated but less so than in H. quadricollis; mesosternal process narrow and linear. Color uniformly rufotestaceous to testaceous, shiny. Male Genitalia (Figs 14 D – F). (Tegmen length = 0.33 mm (n = 2 )) Lobe of basale short, slender, and blunt; shoulders obtuse, rounded, slanted; apicale and basale with distinct suture; parameres with slight sinuation, apices straight, very pointed, thin, narrower than the bases; penis sharply expanded near basal half, narrowed at the base and apex, shorter than tegmen (Fig. 14 D). Variation. Some specimens have slightly straighter anterolateral pronotal margins while in some specimens it is more sinuate. The color varies from testaceous to rufotestaceous.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF986E252A8DF90D6AF92F75.taxon	discussion	Remarks. Hemipeplus pseudoquadricollis resembles H. quadricollis with which it is sympatric; however, the male genital morphology is different, as was discussed above in the remarks for H. quadricollis. Geographical Distribution (Fig. 62). Brazil (Rondônia), Peru (Madre de Dios). Natural History. According to specimen label data, specimens of H. pseudoquadricollis were collected from light traps (including blacklight trap) and by rainforest canopy fogging.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF9F6E272A8DFA316D932D70.taxon	materials_examined	Type Material. Holotype, male, in FSCA (Florida, Collier Co.: Naples); not studied, see Thomas (1985) for details. Other Material Examined. UNITED STATES OF AMERICA. Florida. Alachua Co. Hwy 235, 2 mi N. Cross Creek 7. vii. 1991, C. M. Mills leg., (DAPC, 1 ♂, 1 sex unknown). Collier Co. Collier Seminole St. Park, 9. iii. 1989, M. C. Thomas leg., beating, (DAPC, 1); Immokalee, SW FL REC 10 – 17. v. 2012, Susan Halbert leg., suction trap – middle / short, (FSCA, 2); same locality, 3 – 10. v. 2012, (FSCA, 1); same locality, 15. xii. 2011 – 22. xii. 2011, (FSCA, 1); same locality, 24 – 31. v. 2012, (FSCA, 1); same locality, 2. ii. 2012 – 9. ii. 2012, (FSCA, 1); same locality, 22 – 29. ix. 2011, (FSCA, 1); same locality, 21 – 28. vii. 2014, (FSCA, 1); same locality, 31. v. 2012 – 7. vi. 2012, (FSCA, 1); same locality, 3. v. 2012 – 10. v. 2012, (FSCA, 1); same locality, 18 – 25. ix. 2014, suction trap, short / open field, (FSCA, 1). Gulf Co. 3 mi. W. jct. 30 - C & 30 - E, Cape San Blas, 4. vii. 1982, E. G. Riley leg., (EGRC, 1). Hendry Co. nr. La Belle, 20 – 27. viii. 2001, N. Terrell leg., suction trap, (FSCA, 2). Hillsborough Co. Tampa, iv. 30.?, (DAPC, 1); Tampa,?. x. 30, D. M. Castie leg., (DAPC, 1). Miami-Dade Co. Royal Palm Park, 4. i. 1930, W. S. Blatchley leg., (BMNH, 1); same locality, no date, (BMNH, 9); Biscayne, (BMNH, 3); Miami, H. C. Fall collection, H. chamaeropis, (MCZC, 1). Pinellas Co. Dunedin, W. S. Blatchley collection, (BMNH, 1); Fort de Soto Co. Park 1. vi. 1993, R. Andrew leg., beat palmetto, (DAPC, 1); Ft. de Soto Pk., 9. vi. 1991, R. Morris leg., (FSCA, 1). Dunedin, H. C. Fall collection, (MCZC, 1); Dunedin, 3. iv. 1923, H. C. Fall collection, (MCZC, 1); same locality, 3. ii. 1918, (MCZC, 1); Dunedin, 3. iv. 1921, W. S. B. leg., (MCZC, 1); Polk Co. Lake Marion Estates, 17. vi. 1995, R. F. Morris II leg., (FSCA, 1). Putnam Co. Georgetown, iii. 1948, C. T. Brues leg., (MCZC, 1). St. Lucie Co. Ft. Pierce, 2. vii. 2002, K. Hibbard leg., under fallen palm frond, (FSCA, 1). Volusia Co. Enterprise?. v. 28, (DAPC, 1); Enterprise 10.6, F. C. Bowditch collection, (MCZC, 2); same locality, 3. vi.?, Liebeck collection, (MCZC, 2); same locality 15. v.?, collection of Fredrick Allen Eddy, Holopeplus chamaeropis Schwarz MS n. sp., (MCZC, 1). Unknown locality. G. Lewis coll. B. M. 1910 — 248, (BMNH, 1); (MCZC, 8). Georgia. Chatham Co. Tybee Is [land], H. C. Fall collection, (MCZC, 4); Liebeck collection, (MCZC, 1). Unknown locality. H. C. Fall collection, (MCZC, 1); gift of F. C. Bowditch, (MCZC, 2). Texas. Cameron Co. Brownsville, ex Tamazunchale SLP. Mexico, on palm frond, (DAPC, 1) (Fig. 15); Sabal Palm Grove Sanct., 16. x. 1993, Blackmon, Quinn & Riley leg., UV light, (DAPC, 2); same locality, 8. iv. 1994., E. G. Riley leg., (EGRC, 1); same locality, 16. x. 1993, Blackmon, Quinn & Riley leg., UV light, (EGRC, 12; TAMU 7); same locality (site 11) 25.85601 ° N, 97.41726 ° W, 18. x. 2008, E. G. Riley [Riley, Räber & Heffern leg. for 1] leg., beating dead palm fronds or beating open-revegetated area, (TAMU, 6); same locality, 4. iv. 2009, E. Riley leg., beating open-revegetated area, (TAMU, 2); same locality (site 10) 25.84964 ° N, 97.41798 ° W, 5. vi. 2010, E. G. Riley leg., palm forest beating etc., (TAMU, 24); same locality, 29 – 30. v. 2009, Heffern & Räber leg., (TAMU, 5). MEXICO. Campeche. 29 km E Xpujil 29. vi. 1990, M. C. Thomas leg., (DAPC, 1); same locality, 19. vi. 1990., (DAPC, 2). San Luis Potosi. Tamazunchale SLP, (DAPC, 1); Quintana Roo. 3 km N X-Thobil 17. vi. 1990, M. C. Thomas leg., beating, (DAPC, 1); 54 km NW Felipe Carrillo Puerto 17. vi. 1990, R. Turnbow [for one] & M. C. Thomas [for one] leg., beating palms [by M. C. Thomas], (DAPC, 2). Unknown locality. (DAPC, 1). UNKNOWN LOCALITY. W. S. Blatchley Coll. B. M. 1935 – 440, (BMNH, 2); W. G. Dietz collection, (MCZC, 1); (MCZC, 2).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF9F6E272A8DFA316D932D70.taxon	diagnosis	Diagnosis. Hemipeplus chaos can be distinguished among US species by its large wide eyes, short temples, and full elytra. Hemipeplus chaos has the following diagnostic features: temples small to moderate, rounded; scape submoniliform; eyes large (longer than scape + pedicel), wide and convex; pronotum subcordiform, widened anteriorly with a distinct anterior emargination; anterolateral angles produced, broadly rounded; pronotal pad distinct to moderate; posterolateral angles obtuse, rounded; pronotal lobe indistinct without median notch; scutellar shield elliptical, transverse, not twice as wide as long; body elongate; elytra more or less flat; color uniformly testaceous to rufotestaceous; distribution: USA (Florida, Georgia, Texas), Mexico. Male Genitalia (Figs 15 D – F). (Tegmen length = 0.66 mm (n = 1 )) Lobe of basale elongate, blunt at the apex; shoulders obtusely angled, barely slanted; apicale and basale with distinct suture; parameres surrounded by lateral sheaths, sinuate, depressed near the bases, arcuate near the apices; apices of parameres sharp; penis subequal to or slightly shorter than tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF9F6E272A8DFA316D932D70.taxon	description	Variation. Some specimens have a slightly deeper anterior pronotal emargination than others; the pronotal pad is more distinct in some specimens than others.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF9F6E272A8DFA316D932D70.taxon	discussion	Remarks. As mentioned by Pollock (1999), a specimen (also examined in this study) of Hemipeplus chaos from Miami, Florida (MCZC) bears a label ‘ Holopeplus chamaeropis Sz’, presumably a manuscript name. This name has been seen neither in the literature nor on any other examined Hemipeplus specimen. Geographical Distribution (Fig. 61). Southeast USA (Florida, Georgia) extending west to Texas, and farther south to Mexico. Natural History. According to Thomas (1985), Pollock (1999), and specimen label data, H. chaos has been found associated with Sabal palmetto (Arecaceae). According to Thomas (1985), adults are found in the unopened fronds of Sabal Palm. Pollock (1999) mentioned that the Mexican specimens were obtained from burned Cabbage Palms. Adults have also been collected by sweeping Salicornia at night, beating dead palm fronds, beating in an open-revegetated area, suction traps, and light traps, according to specimen label data. FIGURE 15. Hemipeplus chaos Thomas, 1985. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax C. Lateral view of head and prothorax; D. Genitalia (from different specimen) dorsal view with spiculum gastrale; E. Male tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF926E292A8DFF00688429E1.taxon	description	FIGURE 16. Hemipeplus insularis Grouvelle, 1896. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF926E292A8DFF00688429E1.taxon	materials_examined	Type Material. Not found by Pollock (1999) or in this study. Other Material Examined. DOMINICAN REPUBLIC. Puerto Plata. Puerto Plata, 18 – 28. vi. 1993, R. E. Woodruff leg., palm thatched building, (DAPC, 1 ♂ (Fig. 16), 8 sex unknown). San Juan. 12913, 1. ii. 1957 – 5994, with pineapple slips, (DAPC, 1); 12403, lot 56 – 10028, 31. vii. 1956, Lewis and Price, leg., with pineapple slips, (FSCA, 1).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF926E292A8DFF00688429E1.taxon	diagnosis	Diagnosis. Hemipeplus insularis can be distinguished by its large eyes, very short temples, and lateral margins of the body piceous. Hemipeplus insularis has the following diagnostic features: temples scarcely present as slight roundish expansion behind eyes; scape submoniliform; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum subcordiform, elongate without distinct anterior emargination, anterior half not much widened; anterolateral angles not very produced, broadly rounded; pronotal pad very distinct; posterolateral angles obtuse, rounded; pronotal lobe slight with median notch; scutellar shield subpentagonal to subhexagonal, twice as wide as long; body slender and elongate; elytra more or less flat; color rufotestaceous – rufopiceous dorsally and piceous laterally; distribution: Dominican Republic (Peravia, Puerto Plata, San Juan). Male Genitalia (Figs 16 D – F). (Tegmen length = 0.42 mm (n = 1 )) Lobe of basale short, thin, slightly widened apically and rounded; shoulders straight and acutely angled; apicale and basale with distinct suture; parameres surrounded by lateral sheaths, fairly straight with inner margins arcuate; apices of parameres sharp and pointed; penis longer than tegmen. Geographical Distribution (Fig. 61). Dominican Republic (Peravia (Pollock 1999), Puerto Plata, San Juan). Natural History. According to specimen label data, H. insularis is associated with palms (Arecaceae) and pineapples (Bromeliaceae).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF936E2B2A8DFB846D782DDA.taxon	description	(Figs 17, 61)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF936E2B2A8DFB846D782DDA.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ BELIZE: Cayo; Las Cuevas Research Station; 550 m 16 ° 43.99 N, 88 ° 59.20 W V / 20 - 21 / 2000 M. Caterino / BMNH {E} 2000 - 124 M. Caterino / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus belizensis KC & Pollock’, in BMNH. Three paratypes: 2 specimens, 1 female, 1 sex unknown, same label as in holotype, in BMNH; 1 specimen, male [without head and prothorax], labeled: ‘ GUATE PETEN 5 km S. Santa Elena, 200 - 300 m; beating; 16 ° 52.350 ’ N 89 ° 49.899 ’ W 20 - 25. vi. 2012., E. Fuller 12 - 18 ’, in FSCA. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the type locality of the holotype, i. e., Belize.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF936E2B2A8DFB846D782DDA.taxon	diagnosis	Diagnosis. Hemipeplus belizensis sp. nov. can be distinguished by its large eyes, short temples, and dark, rufopiceous body with coarse dorsal texture. Hemipeplus belizensis sp. nov. has the following diagnostic features: temples present, short and rounded; scape elongate (about as long as antennomeres II + III); eyes large (as long as scape + pedicel), moderately wide and moderately convex; pronotum subcordiform without distinct anterior emargination, anterior half widened, lateral margins sinuate; anterolateral angles moderately produced, rounded; pronotal pad distinct; posterolateral angles obtuse, rounded; pronotal lobe indistinct without noticeable median notch; body slender and elongate; elytra more or less flat; color dull, dark rufopiceous; distribution: Belize, Guatemala.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF936E2B2A8DFB846D782DDA.taxon	description	Description. Measurements (in mm) — GHW: 0.70 – 0.73; HL: 0.66; AL: 1.16 – 1.18; GPW: 0.70 – 0.76; PL: 0.66 – 0.70; GEW: 0.96 – 1.00; EL: 3.50 – 3.72; TL: 4.82 – 4.88. Head (Fig. 17 B) wide, (GHW / HL: 1.06 – 1.10); eyes large (as long as scape + pedicel), moderately wide and moderately convex; temples short and rounded; occiput with slight raised areas; antennae shorter than head and pronotum length combined; antennal scape elongate (about as long as antennomeres II + III); pedicel small, submoniliform; antennomeres III – V more elongate and apically dilated; VI – X subtriangular, gradually increasing in size; XI subpyriform with apex narrowed; lateral mandibular tooth absent. Pronotum (Fig. 17 B) wide (GPW / PL: 1.06 – 1.09); anterior margin without noticeable emargination; anterolateral angles rounded, slightly produced; lateral margins sinuate; posterior half contracted; posterolateral angles obtuse and rounded; pronotal lobe slight without prominent median notch; pronotal pad (Fig. 17 C) present; pronotal pits shallow without noticeable grooves. Scutellar shield subhexagonal, transverse, less than twice as wide as long. Elytra (Fig. 17 A) elongate (EL / GEW: 3.65 – 3.72), more or less flat; vestiture short and dense; apical patch long and piceous, appears faint in piceous elytra. Ventral surface. Prosternal process short, not extending to the posterior of procoxae; hypomeral edges broadly separated; mesosternal process narrow and linear. Color unevenly rufopiceous, dull; ventrally rufopiceous with paler palpi, glossal sclerite, and galea; legs rufotestaceous. FIGURE 17. Hemipeplus belizensis sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale (apices of parameres and penis broken); E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. Male Genitalia (Figs 17 D – F). (Tegmen length = 0.40 mm (n = 2 )) Lobe of basale short and blunt, slender to very thin; shoulders almost straight and obtusely angled; apicale and basale with distinct suture; parameres slightly depressed near the bases and slightly arcuate apically; apices of parameres sharp and pointed; penis subequal to tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF936E2B2A8DFB846D782DDA.taxon	discussion	Remarks. Hemipeplus belizensis sp. nov. occurs sympatrically with H. anishae sp. nov. in Belize. Geographical Distribution (Fig. 61). Belize, Guatemala. FIGURE 18. Hemipeplus longiscapus Pollock, 1999 (Holotype). A. Dorsal habitus; B. Dorsal head and prothorax; C. Lateral view of head and prothorax. Image Credit. NMNH / Lourdes Chamorro.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF966E2C2A8DFC8E6A6C2FF5.taxon	materials_examined	Type Material. Lectotype, designated by Pollock (1999), in MNHN (Brazil, Goiás State); not studied, see Pollock (1999) for details. One paralectotype, female, labeled: ‘ [round label with yellow margin] Para-type / Brazil. S. Antonio da Barra (Prov. de Bahia) 1 l. xii. 1888 Gounelle coll. / Hemipeplus gounellei Grouvelle PARATYPE. / [red label] PARALECTOTYPE Hemipeplus gounellei Grouvelle design. DA Pollock 1997 / Brit. Mus. 1933 - 348. / [with a barcode] NHMUK 014663487 / [blue label] ♀ ’, in BMNH.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF966E2C2A8DFC8E6A6C2FF5.taxon	diagnosis	Diagnosis. Hemipeplus gounellei can be distinguished by its subcylindrical scape, strongly convex eyes, short temples, rounded pronotal lobe, and rufopiceous color. Hemipeplus gounellei has the following diagnostic features: temples short, rounded; scape subcylindrical; eyes medium (about as long as scape + pedicel) and wide, strongly convex; pronotum subcordiform, wide with moderate anterior emargination, anterior half widened than posterior; anterolateral angles produced, broadly rounded, slightly sinuate; pronotal pad very distinct; lateral margins sinuate; posterolateral angles obtuse, rounded; pronotal lobe distinct and rounded, without median notch; scutellar shield subpentagonal, transverse, not twice as wide as long; body elongate; elytra flat; color unevenly rufopiceous, dull; distribution: Brazil (Goiás). Geographical Distribution (Fig. 62). Brazil (Goiás).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF966E2C2A8DFF006A8028E8.taxon	materials_examined	Type Material. Holotype, male, only images sent from NMNH, labeled: ‘ Cayamas 6.1 Cuba / [red label] HOLOTYPE [in Pollock’s handwriting] Hemipeplus longiscapus Pollock [printed] D. A. Pollock 1997 / [with a barcode] USNMENT, 01595471 ’, in NMNH.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF966E2C2A8DFF006A8028E8.taxon	diagnosis	Diagnosis. Hemipeplus longiscapus can be distinguished by its elongate scape longer than eyes, short temples, and pale color. Hemipeplus longiscapus has the following diagnostic features: temples short, rounded; scape long, subequal to the combined length of antennomeres II – IV; eyes small in comparison to its long scape, moderately wide and moderately convex; pronotum subcordiform with shallow anterior emargination; anterior half widened with rounded anterolateral angles; pronotal pad present, slightly raised; posterolateral angles rounded but distinct; pronotal lobe slight; body slender and elongate; elytra flat; color uniformly testaceous; distribution: Cuba (Cayamas) (Pollock 1999). Geographical Distribution (Fig. 61). Cuba (Cayería las Cayamasa).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF966E2F2A8DF9B06AB52F11.taxon	materials_examined	Type Material. Holotype, sex unknown, only images sent from NMNH, labeled: ‘ ex VILLA GONZALEZ TAMP. MEX. / with SABAL SP. fronds / at PROGRESSO # 1219 / 11. II. 74 74.12763 / Hemipeplus n. sp. 1 Det. M. C. Thomas 1991 / NMNH / [purple blank label] / [red blank label] / [red label] HOLOTYPE [in Pollock’s handwriting] Hemipeplus thomasi Pollock [printed] D. A. Pollock 1997 / [with a barcode] USNMENT, 01595476 ’, in NMNH. Other Material Examined. UNITED STATES OF AMERICA. Texas. Cameron Co. Sabal Palm Grove Ref. (Site 9) 25.85092 ° N 97.41604 ° W 21. ix. 2008, E. G. Riley- 111 leg., re-vegetated area, (TAMU, 1 ♀); same locality, (Site 10) 25.84964 ° N 97.41798 ° W 5. vi. 2010, E. G. Riley- 2100 leg., palm forest beating etc., (TAMU, 1 ♂) (Fig. 20). FIGURE 19. Hemipeplus gounellei Grouvelle, 1896 (Paralectotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax. FIGURE 20. Hemipeplus thomasi Pollock, 1999. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF966E2F2A8DF9B06AB52F11.taxon	diagnosis	Diagnosis. Hemipeplus thomasi can be distinguished from the similar-appearing H. pollocki sp. nov. (Fig. 21) by its rufopiceous color and duller appearance. Hemipeplus thomasi distantly resembles H. chaos (Fig. 15) but the latter has the anterolateral pronotal margin less produced, much larger eyes, and pronotal lobe much less significant without median notch. Hemipeplus thomasi has the following diagnostic features: temples moderate, rounded; scape submoniliform; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum subcordiform, wide with deep anterior emargination in center; anterolateral angles strongly produced, broadly rounded; pronotal pad not distinct or significantly raised; posterolateral angles obtuse, rounded; pronotal lobe not very distinct but with median notch; body wide and elongate; elytra flat; color rufous with darker elytra; distribution: Mexico (Tamaulipas), USA (Texas: Cameron Co.). Re-description. Measurements (in mm) — GHW: 0.85 – 0.90; HL: 0.85 – 0.96; AL: 1.28 – 1.40; GPW: 0.98 – 1.03; PL: 0.80 – 0.83; GEW: 1.26 – 1.33; EL: 4.36 – 4.64; TL: 6.01 – 6.43. Head (Fig. 20 B) slightly longer than wide (GHW / HL: 0.94 – 1.00); eyes large (longer than scape + pedicel), moderately wide and moderately convex; temples, short and round; occiput without much raised areas; antennae shorter than head and pronotum length combined; antennal scape short, submoniliform, apically dilated, and produced externally on one side; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter, moniliform, apically dilated; V rounder, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after median notch; lateral mandibular tooth absent. Pronotum (Fig. 20 B) wide (GPW / PL: 1.23 – 1.24); anterior margin with deep median emargination; anterolateral angles rounded and strongly produced; lateral margins sinuate; posterior half contracted; posterolateral angles obtuse and rounded, very slightly produced; pronotal lobe slight with median notch; pronotal pad (Fig. 20 C) not significant; pronotal pits deep with significant elliptical grooves. Scutellar shield subpentagonal to subhexagonal with convex anterior margin, transverse, less than twice as wide as long. Elytra (Fig. 20 A) elongate (EL / GEW: 3.46 – 3.49) and flat; vestiture short and dense; apical patch long and rufous (not visible on rufous elytra) to rufopiceous. Ventral surface. Prosternal process short, not extending to the posterior of procoxae; hypomeral edges widely separated; mesosternal process narrow and linear. Color rufous with weak piceous shades on elytra; about uniformly rufous ventrally. Male Genitalia (Figs 20 D – F). (Tegmen length = 0.58 mm (n = 1 )) Lobe of basale short, slender with a clubbed apex; shoulders slanted, obtusely rounded; apicale and basale with distinct suture; parameres with strongly arcuate apices; penis broad, expanded near basal half and reduced at the apex, forming a wedge-shaped tip, length subequal to tegmen. Geographical Distribution (Fig. 61). Mexico (Tamaulipas), USA (Texas: Cameron Co.) (new country record). Natural History. According to specimen label data, H. thomasi has been found associated with Sabal palmetto; adults have been collected by beating in palm-forest and re-vegetated area (s).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF956E312A8DFA15688C2EE5.taxon	description	(Figs 21, 61)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF956E312A8DFA15688C2EE5.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ MEX: Sonora: Nacori Chico, Rancho los Pescados, 13 APR 92 Col: E. Joyal (ASU) / 29 ° 29 ’ N, 108 ° 59 ’ W, 780 m. elev., in new leaves Sabal mesana [uresana] (Arecaceae) / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus pollocki KC’, in ASU. Twenty-four paratypes: 2 females, same label as holotype, in ASU; 12 specimens [1 without head, 2 without head and prothorax], sex unknown, same label as holotype, ‘ / Oedmeridoe? [Oedemeridae] Det. M. Catier’ [on one specimen], in ASU; 10 specimens [1 without head], sex unknown, labeled: ‘ MEX: Sonora, Yecora, Rancho el Palmer de Onavas, 16 OCT 91 Col: E. Joyal (ASU) / 28 ° 29 ’ 30 ’’ N 109 ° 23 ’ W 700 m. elevation, in new leaves Sabal mesana [uresana] (Arecaceae) / Hemipeplus det. M. Gimmel 2015 [in one specimen] ’, in ASU. Derivation of Specific Epithet. The specific name (noun in the genitive case) is given in allusion to Darren A. Pollock, for the help he provided to the first author during his Master’s graduate studies on Hemipeplinae, and for his contributions to the taxonomy of Hemipeplinae. FIGURE 21. Hemipeplus pollocki sp. nov. (Holotype) A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF956E312A8DFA15688C2EE5.taxon	diagnosis	Diagnosis. Hemipeplus pollocki sp. nov. can be distinguished by its rufous shiny color compared to the duller rufopiceous color of the similar-appearing H. thomasi (Fig. 20). Hemipeplus pollocki sp. nov. has the following diagnostic features: temples present, prominent and rounded; scape submoniliform; eyes moderately large (as long as scape + pedicel) but very shallow, moderately convex; pronotum subcordiform, wide with a deep anterior emargination, anterior half widened, lateral margins sinuate; anterolateral angles produced, broadly rounded; pronotal pad narrow, not significantly raised; posterolateral angles square, rounded; pronotal lobe indistinct without median notch; pronotal pits deep with strong grooves; body long and wide; elytra flat; color shiny, rusty brown / rufous dorsally, elytra dark rufous; distribution: Mexico (Sonora).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF956E312A8DFA15688C2EE5.taxon	description	Description. Measurements (in mm) — GHW: 0.82 – 0.96; HL: 0.75 – 0.91; AL: 1.18 – 1.30; GPW: 0.91 – 1.05; PL: 0.75 – 0.93; GEW: 1.26 – 1.50; EL: 4.04 – 4.84; TL: 5.54 – 6.68. Head (Fig. 21 B) wide (GHW / HL: 1.05 – 1.09); eyes moderately large (as long as scape + pedicel), shallow and moderately convex; temples, short and round; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape, submoniliform, apically dilated, and produced externally on one side; pedicel small moniliform; antennomere III elongate and apically dilated; IV much shorter, moniliform, apically dilated; V short, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after median notch; lateral mandibular tooth absent or extremely small. Pronotum (Fig. 21 B) wide (GPW / PL: 1.13 – 1.21); anterior margin with deep median emargination; anterolateral angles rounded and produced; lateral margins sinuate; posterior half contracted; posterolateral angles square and rounded; pronotal lobe slight / truncate without median notch; pronotal pad (Fig. 21 C) not significant or well raised; pronotal pits deep and dark with significant oval grooves. Scutellar shield convex or subpentagonal, transverse, about twice as wide as long. Elytra (Fig. 21 A) elongate (EL / GEW: 3.20 – 3.23) and flat; vestiture short and scarce; apical patch long and darker than elytra; Ventral surface. Prosternal process short, not extending to the posterior of procoxae; hypomeral edges widely separated; mesosternal process narrow and linear. Color rufous, shiny, elytra darker; about uniformly brown ventrally, inner edge of hypomeron piceous, elytral epipleura piceous, tarsi slightly paler. Male Genitalia (Figs 21 D – F). (Tegmen length = 0.65 mm (n = 1 )) Lobe of basale very short and apically rounded; shoulders almost straight, rectangular, without a slant; apicale and basale with distinct suture; apicale long, weakly sinuate, each half forked into two parts, a shorter paramere and a much longer arcuate paramere; apices of parameres sharp and pointed; penis shorter than tegmen. Variation. Some specimens have anterolateral pronotal angles more produced compared to others.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF956E312A8DFA15688C2EE5.taxon	discussion	Remarks. Despite significant morphological differences, the sheen and body shape of Hemipeplus pollocki sp. nov. are similar to those of H. anishae sp. nov. from Belize, suggesting a possible affinity between the two species. Geographical Distribution (Fig. 61). Mexico (Sonora). Natural History. According to specimen label data, H. pollocki sp. nov. was collected associated with Sabal uresana Trel. (Arecaceae).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF8B6E322A8DFA816A7F2DBF.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ ARGENTINA, Cha., 2 km. N. Jcn. to I. Cerrito 1 - 24 - 1989, CW & L. O’Brien & G. Wibmer / Hemipeplus n sp. 5 Det. M. C. Thomas 1992 / MLPA / [blank red label] / [green label] FSCA / [blue label] ♀ / [red label] HOLOTYPE Hemipeplus argentinus Pollock D. A. Pollock 1997 ’, in FSCA.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF8B6E322A8DFA816A7F2DBF.taxon	diagnosis	Diagnosis. Hemipeplus argentinus can be distinguished from the similar-appearing H. bolivianus (Figs 23.1, 23.2) by its slightly elongate scape and distinct anterior emargination on the pronotum. Hemipeplus argentinus has the following diagnostic features: temples moderately long, rounded; scape submoniliform, slightly elongate, apically dilated; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum subcordiform, with a deep anterior emargination, anterior half much more widened than posterior; anterolateral angles strongly produced, broadly rounded; pronotal pad distinct; posterolateral angles obtuse, rounded; pronotal lobe not very prominent but with a slight median notch; scutellar shield subhexagonal, transverse, about twice as wide as long; body elongate; elytra slightly convex; color rufous, dull; distribution: Argentina (Entre Ríos Province). Geographical Distribution (Fig. 62). Argentina (Entre Ríos Province). FIGURE 22. Hemipeplus argentinus Pollock, 1999 (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF896E352A8DFF006DD72F0F.taxon	description	FIGURE 23. 1. Hemipeplus bolivianus Pollock, 1999 (Ecuador). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. FIGURE 23. 2. Hemipeplus bolivianus Pollock, 1999 (Bolivia). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF896E352A8DFF006DD72F0F.taxon	materials_examined	Type Material. Holotype, female, in SEMC (Bolivia, Beni Department); see Pollock (1999) for details. One paratype, female, labeled: ‘ BRAZIL: Rondonia. 62 km SW Ariquemes, nr Fzda. Rancho Grande 20 – IX – 1992 U. Schwitz, BL Trap / [blue label] PARATYPE Hemipeplus bolivianus Pollock’, in FSCA. Other Material Examined. BOLIVIA. Cochabamba Department. Villa Tunari, Hotel el Puente 16 ° 59 ’ S, 65 ° 324 ’ W, 1158 m. 10 – 12. ix. 2012 P. Skelly J. Wappes, T. Bonaso, C. Hamel leg., (FSCA, 2 ♂, 2 sex unknown) (Fig. 23.2); Santa Cruz Department. Potrerillos del Guenda; 40 km. NW. Santa Cruz de la Sierra; 350 – 400 m; 17 ° 40 ’ S 063 ° 27 ’ W 7 – 9. ix. 2012, P. Skelley, J. Wappes, T. Banaso, C. Hamel leg., (FSCA, 1); Santa Cruz 4 – 6 k SSE Buena Vista F & F Hotel, 1 – 8. xi. 2002, J. E. Wappes leg., (FSCA, 1 ♂). ECUADOR. Napo. 25 km. e. Puerto Napo, 450 m.?. ix. 1997, B & B. Valentine leg., Cabanas Alinahui u-v light, (FSCA, 1 ♂) (Fig. 23.1). COLOMBIA (photograph only, needs verification). Vichada Department. PNN El Tuparro — Centro Administrativo; 5 º 21 ’ N 67 º 51 ’ W 100 m, 08 – 28. viii. 2000, Villalba W. leg., Malaise trap, (IAvH-E, 1).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF896E352A8DFF006DD72F0F.taxon	diagnosis	Diagnosis. Hemipeplus bolivianus can be distinguished from the similar-appearing H. argentinus (Fig. 22) by its moniliform scape and shallower anterior emargination on the pronotum. Hemipeplus bolivianus has the following diagnostic features: temples moderately long, rounded; scape very short and moniliform; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum subcordiform, with shallow anterior emargination; punctures wider than in H. argentinus; anterior half more widened than posterior; anterolateral angles produced, broadly rounded; pronotal pad distinct; posterolateral angles obtuse, rounded; pronotal lobe indistinct with faint to no median notch; scutellar shield subhexagonal, transverse, twice as wide as long; body elongate; color uniformly rufotestaceous; elytra slightly convex; distribution: Bolivia, Brazil (Pollock 1999), Colombia?, Ecuador, Peru (Pollock 1999). Male Genitalia (Figs 23. 1 D – F, 23. 2 D – F). (Tegmen length = 0.30 – 0.35 mm (n = 4 )) Lobe of basale moderately long, thin, pin-like with rounded apex; shoulders almost straight, acutely angled; apicale and basale with distinct suture; parameres straight, diverging outward; apices of parameres straight, sharp and pointed; penis much longer than tegmen, slightly narrowed medially.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF896E352A8DFF006DD72F0F.taxon	description	Variation. Some specimens have slightly deeper emargination on anterior pronotal margin; there is a slight median notch on truncate pronotal lobe of some specimens; parameres are longer in some specimens than others.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF896E352A8DFF006DD72F0F.taxon	discussion	Remarks. Hemipeplus bolivianus is a widespread species in South America. Geographical Distribution (Fig. 62). Bolivia, Brazil (Pollock 1999), Colombia?, Ecuador (new country record), Peru (Pollock 1999). Natural History. Hemipeplus bolivianus is most likely associated with palms (Pollock 1999). According to specimen label data, adults have also been collected from UV light trap and malaise trap (s).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF8C6E372A8DFC286AAA2DC7.taxon	description	(Figs 24, 63)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF8C6E372A8DFC286AAA2DC7.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ Madagascar: (Sud) Pays Androy (Nord) Alluaud 1900 | 36 / [blue label] MUSEUM PARIS Coll. Ch. Alluaud / [blue label] ♀ / [red label] HOLOTYPE ♀ Eurypeplus quadrimaculatus KC & Pollock’, in MNHN. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the presence of four pale spots on the elytra of this species.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF8C6E372A8DFC286AAA2DC7.taxon	diagnosis	Diagnosis. Eurypeplus quadrimaculatus gen. nov., sp. nov. can be distinguished by its elongate pronotum with straight anterior margin and four pale spots on the elytra. Eurypeplus quadrimaculatus gen. nov., sp. nov. has the following diagnostic features: temples short and rounded; eyes large (longer than scape + pedicel), wide, and strongly convex; head wide; pronotum elongate, laterally sinuate, without anterior emargination; anterolateral angles indistinct, broadly rounded; pronotal pad absent; posterolateral angles square, rounded; pronotal lobe short, rounded without median notch; elytra less flat than in Hemipeplus, laterally expanded; color rufopiceous with four large elliptical pale spots on elytra; distribution: Madagascar.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF8C6E372A8DFC286AAA2DC7.taxon	description	Description. Measurements (in mm) — GHW: 0.83; HL: 0.66; AL: 1.58; GPW: 0.73; PL: 0.78; GEW: 1.26; EL: 3.09; TL: 4.53. Head (Fig. 24 C) wide, (GHW / HL: 1.26) wide; eyes large (longer than scape + pedicel), wide and strongly convex; temples distinct, short and rounded; occiput without raised areas; antennae longer than head and pronotum length combined; antennal scape short and moniliform; pedicel small moniliform; antennomere III more elongate than II; IV – VII subfiliform and apically dilated; VIII – X subtriangular; XI subpyriform with apex narrowed; lateral mandibular tooth unobservable in the specimen. Pronotum (Fig. 24 B) narrow and elongate (GPW / PL: 0.94); anterior margin without emargination; anterolateral angles rounded, indistinct; lateral margins sinuate; posterior half contracted; posterolateral angles rounded and square; pronotal lobe short, rounded without median notch; pronotal pad (Fig. 24 F) absent; pronotal pits shallow without visible grooves. Scutellar shield raised above elytral level, pentagonal, transverse, less than twice as wide as long. Elytra (Fig. 24 A) elongate (EL / GEW: 2.45), widened laterally at the halfway point, disc convex; vestiture moderate; apical patch absent. Ventral surface (Figs 24 D – E). Prosternal process short, not extending to the posterior of procoxae (Fig. 24 E); hypomeral edges widely separated; mesanepisternum broad and flat; mesosternal plate much narrower than in Hemipeplus and Holopeplus; mesosternal process linear and wider than in most Hemipeplinae; abdominal process wider than in Hemipeplus and Holopeplus; (at least) female with a median fovea in V 4 (Fig. 24 D). Color dull, rufopiceous with four large elliptical pale spots on elytra; antennomeres II – VII pale while rest of the antenna rufopiceous; ventrally labial palpi, glossal sclerite, tibia, tarsi pale; abdominal sternites I – III less dark otherwise rufopiceous.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF8C6E372A8DFC286AAA2DC7.taxon	discussion	Remarks. Eurypeplus quadrimaculatus gen. nov., sp. nov. is the only species of Hemipeplinae with elytra exhibiting distinctive light-colored maculae. Geographical Distribution (Fig. 63). Madagascar. FIGURE 24. Eurypeplus quadrimaculatus gen nov., sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Dorsal head view in horizontal plane; D. Ventral habitus; E. Ventral habitus closeup; F. Lateral view of head and prothorax; G. Female genitalia (with scale bar).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF826E392A8DFF006B6C28CA.taxon	description	FIGURE 25. Hemipeplus reichertae Pollock, 1997 (Paratype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF826E392A8DFF006B6C28CA.taxon	materials_examined	Type Material. Holotype, sex unknown, in MRAC (Congo); not studied, see Pollock (1997) for details. One paratype, female, labeled: ‘ Sangmelima, Cameroons, A. I. Good. / April 1933 / Carn. Mus. Acc 10080 / [blue label] PARATYPE Hemipeplus reichertae Pollock’, in CMNH (Fig. 25).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF826E392A8DFF006B6C28CA.taxon	diagnosis	Diagnosis. Hemipeplus reichertae can be distinguished by its large eyes, lack of temples, subcordiform pronotum without prominent posterior lobe, and pale color. Hemipeplus reichertae has the following diagnostic features: temples absent; scape short, submoniliform, apically dilated and externally produced on one side; eyes large (about as long as scape + pedicel + antennomere III), wide and convex; pronotum subcordiform, wide and flat with a distinct anterior emargination; anterior half wider than posterior; anterolateral angles strongly produced, broadly rounded; pronotal pad present but not very prominent; posterolateral angles obtuse, rounded; pronotal lobe slight without median notch; scutellar shield subheptagonal, transverse, about twice as wide as long; body long, slightly widened at apical half of elytra; elytra slightly convex; color uniformly brown; distribution: west-central Africa (Pollock 1997). Geographical Distribution (Fig. 63). Cameroon, Democratic Republic of Congo, Equatorial Guinea, Namibia, Republic of Congo (Pollock 1997).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF836E3A2A8DFCAC6A972DEC.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ [round label with red margin] Type / [card with glass cover slip and dissected female ovipositor] Hemipeplus egregius Arrow. TYPE. ♀ V. ovipos. / “ Germ ” E. Africa Kilindi. 15. iv. 14. / Pres, by Imp. Bur. Ent. Brit. Mus. 1923 - 383. / Hemipeplus egregius, type Arrow / [red label] HOLOTYPE Hemipeplus egregius Arrow exam. D. A. Pollock ’ 95 / [with a barcode] NHMUK 014663478 ’, in BMNH. Other Material Examined. SOUTH AFRICA. KwaZulu-Natal. Southbroom, 24. vii. 1933, (NHMB, 1 ♂) (Fig. 26). Limpopo. Tvl. Hans Merensky Nat. Res., 23.42 S 30.44 E. 23 – 25. i. 1987, R. Oberprieler leg., leaves of Phoenix reclinata, (BMNH, 1). MALAWI. Chiromo. R. C. Wood leg., Pres. by Imp. Inst. Ent. B. M. 1936 – 27, (BMNH, 1).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF836E3A2A8DFCAC6A972DEC.taxon	diagnosis	Diagnosis. Hemipeplus egregius can be distinguished by its large strongly convex eyes, lack of temples, acutely produced pronotal angles, prominent pronotal lobe, and dark brown color. Hemipeplus egregius has the following diagnostic features: temples absent or not prominent; scape submoniliform; eyes large (longer than scape + pedicel), very wide and strongly convex; pronotum subcordiform, wide and flat with a distinct anterior emargination; pronotum strongly sinuate laterally, anterior half strongly produced; anterolateral angles acutely rounded; pronotal pad absent; posterolateral angles acute; pronotal lobe prominent with a deep median notch; scutellar shield U shaped, slightly wider than long; elytra more or less flat; color uniformly testaceous to rufotestaceous; distribution: southern Somalia to South Africa (Natal) (Pollock 1997). Male Genitalia (Figs 26 D – F). (Tegmen length = 0.62 mm (n = 1 )) Lobe of basale short, thick, and wide, apically dilated; shoulders very slanted and not angulate; apicale and basale fused without a distinct suture; parameres short, narrowed basally, widened apically with short, narrowed, blunt tips; penis longer than tegmen. Geographical Distribution (Fig. 63). Kenya, Malawi, Mozambique, Somalia, South Africa, Tanzania (Pollock 1997). Natural History. According to specimen label data and Pollock (1997), H. egregius adults are associated with Coconut Palm (Syagrus romanzoffiana (Cham.) Glassman) (Arecaceae) and Phoenix reclinata Jacq. (Arecaceae). FIGURE 26. Hemipeplus egregius Arrow, 1930. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF816E3C2A8DFF00683C2924.taxon	description	FIGURE 27. Hemipeplus grouvellei Pollock, 1997 (Holotype). A. Dorsal habitus; B. Dorsal head and prothorax; C. Lateral view of head and prothorax. Image Credit. NMNH / Lourdes Chamorro.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF816E3C2A8DFF00683C2924.taxon	materials_examined	Type Material. Holotype, sex unknown, only images sent from NMNH, labeled: ‘ MADAGASCAR: Prov. Fianarantsoa, 7 km W Ranomafana, 1100 m 1 – 7 November 1988 W. E. Steiner / At black light in montane rain forest / [handwritten] Comp. with type of H. madagascariensis Grouvelle & NOT the same species / [red label] HOLOTYPE [in Pollock’s handwriting] Hemipeplus grouvellei Pollock [printed] Examined by D. A. Pollock 1997 / [with a barcode] USNMENT, 01595470 ’, in NMNH.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF816E3C2A8DFF00683C2924.taxon	diagnosis	Diagnosis. Hemipeplus grouvellei can be distinguished from the similar-appearing H. egregius (Fig. 26) by its obtusely rounded posterolateral pronotal angles and less prominent pronotal lobe. Hemipeplus grouvellei has the following diagnostic features: temples absent; scape submoniliform; eyes very large (subequal to scape + pedicel + antennomere III) moderately wide and moderately convex; pronotum subcordiform, wide and flat without a distinct anterior emargination; pronotum strongly sinuate anterolaterally, anterior half strongly produced, acutely rounded; pronotal pad absent; posterolateral angles obtusely rounded; pronotal lobe broad and arcuate; elytra slightly convex apically; color uniformly light brown; distribution: Madagascar (Pollock 1997). Geographical Distribution (Fig. 63). Madagascar. Natural History. According to Pollock (1997), adults of H. grouvellei were collected from blacklights in montane rainforest.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF866E3D2A8DFCCF6DCB2DBD.taxon	materials_examined	Type Material. Holotype, sex unknown, labeled: ‘ [round label with red margin] Type H. T. / Rodrigues 1. viii-xi, 1918, H. J. Snell and H. P. Thomasset, / [label upside down] Brit. Mus. 1933 - 349. / Hemipeplus rodericensis H. Scott TYPE. / [red label] HOLOTYPE Hemipeplus rodericensis Scott exam. D. A. Pollock ’ 95 / [with a barcode] NHMUK 014663479 ’, in BMNH. Twenty-three paratypes: 8 males, 6 females, 9 sex unknown, labeled same as holotype, in BMNH (Fig. 28).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF866E3D2A8DFCCF6DCB2DBD.taxon	diagnosis	Diagnosis. Hemipeplus rodericensis can be distinguished by its elongate subquadriform pronotum and dark brown, flat, and elongate body. Hemipeplus rodericensis has the following diagnostic features: temples broad and rounded; scape subcylindrical and externally angulate on one side; eyes moderately large (as long as scape + pedicel), wide and convex; pronotum subquadriform, longer than wide to as long as wide with flat disc; laterals subparallel; anterior half about as wide as posterior half with deep anterior notch in center; anterolateral angles indistinct, rounded; pronotal pad indistinct; posterolateral angles obtuse; pronotal lobe prominent and rounded without median notch; scutellar shield broadly convex, transverse, more than twice as wide as long; body wide and long; elytra flat; color uniformly dark brown, shiny; distribution: Rodrigues Island (Mauritius). Male Genitalia (Figs 28 D – F). (Tegmen length = 0.83 mm (n = 1 )) Lobe of basale short, wide, and rounded; shoulders very slanted and not angulate; apicale and basale fused without a distinct suture; parameres very long and straight; apices slightly wider than rest of the apical halves with narrowed blunt tips; penis slightly longer than tegmen, tapered at the apex.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF866E3D2A8DFCCF6DCB2DBD.taxon	discussion	Remarks. Hemipeplus rodericensis is thought to be closely related to H. hemipterus and H. abditus from the New World based on similarities in structure of pronotum and male genitalia. Geographical Distribution (Fig. 63). Rodrigues Island (Mauritius). Natural History. Hemipeplus rodericensis is associated with Pandanus Parkinson (Pandanaceae) species (Screw Pines) (Scott 1933; Pollock 1997). FIGURE 28. Hemipeplus rodericensis Scott, 1933 (Paratype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF846E3F2A8DFF006A462841.taxon	materials_examined	Type Material. Holotype, sex unknown, only images sent from MNHN, labeled: ‘ [light green blue label] Madagasc. / [in Grouvelle’s handwriting] Hemipeplus madagascariensis 5 Grouv / [red label] TYPE / [light blue label] MUSEUM PARIS 1917 Coll. GROUVELLE / [red label] HOLOTYPE [handwritten] Hemipeplus madagascariensis Grouv. Examined by D. A. Pollock 1997 / [with a barcode] MNHN, Paris EC 26369 ’, in MNHN (Grouvelle Collection).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF846E3F2A8DFF006A462841.taxon	diagnosis	Diagnosis. Hemipeplus madagascariensis can be distinguished by its large eyes, large, rounded temples, and distinct pronotal lobe. Hemipeplus madagascariensis has the following diagnostic features: temples very large (about 1 / 3 eye length) and rounded; scape moniliform externally produced on one side; eyes very large (subequal to scape + pedicel + antennomere III), very wide and convex; pronotum subcordiform, wide and flat without distinct anterior emargination; pronotum strongly sinuate anterolaterally, anterior half strongly produced and wider than posterior half; anterolateral angles narrowly rounded; pronotal pad present; posterolateral angles rounded; pronotal lobe broad and rounded without median notch; scutellar shield slightly wider than long, shield-shaped; elytra slightly convex apically; color uniformly light brown; distribution: Madagascar (Pollock 1997). Geographical Distribution (Fig. 63). Madagascar.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF856E402A8DFD256DCB2D1E.taxon	materials_examined	Type Material. Holotype, sex unknown, in ZMHB (Guinea); not studied, see Grouvelle (1919) or Pollock (1997) for details. Other Material Examined. REPUBLIC OF THE CONGO. Sangha Department. Kabo. 2.10 ° N 16.10 ° E environs Bomassa 07 – 09. iv. 1993, D. H. Chadwick leg., (MAIC, 1 ♂ (Fig. 30), 2 sex unknown). UGANDA. Ruwenzori Range. Semliki Forest. 2,850 ft. 22. viii. – 3. ix 1952. D. S. Fletcher. leg., Ruwenzori Exped. B. M. 1952 – 566, (BMNH, 1). GABON. Woleu-Ntem Province. Okano. Mikongo (Rougier), Monts de Cristal, secondary forest 0 ° 29 ’ 47.0 ” N 11 ° 10 ’ 42.0 ” E 430 m, 28. vii – 12. viii. 2019, Albert, J-L., Aristophanous, M., Biemba, J., Dérozier, V., Morerro, P. leg., MV light, BMNH (E) 2020 – 19, (BMNH, 1). CAMEROON. Southwest Region. Fako. 2 km N Bakingili 200 m. 28 – 30. vii. 1984, J. Rawlins, J. DiGiulio leg., (CMNH, 1).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF856E402A8DFD256DCB2D1E.taxon	diagnosis	Diagnosis. Hemipeplus suturalis can be distinguished by its acutely pointed temples and pronotum without pronotal pad. Hemipeplus suturalis has the following diagnostic features: temples short and strongly acute; scape submoniliform and externally produced on one side; eyes moderately large (as long as scape + pedicel), wide and convex; pronotum wide, flat, and subcordiform with distinct emargination on anterior margin; anterior half wider than posterior half; anterolateral angles strongly produced, broadly rounded; pronotal pad absent; posterolateral margins subparallel; posterolateral angles obtuse; pronotal lobe not prominent / truncate and without median notch; scutellar shield subhexagonal, transverse, less than twice as wide as long; body short and flat; elytra more or less flat; color dark testaceous, shiny, pronotum piceous – rufopiceous, head rufous – rufopiceous; distribution: west-central Africa (Cameroon, Republic of the Congo, Uganda) (Pollock 1997). Male Genitalia (Figs 30 D – F). (Tegmen length = 0.50 mm (n = 1 )) Lobe of basale extremely short, thick, and wide; shoulders very slanted and obtusely angled; apicale and basale with distinct suture; parameres very long and almost straight with minimum sinuation near basal half; apices very thin and sharp, directed inward; penis shorter than tegmen tapering toward apex.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF856E402A8DFD256DCB2D1E.taxon	discussion	Remarks. Hemipeplus suturalis resembles H. quadricollis and H. pseudoquadricollis from the New World except for the subcordiform pronotum. Geographical Distribution (Fig. 63). West-central Africa (Cameroon, Democratic Republic of the Congo, Gabon, Republic of the Congo, Uganda) (Pollock 1997). Natural History. According to specimen label data, adults of H. suturalis have been collected at mercury vapor light and in secondary forest. FIGURE 30. Hemipeplus suturalis Grouvelle, 1919. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFFB6E422A8DFF006AD52DCE.taxon	materials_examined	Type Material. Holotype, sex unknown, labeled: ‘ [round label with red margin] Type H. T. / [underlined in blue] N. W Rhodesia [Zimbabwe]: Nama-ula. 28. VIII. 1914. H. C. Dollman. / [label upside down] H. C. Dollman Coll. 1919 – 79 / Hemipeplus dollmani H. Scott TYPE. / [red label] HOLOTYPE Hemipeplus dollmani Scott exam. D. A. Pollock ‘ 95 ’, in BMNH. Eight paratypes: 3 specimens, female, same label as holotype, in BMNH; 1 specimen, female, labeled: ‘ [underlined in blue] N. W. Rhodesia: Nama-ula. 1. IX. 1914. H. C. Dollman. / H. C. Dollman coll. 1919 – 79 / Hemipeplus dollmani H. Scott Paratype / [abdomen in a slide] Hemipeplus dollmani H. Scott. ♀ B. ’, in BMNH; 1 specimen, female, labeled: ‘ [underlined in blue] N. W. Rhodesia: Nama-ula. 2. IX. 1914. H. C. Dollman. / H. C. Dollman coll. 1919 – 79 / Hemipeplus dollmani H. Scott Paratype’, in BMNH; 1 specimen, female, labeled: ‘ [underlined in blue] N. W. Rhodesia Lukendo. 27. viii. 1913. H. C. Dollman. [underside of label] mopani bark / H. C. Dollman coll. 1919 – 79 / Hemipeplus dollmani H. Scott Paratype’, in BMNH; 2 specimens, female, labeled: ‘ BELGIAN CONGO Lunda, 20. v. 1929, Dr. H. S. Evans [something illegible] / Pres. by Imp. Inst. Ent. 1933 - 382 / Hemipeplus dollmani H. Scott Paratype / 77.29 ’, in BMNH. Other Material Examined. SOUTH AFRICA. Gauteng. Pretoria. Renosterkop N of Faerie Glen, 1400 m. 25.46 S 28.19 E, 01. ix. 1986, R. Oberprieler leg., collected off Setaria lindenbergiana?, (BMNH, 2); 10 – 12. x. 1986, W. Wittmer leg., (NHMB, 10); Wapadrand 17. xi. 1987 [no year on 2 specimens], W. Wittmer leg., (NHMB, 5). KwaZulu-Natal. uThukela District. Frere, 8. xi. 1992, W. Wittmer leg., (NHMB, 1 ♂) (Fig. 31). ZAMBIA. Congo-Zambesi Watershed. 1929. Dr. H. S. Evans, Pres. By Imp. Inst. Ent. 1932 – 147, (BMNH, 1). Southern Province. Choma District. Bruce-Miller Farm Campsite (Southern Miombo Woodland) 16 ° 38 ’ 12 ’’ S, 27 ° 01 ’ 30 ’’ E, 10 – 14. x. 2018, Carter, M., McDougall, S., Miles, W. Mulvaney, L. leg., LepiLED Light Trap, ANHRT: 2018.35 BMNH (E) 2020 – 19, (BMNH, 2). Muchinga Province. Kalungu District. 1280 m. Kalungu, north of Iskoa, 9 ° 40 ’ 52 ’’ S, 32 ° 42 ’ 50 ’’ E, 22 – 24. xi. 2016, Smith, Takano & Oram leg., light trap, BMNH (E) 2016 – 251 ANHRT 23: ZM- 009, (BMNH, 1). ZIMBABWE. Mashonaland West. Kariba District. Southern Rhodesia, Kariba, 1675 ft., 4. xi. 1968, D. R. Birkenmeyer leg., blacklight trap, Rhodesian bush, (FSCA, 1).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFFB6E422A8DFF006AD52DCE.taxon	diagnosis	Diagnosis. Hemipeplus dollmani can be distinguished by its small temples, distinct pronotal pad, and shiny, pale color. Hemipeplus dollmani has the following diagnostic features: temples short and indistinct, often barely noticeable; scape short and moniliform; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum short, slightly wide, and subcordiform with distinct emargination anteriorly and sinuation laterally, anterior half wider than posterior half with deep anterior emargination in center; anterolateral angles produced, rounded; pronotal pad very distinct, broad and setose; posterolateral angles obtuse, rounded; pronotal lobe indistinct without median notch; scutellar shield subhexagonal, transverse, about twice as wide as long; body narrow and long; elytra slightly convex; color pale, shiny; distribution: Africa between 7 ° N and 25 ° S (Pollock 1997). Male Genitalia (Figs 31 D – F). (Tegmen length = 0.28 mm (n = 1 )) Lobe of basale short, slender with rounded end; shoulders straight and rounded; apicale and basale subequal (excluding the basale lobe) with distinct suture; parameres long and about straight with slightly arcuate apices; apices thin and sharp; penis very wide and longer than tegmen, narrowed apically.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFFB6E422A8DFF006AD52DCE.taxon	description	Variation. Temples are more distinct in some specimens of H. dollmani than others.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFFB6E422A8DFF006AD52DCE.taxon	discussion	Remarks. Hemipeplus dollmani resembles H. australicus from Australia but has temples; it resembles H. labuanensis sp. nov. (Malaysia) and H. sinensis sp. nov. (China) but the pronotal pad is wider with denser setae. Geographical Distribution (Fig. 63). Cameroon, Democratic Republic of the Congo, Gabon, Republic of the Congo, South Africa, Zambia, Zimbabwe [Africa between 7 ° N and 25 ° S] (Pollock 1997). Natural History. According to specimen label data, H. dollmani is associated with Setaria lindenbergiana? (Nees) Stapf (Poaceae). FIGURE 31. Hemipeplus dollmani Scott, 1933. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF96E452A8DFF006D6A2D9E.taxon	materials_examined	Type Material. Holotype, sex unknown, only images sent from MNHN, labeled: ‘ [blue label] Gabon [with other illegible word (s)] / [pink] Type / Hemipeplus africanus Grouv. / MUSEUM PARIS 1917 Coll. GROUVELLE / [red label] HOLOTYPE Hemipeplus africanus Grouv. Examined by D. A. Pollock 1997 / [with a barcode] MNHN, Paris EC 26370 ’, in MNHN (Grouvelle Collection). Other Material Examined. CAMEROON. Southwest Region. Fako Division. Upper Farm Buea elevation 4500 ft., 9 – 20. v. 1949, B. Malkin leg., (DAPC, 1); 2 km N Bakingili 200 m. 28 – 30. vii. 1984, J. Rawlins, J. DiGiulio leg., (CMNH, 1). CÔTE D’IVOIRE. Danané Department. Yeale Village, Mt. Nimba, 07 ° 31 ’ 35.3 ’’ N, 08 ° 25 ’ 20.1 ’’ W, 18 – 29. iv. 2016, Aristophanous, M., Geiser, M., Moretto, P., leg., light trap, BMNH (E) 2016 – 109, TripRef: CI- 003 (ANHRT 17), (BMNH, 1 ♂) (Fig. 32.2). GABON. Woleu-Ntem Province. Okano. Mikongo (Rougier), Monts de Cristal, secondary forest 0 ° 29 ’ 47 ’’ N, 11 ° 10 ’ 42 ’’ E 430 m, 28. vii – 12. viii. 2019, Albert, J-L., Aristophanous, M., Bie Mba, J., Dérozier, V., Morerro, P. leg., MV light, BMNH (E) 2020 – 19, (BMNH, 1 ♂). GHANA. Western Region. Prestea-Huni Valley District. Pretsia, 24 m 4 ° 55 ’ N 1 ° 52 ’ W, 25. viii. 1969, Dr. S. Endrody-Younga leg., (DAPC, 1 ♀). Ashanti Region. Kumasi Metropolitan. Kumasi, Nhiaau 330 m, 6 ° 43 ’ N 1 ° 36 ’ W, 1 – 7. iii. 1967, Dr. S. Endrody-Younga leg., light trap, (DAPC, 1). GUINEA. Kindia Region. Coyah Prefecture. 1. viii – 12. xii. 1968, K. Perenoz leg., (DAPC, 1). Kindia Prefecture. Damakanya, 29 – 30. iv. 1951, Exped. Mus. G. Frey, Franz Guinea 1951 W. Afr., Bechvne leg., (NHMB, 1 ♂) (Fig. 32.1). NIGERIA. Oyo. Ibadan, iv. 1956, V. F. Eastop leg., C. E. Tottenham Collection B. M. 1974 – 587, (BMNH, 1). Niger. Tafa, 21. v. 1957, V. FE leg., C. E. Tottenham Collection, BM 1974 – 587, (BMNH, 1). UGANDA. Ruwenzori Range. Semliki Forest. 2850 ft. 22. vii. – 3. ix. 1952. D. S. Fletcher leg., (BMNH, 1). Bwamba. Hakitengya,?. ii – iii. 1949, W. H. R. Lumsden leg., from suction trap, Brit. Mus. 1949 – 576, (BMNH, 1). Kampala. Kampala, 7. vi. 192, H. Hargreaves leg., (BMNH, 1 ♀).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF96E452A8DFF006D6A2D9E.taxon	diagnosis	Diagnosis. Hemipeplus africanus can be distinguished from the similar-appearing H. alluaudi (Figs 33.1, 33.2) by its subparallel posterolateral pronotal margins and darker color. Hemipeplus africanus has the following diagnostic features: temples short and angulate to somewhat rounded, slightly larger in some specimens; scape moniliform; eyes large (longer than scape + pedicel), wide to moderately wide; pronotum wide to relatively narrow, subcordiform; anterior half wider than posterior half with deep to shallow anterior emargination in center; anterolateral angles moderately produced, broadly rounded; pronotal pad present but narrow; hind angles sharply acute to less so; posterolateral margins of pronotum straight and subparallel; pronotal lobe indistinct and rounded without median notch; scutellar shield subhexagonal to elliptical, transverse, less than twice as wide as long; body long; elytra slightly convex; color rufotestaceous to rufopiceous; distribution: widespread in Africa between 10 ° N and 5 ° S (Pollock 1997). Male Genitalia (Figs 32.1 D – F, 32.2 D – F). (Tegmen length = 0.35 – 0.43 mm (n = 2 )) Lobe of basale short with rounded end, a little more elongate than in H. alluaudi; shoulders slanted and obtusely angled (an extra layer from basale pointed upward sometimes covers it making it look acutely pointed (Fig. 32.2 E )); apicale and basale with somewhat clear suture; basale subequal to apicale (not including basal lobe); parameres long and almost straight, apices curved inward; apices thin and sharp; penis wide, much longer than tegmen, apically narrowed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF96E452A8DFF006D6A2D9E.taxon	description	Variation. Some specimens have pointed and acute hind angles of pronotum while it is less so in some; emargination on anterior pronotum is fairly deep to shallow; body color ranges from uniformly lighter brown to dark brown with piceous shade; the size of eye varies as some specimens with shallower eyes were seen; females probably have smaller / shallower eyes.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF96E452A8DFF006D6A2D9E.taxon	discussion	Remarks. Hemipeplus africanus closely resembles H. klematanicus, H. pseudoklematanicus sp. nov. and H. nigerrimus sp. nov. from the Austro-Oriental region and might be difficult to distinguish without examination of male genitalia or locality data. Geographical Distribution (Fig. 63). Cameroon, Côte d’Ivoire, Democratic Republic of the Congo, Gabon, Ghana, Guinea, Nigeria, Rwanda, Tanzania, Uganda (Pollock 1997). Natural History. According to specimen label data, adults of H. africanus have been collected from light traps including mercury vapor light (s) in secondary forest (s), and suction trap (s). Specimens have also been collected by beating and aerial collecting according to Pollock (1997). FIGURE 32. 1. Hemipeplus africanus Grouvelle, 1915. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. FIGURE 32.2. Hemipeplus africanus Grouvelle, 1915. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFFC6E482A8DFF006B412F50.taxon	description	FIGURE 33. 2. Hemipeplus alluaudi Grouvelle, 1923 (Malawi). A. Dorsal habitus; B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view; F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFFC6E482A8DFF006B412F50.taxon	materials_examined	Type Material. Lectotype, designated by Pollock (1997), sex unknown, only images sent from MNHN, labeled: ‘ Cote d’Afrique or. angl. RIVIERE RAMISI ALLUAUD ET JEANNEL NOV 1911 St. 8 / [pink label] Type / Hemipeplus alluaudi Grouv. / MUSEUM PARIS 1917 Coll. GROUVELLE / LECTOTYPE Hemipeplus alluaudi Grouvelle design D. A. Pollock 1996 / [with a barcode] MNHN, Paris EC 26371 ’, in MNHN (Grouvelle Collection). Paralectotype, sex unknown, only images sent from MNHN, labeled: ‘ Cote d’Afrique or. angl. RIVIERE RAMISI ALLUAUD ET JEANNEL NOV 1911 St. 8 / Hemipeplus alluaudi Grouv. / MUSEUM PARIS Coll. A. GROUVELLE 1917 / [red label] Type / [red label] PARALECTOTYPE Hemipeplus alluaudi Grouv. Designated by D. A. Pollock 1997 / [with a barcode] MNHN, Paris EC 26372 ’, in MNHN (Grouvelle Collection). Other Material Examined. KENYA. Kwale District. Shimba Hills Res. Forest 28. viii. 1986, B. D. Valentine family coll., Valentine coll’n Rec. 2016, (FSCA, 1 ♂); same locality, 26. viii. 1986, (FSCA, 1 ♂ (Fig. 33.1), 3 ♀). REPUBLIC OF MALAWI. Mulanje District. Likhubula Forest Lodge 24 – 25. ii. 2009 700 m, J. Heppner & R. Murphy leg., host-Leguminosae: Brachystegia, (FSCA, 1 ♂) (Fig. 33.2).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFFC6E482A8DFF006B412F50.taxon	diagnosis	Diagnosis. Hemipeplus alluaudi can be distinguished from the similar-appearing H. africanus (Figs 32.1, 32.2) by its posterolateral pronotal margins not subparallel and uniformly paler color. Hemipeplus alluaudi has the following diagnostic features: temples short and angulate to somewhat rounded, slightly larger in some specimens; scape submoniliform, somewhat more elongate than in H. africanus (Figs 32.1, 32.2); eyes large (longer than scape + pedicel), wide and convex; pronotum wide to relatively narrow, subcordiform; anterior half wider than posterior half with deep to shallow anterior emargination in center; anterolateral angles moderately to strongly produced, broadly rounded; pronotal pad present, more distinct than in H. africanus; hind angles obtuse, rounded to somewhat angled; posterolateral margins of pronotum not straight or subparallel; pronotal lobe indistinct and without median notch; scutellar shield transverse, elliptical to subhexagonal, less than twice as wide as long; body long; elytra slightly convex but distinctly flatter than in H. africanus; color uniformly brown; distribution: Kenya, Malawi. Male Genitalia (Figs 33. 1 D – F, 33. 2 D – F). (Tegmen length = 0.33 – 0.42 mm (n = 3 )) Lobe of basale very short, and wide, shorter than in H. africanus; shoulders slanted and obtusely angled; apicale and basale with somewhat clear suture; basale subequal to apicale (not including basal lobe); parameres long and almost straight, apices curved inward; apices thin and sharp; penis wide and longer than tegmen, apically pointed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFFC6E482A8DFF006B412F50.taxon	description	Variation. The specimen of H. alluaudi from Malawi is larger than those from Kenya.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFFC6E482A8DFF006B412F50.taxon	discussion	Remarks. Hemipeplus alluaudi closely resembles the lighter form of H. africanus, and even male genitalia are virtually inseparable except for the shorter, wider lobe of the basale that appears consistent among the specimens. Externally, it can be distinguished from H. africanus using the keys provided in the diagnosis section. Geographical Distribution (Fig. 63). Kenya (south), Malawi (new country record). Natural History. A label on one specimen from Malawi in FSCA reads that the host is Brachystegia Benth. (Leguminosae). If so, this is the first record of a dicot plant being “ host ” to Hemipeplinae, although they have been collected by sweeping dicots such as Salicornia before (Pollock 1999). Apart from that, nothing is known regarding the natural history of H. alluaudi.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF06E4C2A8DFCC06AA62B60.taxon	description	(Figs 34, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF06E4C2A8DFCC06AA62B60.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ SOLOMON IS. BOUGAINVILLE (S.) Kokure, 690 m. June 9, 1956 / E. J. Ford. Jr. Collector / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus antennatus KC & Pollock’, in BPBM. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the species’ distinctive subcylindrical apical antennomeres, differing from the semi-triangular antennomeres characteristic of its congeners.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF06E4C2A8DFCC06AA62B60.taxon	diagnosis	Diagnosis. Hemipeplus antennatus sp. nov. is the only known species in Hemipeplinae with subcylindrical apical antennomeres. Hemipeplus antennatus sp. nov. has the following diagnostic features: temples absent; antennomeres V to basal half of XII subcylindrical; eyes large (almost as long as antennomeres I + II + III), wide and strongly convex; pronotum subcordiform and rufopiceous; anterior emargination moderate; anterior half wider than posterior half; pronotal pad distinct; pronotal angles broadly rounded; pronotal lobe indistinct without median notch; body elongate; elytra more or less flat; color piceous with shades of lighter brown; distribution: Solomon Islands (Bougainville).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF06E4C2A8DFCC06AA62B60.taxon	description	Description. Measurements (in mm) — GHW: 0.68; HL: 0.65; AL: 1.13; GPW: 0.70; PL: 0.61; GEW: 0.76; EL: 3.16; TL: 4.42. Head (Fig. 34 B) wide, (GHW / HL: 1.05); eyes large (almost as long as antennomeres I + II + III), wide and strongly convex; temples absent; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape submoniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV short and moniliform; V – X subcylindrical, each larger than the next; XI subcylindrical with apex narrowed after median notch; lateral mandibular tooth absent in the specimen. Pronotum (Fig. 34 B) relatively wide (GPW / PL: 1.15); anterior margin with moderate emargination; anterolateral angles broadly rounded and moderately produced; lateral margins sinuate; posterior half much more contracted than apical half; posterolateral angles obtuse and rounded; pronotal lobe truncate without median notch; pronotal pad (Fig. 34 C) distinctly present; pronotal pits shallow with moderate grooves. Scutellar shield convex, transverse, not twice as wide as long. Elytra (Fig. 34 A) elongate (EL / GEW: 4.16) and more or less flat; vestiture short and moderate; apical patch cannot be viewed because of the condition of the specimen. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges appear well separated in the specimen but cannot be sure because of the condition of the specimen. Color dull, head and pronotum rufopiceous; labrum, clypeus lighter; elytra piceous with shades of lighter brown; ventrally head and prothorax darker, palpi lighter. FIGURE 34. Hemipeplus antennatus sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia tegmen left side. Male Genitalia (Figs 34 D). (Tegmen length not available (lost before measurement was done )) Lobe of basale moderately long and slender; shoulders sharply acute, pointed upward; apicale and basale with distinct suture; parameres short and arcuate at the apices; apices of parameres narrower than the bases with sharp thin tips; length of penis not known. Geographical Distribution (Fig. 64). Solomon Islands (Bougainville).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF66E4E2A8DFE056D532AD8.taxon	description	(Figs 35, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF66E4E2A8DFE056D532AD8.taxon	materials_examined	Type Material. Holotype, male, right specimen of the two specimens on same pin, labeled: ‘ F. W. Lake, 10 ml. N. of Rocky R., Via Coen, N. Qld. 17. xii. 1964. G. Monteith / [light green label] U. Q. I. C. Loan 797 / [dark green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus monteithi right specimen KC & Pollock’, in UQIC. Twenty-one paratypes: 1 specimen, male, labeled same as holotype, on the left on the same pin, in UQIC; 1 specimen, male, labeled: ‘ 11.45 S 142.35 E QLD Heathlands nr. gravel pit 9 Dec. 1992 W. Dressler, P. Zborowski, at light’, in ANIC; 2 specimens, sex unknown, labeled: ‘ 11.43 S 142.41 E 12 km EbyN Heathlands, QLD 15 - 26 Jan. 1992 I. D. Naumann, T. A. Weir rainforest’, in ANIC; 1 specimen, sex unknown, labeled: ‘ 11.41 S 142.42 E 15 km NEbyN Heathlands QLD 15 - 26 Jan. 1992 I. Naumann, T. A. Weir / rainforest and rainforest margin’ in ANIC; 1 specimen, sex unknown, labeled: ‘ 11.41 S 142.42 E 14 km ENE Heathlands, 7 Dec. 1992 W. Dressier P. Zborowski, beating rainforest / HEMIPEPLUS AUSTRALICUS ARROW det. JF Lawrence’, in ANIC; 1 specimen, sex unknown, labeled: ‘ 12.44 S 143.14 E, QLD 3 km ENE Mt. Tozer, 28 June — 4 July 1986 J. C. Cardale ex ethanol’, in ANIC; 12 specimens, sex unknown, labeled: ‘ F. W. Lake, 10 ml. N. of Rocky R., Via Coen N. Qld. 17. xii. 1964. G. Monteith’, in UQIC; 2 specimens, sex unknown, labeled: ‘ F. W. Lake, 10 ml. N. of Rocky R., Via Coen, N. Qld. 17. xii. 1964. G. Monteith / EX BASES OF LOMANDRA / On loan from Ent. Dept. U. of Qld. ’, in ANIC. Derivation of Specific Epithet. The specific name (noun in the genitive case) is given in allusion to Geoff Monteith who collected the holotype in 1964.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF66E4E2A8DFE056D532AD8.taxon	diagnosis	Diagnosis. Hemipeplus monteithi sp. nov. resembles H. australicus (Figs 39.1, 39.2) with which it might be sympatric, but it is wider and shorter in appearance. Hemipeplus monteithi sp. nov. has the following diagnostic features: temples absent; scape moniliform; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum subcordiform, wide and flat; anterior emargination broad and shallow; anterior half much wider than posterior half; pronotal pad distinct, wide with profuse setae; pronotal angles rounded; pronotal lobe indistinct without median notch; body short and elongate; elytra more or less flat; color uniformly pale and shiny; distribution: Australia (Queensland).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF66E4E2A8DFE056D532AD8.taxon	description	Description. Measurements (in mm) — GHW: 0.50 – 0.55; HL: 0.45 – 0.52; AL: 0.80 – 0.88; GPW: 0.60 – 0.67; PL: 0.45 – 0.50; GEW: 0.65 – 0.75, EL: 2.23 – 2.43; TL: 3.13 – 3.45. Head (Fig. 35 B) wide, (GHW / HL: 1.06 – 1.11); eyes large (longer than scape + pedicel), moderately wide and moderately convex; temples absent; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape short and moniliform; pedicel small moniliform; antennomere III more elongate and apically dilated; IV short and moniliform; V slightly larger, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after notch; lateral mandibular tooth very small. Pronotum (Fig. 35 B) wide (GPW / PL: 1.33 – 1.34); anterior margin with broad shallow emargination; anterolateral angles rounded, produced; lateral margins weakly sinuate; posterior half much more contracted than anterior half; posterolateral angles obtusely rounded; pronotal lobe truncate without median notch; pronotal pad (Fig. 35 C) broad and distinct; pronotal pits moderately developed with moderate grooves. Scutellar shield subhexagonal, transverse, nearly twice as wide as long. Elytra (Fig. 35 A) short (EL / GEW: 3.24 – 3.43) and flat; vestiture short and dense; apical patch long and only slightly darker than elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges widely separated; mesosternal process narrow and linear. Color uniformly light brown / pale, shiny. Male Genitalia (Figs 35 D – F). (Tegmen length = 0.26 mm (n = 1 )) Lobe of basale moderately long, slender with blunt apex; shoulders with small degree of slant, rectangular; apicale and basale with distinct suture; parameres arcuate with apices thick and blunt, as if cut; penis much longer than tegmen. Note: the parameres of H. monteithi sp. nov. are similar to those of H. australicus but less curved. It should be noted that all Australian species have very similar male genitalia only varying in the degree of curvedness of the parameres, suggesting possible common ancestry. FIGURE 35. Hemipeplus monteithi sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar) F. Tegmen lateral view with one paramere removed. Geographical Distribution (Fig. 64). Australia (Queensland). Natural History. According to specimen label data, H. monteithi sp. nov. is associated with Lomandra Labill. (Asparagaceae) and adults have been collected from light and beating rainforest.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF46E4F2A8DFEDD6B8B2DA6.taxon	description	(Figs 36, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF46E4F2A8DFEDD6B8B2DA6.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ NEW GUINEA (Neth.) Hollandia 100 m., Aug. 24. 1955 / Light Trap J. L. Gressitt / [blue label] ♀ / [red label] HOLOTYPE ♀ Hemipeplus gressitti KC & Pollock’, in BPBM. One paratype, female, labeled: ‘ New Guinea: Voqelkop Manokwari NE of town 10 - 25 m 10. vi. 1979 / J. L. Gressitt Collector BISHOP MUSEUM / Palms Sago / V. large eyes Det. D. A. Pollock’, in BPBM. Derivation of Specific Epithet. The specific name (noun in the genitive case) is given in allusion to Judson Linsley Gressitt who collected the holotype in 1955.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF46E4F2A8DFEDD6B8B2DA6.taxon	diagnosis	Diagnosis. Hemipeplus gressitti sp. nov. can be distinguished by its very large eyes and lack of temples. Hemipeplus gressitti sp. nov. has the following diagnostic features: temples absent; scape elongate, apically dilated; largest eyes among all Hemipeplinae species (longer than antennomeres I + II + III), wide and convex; pronotum subcordiform, relatively long; anterior emargination almost absent; anterior half much wider than posterior half; pronotal pad distinct; pronotal angles rounded; pronotal lobe truncate without visible median notch; body long; elytra slightly convex; color brown, dull, head and pronotum slightly darker; rufopiceous head and pronotum in paratype; distribution: Indonesia (Hollandia, Manokwari).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF46E4F2A8DFEDD6B8B2DA6.taxon	description	Description. Measurements (in mm) — GHW: 0.55 – 0.68; HL: 0.50 – 0.63; AL: 0.88 – 0.92; GPW: 0.55 – 0.70; PL: 0.50 – 0.60; GEW: 0.60 – 0.95, EL: 2.70 – 3.16; TL: 3.70 – 4.39. Head (Fig. 36 B) wide, (GHW / HL: 1.08 – 1.10); largest eyes among all Hemipeplinae species (longer than antennomeres I + II + III), wide and convex; temples absent; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape elongate, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV short and moniliform, apically dilated; V broader, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after median notch; lateral mandibular tooth absent. Pronotum (Fig. 36 B) wide (GPW / PL: 1.10 – 1.17); anterior margin without prominent emargination; anterolateral angles rounded, produced; posterior half much more contracted than anterior half; posterolateral margins subparallel; posterolateral angles obtusely rounded; pronotal lobe truncate without visible median notch; pronotal pad (Fig. 36 C) distinct; pronotal pits shallow with faint grooves. Scutellar shield elliptical, transverse, not twice as wide as long. Elytra (Fig. 36 A) long (EL / GEW: 3.33 – 4.50) and slightly convex; vestiture short and moderate; apical patch long but faint. Ventral surface. Prosternal process short and pointed, not extending posterior of procoxae; hypomeral edges almost fused, very narrowly separated; mesosternal process narrow and linear. Color dull, brown, head and pronotum slightly darker (rufopiceous in paratype) than elytra; ventrally same as above, palpi and tarsi lighter in holotype; uniformly rufous in paratype with edges of pro- and mesosternum darker. Male Genitalia. Even though a male specimen was not available, we hypothesize that the male genitalia are very similar to those of species such as H. australicus, H. enarotaliensis sp. nov., and H. neoguineensis sp. nov. based on the external morphological characters.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFF46E4F2A8DFEDD6B8B2DA6.taxon	discussion	Remarks. Hemipeplus gressitti sp. nov. has the largest eyes relative to its antennomeres in the entire subfamily. Geographical Distribution (Fig. 64). Indonesia (Hollandia, Manokwari). Natural History. According to specimen label data, H. gressitti sp. nov. has been collected from Sago Palm and light trap (s). FIGURE 36. Hemipeplus gressitti sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFEA6E522A8DFF006B8B2DF3.taxon	description	(Figs 37, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFEA6E522A8DFF006B8B2DF3.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ NEW GUINEA: NE LAE AREA: Lae-Bubia 6 – 20 m. VII- 21 - 1959 / K. beccarii / J. L. Gressitt Collector. / No temples Det. D. A. Pollock 19 / [light green label] BPBM / [dark green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus neoguineensis KC & Pollock’, in BPBM. Ten paratypes; 2 specimens, sex unknown, labeled: ‘ NEW GUINEA: SE Murua River (S. side), 2 m 17. XII. 1964 / J. Sedlacek Collector BISHOP MUS. ’, in BPBM; 1 specimen, female, labeled: ‘ NEW GUINEA: PAPUA Brown River 5. XI. 1960 / Calamus / J. L. Gressitt Collector’, in BPBM; 1 specimen, sex unknown, labeled: ‘ N. GUINEA: NE Bulolo 760 m, 18.6. [illegible year] / J. Sedlacek Collector BISHOP MUS. / Hemipeplus sp. det. A. S. Slipinski 96 ’, in BPBM; 1 specimen, male, labeled: ‘ NEW GUINEA (SE) Cape Rodney 3. XI. 1960. / J. L. Gressitt Collector / Heterospathe’, in BPBM; 1 specimen, female, labeled: ‘ NEW GUINEA: PAPUA Keparra-Sengi, Nr. Kokoda 500 m. III- 26 - 1956 / J. L. Gressitt Collector / Sago palm’, in BPBM; 2 specimens (1 inside a capsule), sex unknown, labeled: ‘ NEW GUINEA: PAPUA Daradae Pl’n. 500 m, 80 km N. to Port Moresby, IX- 4 - 1959 / T. C. Maa Collector BISHOP / Sweeping’, in BPBM; 1 specimen, sex unknown, labeled: ‘ NEW GUINEA: SE Moresby, Brown R. 10 m. 16. XII. 1964 / J. Sedlacek Collector BISHOP MUS. ’, in BPBM; 1 specimen, sex unknown, labeled: ‘ NEW GUINEA: NETH. Waris, S. of Hollandia, 450 - 500 m VIII- 8 - 15 - 1959 / T. C. Maa Collector’, in BPBM. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to this species’ type locality, i. e., New Guinea.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFEA6E522A8DFF006B8B2DF3.taxon	diagnosis	Diagnosis. Hemipeplus neoguineensis sp. nov. can be distinguished from the similar-appearing H. australicus (Figs 39.1, 39.2) by its larger eyes, browner color, and less setose pronotal pad. Hemipeplus neoguineensis sp. nov. has the following diagnostic features: temples absent; scape submoniliform, apically dilated, externally produced on one side; eyes large (as long as scape + pedicel + antennomere III), moderately wide and moderately convex; pronotum subcordiform; anterior emargination almost absent; anterior half much wider than posterior half; pronotal pad distinct; pronotal angles rounded; pronotal lobe truncate without median notch; body long; elytra slightly convex; color uniformly rufotestaceous, not shiny; distribution: New Guinea.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFEA6E522A8DFF006B8B2DF3.taxon	description	Description. Measurements (in mm) — GHW: 0.55; HL: 0.43 – 0.45; AL: 0.80 – 0.88; GPW: 0.58; PL: 0.48; GEW: 0.65 – 0.68; EL: 2.23 – 2.38; TL: 3.14 – 3.31. Head (Fig. 37 B) wide (GHW / HL: 1.22 – 1.28); eyes large (as long as scape + pedicel + antennomere III), moderately wide and moderately convex; temples absent; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape submoniliform, apically dilated, externally produced on one side; pedicel small moniliform; antennomere III slightly more elongate and apically dilated; IV short and moniliform, apically dilated; V broader, apically dilated; VI same as V but larger; VII moniliform, apically dilated; VIII – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 37 B) wide (GPW / PL: 1.21); anterior margin without prominent emargination; anterolateral angles rounded, produced; lateral margins moderately sinuate; posterior half much more contracted than anterior half; posterolateral angles obtusely rounded; pronotal lobe truncate without visible median notch; pronotal pad (Fig. 37 C) distinct; pronotal pits shallow without well-developed grooves. Scutellar shield subhexagonal, transverse, about twice as wide as long. Elytra (Fig. 37 A) long (EL / GEW: 3.43 – 3.50) and slightly convex; vestiture short and dense; apical patch long and faint. Ventral surface. Prosternal process short, not extending to posterior of procoxae; hypomeral edges very narrowly separated to fused; mesosternal process narrow and linear. Color uniformly rufotestaceous, dull. Male Genitalia (Figs 37 D – F). (Tegmen length = 0.28 mm (n = 1 )) Lobe of basale short, slender with rounded apex; shoulders obtusely angled to slightly smooth with moderate slant; apicale and basale with distinct suture; parameres arcuate with apices thick and blunt, as if cut; penis slightly longer than tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFEA6E522A8DFF006B8B2DF3.taxon	discussion	Remarks. Although the male genitalia of H. neoguineensis sp. nov. is similar to that of H. australicus, there are significant morphological differences, such as those provided in the diagnosis section, which are consistent throughout the New Guinean population. Geographical Distribution (Fig. 64). New Guinea. Natural History. According to specimen label data, H. neoguineensis sp. nov. is associated with Calamus L. (Acoraceae), Cycas revoluta Thunb. or Sago Palm (Cycadaceae), K. beccarii [Coelogyne beccarii? Rchb. f. (Orchidaceae)], and Heterospathe Scheff. (Arecaceae). FIGURE 37. Hemipeplus neoguineensis sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. FIGURE 38. Hemipeplus manusicus sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE96E532A8DFF006C2C2F75.taxon	description	(Figs 38, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE96E532A8DFF006C2C2F75.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ BISMARCK ARCH: MANUS I., Lorengau nr. sea level Dec. 15 - 29, 1959 / SAGO PALM / T. C. Maa Collector BISHOP / [blue label] ♀ / [red label] HOLOTYPE ♀ Hemipeplus manusicus KC & Pollock’, in BPBM. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to this species’ type locality, i. e., Manus Island in Papua New Guinea.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE96E532A8DFF006C2C2F75.taxon	diagnosis	Diagnosis. Hemipeplus manusicus sp. nov. can be distinguished from the similar-appearing H. gressitti sp. nov. (Fig. 36) by its relatively smaller eyes and darker color. Hemipeplus manusicus sp. nov. has the following diagnostic features: temples absent; scape submoniliform, slightly elongate, apically dilated; eyes large (longer than scape + pedicel), wide and convex; pronotum subcordiform; anterior emargination weak; anterior half much wider than posterior half; pronotal pad distinct; pronotal angles rounded; pronotal lobe indistinct with faint median notch; body long; elytra slightly convex; color dark brown, dull, head and pronotum rufopiceous, darker than the elytra; distribution: Papua New Guinea (Manus Island).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE96E532A8DFF006C2C2F75.taxon	description	Description. Measurements (in mm) — GHW: 0.60; HL: 0.60; AL: 0.87; GPW: 0.68; PL: 0.58; GEW: 0.80, EL: 2.95; TL: 4.13. Head (Fig. 38 B) about as long as wide (GHW / HL: 1.00); eyes large (longer than scape + pedicel), wide and convex; temples absent; occiput with strongly raised areas; antennae shorter than head and pronotum length combined; antennal scape submoniliform, slightly elongate, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV short and moniliform, apically dilated; V broader, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed; lateral mandibular tooth absent. Pronotum (Fig. 38 B) wide (GPW / PL: 1.17); anterior margin with slight emargination; anterolateral angles rounded, produced; posterior half much more contracted than anterior half; posterolateral margins subparallel; posterolateral angles obtusely rounded; pronotal lobe truncate with a slight median notch; pronotal pad (Fig. 38 C) distinct; pronotal pits shallow with faint grooves. Scutellar shield subpentagonal, transverse, less than twice as wide as long. Elytra (Fig. 38 A) long (EL / GEW: 3.69) and slightly convex; vestiture short and dense; apical patch long and much darker than elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges narrowly separated, very close to each other; mesosternal process narrow and linear. Color dull, dark brown with head and pronotum darker than elytra; ventrally about uniformly rufous; inner edges of pro- and mesosternum (including hypomeral edges) darker; tarsi, palpi, glossal sclerite, and galea paler. Geographical Distribution (Fig. 64). Papua New Guinea (Manus Island). Natural History. According to specimen label data, H. manusicus sp. nov. is associated with Sago Palm.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE96E562A8DFA306CDE2C0D.taxon	materials_examined	Type Material. Lectotype (here designated; Article 74 (ICZN, 1999 )); 1 specimen (left), sex unknown, pinned along with another specimen on same card, labeled: ‘ [round label with red margin] Type / [underlined in pink] Port Darwin. [/ underlined in pink] 92 — 2. / [label upside down] 4713 / Hemipeplus australicus type arrow / [red label] LECTOTYPE left spec. Hemipeplus australicus Arrow designated by D. A. Pollock 1997 / [with a barcode] NHMUK 014663483 / [with a barcode] NHMUK 014663484 ’, in BMNH (Fig. 39.1); Paralectotype: 1 specimen (right on same card as lectotype, ventral side up), sex unknown, labeled same as lectotype, in BMNH. FIGURE 39.1. Hemipeplus australicus Arrow, 1930 (Lectotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax. FIGURE 39.2. Hemipeplus australicus Arrow, 1930. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar). Other Material Examined. AUSTRALIA. Northern Territory. Adelaide River 91 — 49, (BMNH, 3); Port Darwin, NW Australia, J. J. Walker, G. C. Champion leg., B. M. 1927 – 409, (BMNH, 1 ♀, 1 ♂ (Fig. 39.2), 1 sex unknown); Port Keats, N. T. 14.06 S, 129.33 E, 19. viii. 1968, M. Mendum leg., (ANIC, 1); Umbakumba, Little Lagoon, Groote Eylandt, 17. vii. 1952, J. L. Wassell leg., at light, (UQIC, 1). Western Australia. W. Austl. N side Carson, R. on Gibb Rd., 27. x. 1976, FMHD # 76 – 5016, at light nr pool, J. B. Kethley leg., (FMNH, 3 ♀); same locality and date, FMHD # 76 – 5044, at light, J. B. Kethley leg., (FMNH, 1 ♂, 2 ♀). Queensland. Mogill, near Brisbane, 27. ix. 1958 x, J. L. Gressitt leg., Hemipeplus sp. Det. J. F. Lawrence, (BPBM, 1 ♀); Fossill Site “ B ” Mt. Crosby, S. E. QU., 16. x. 1963, E. Exley leg., (UQIC, 1 ♀); Millstream Falls Via Ravenshoe, 24. xii. 1964. N. Qld. H. A. Rose leg., (UQIC, 1); Iron Range, Cape York Pen. N. Qld. 16 – 23. xi. 1965, G. Monteith leg., (UQIC, 1); 15.41 ° S 145.12 ° E Annan R. 3 km WbyS Black Mt. 27. ix. 1980 Q, J. C. Cardale leg., ex ethanol, (ANIC, 1); Babinda NQ 11 / 50 93, (ANIC, 1 ♀); Tolga, 20. x. 1985, J. D. Brown leg., ex collection A Walford-Huggins, E. Gowing-Scopes collection BMNH (E) 2005 – 4, (BMNH, 7). Unknown locality. (UQIC, 6).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE96E562A8DFA306CDE2C0D.taxon	diagnosis	Diagnosis. Hemipeplus australicus can be distinguished by its lack of temples, wide pronotal pad, elongate body, and shiny, pale color. Hemipeplus australicus has the following diagnostic features: temples absent; scape moniliform, apically dilated, externally produced on one side; eyes large (longer than scape + pedicel), wide and convex; pronotum subcordiform; anterior emargination almost absent; anterior half much wider than posterior half; pronotal pad broad and distinct; pronotal angles rounded; pronotal lobe truncate without visible median notch; body long; elytra slightly convex; color uniformly light brown / pale, shiny; distribution: Australia (Northern Territory, Queensland, Western Australia). Re-description. Measurements (in mm) — GHW: 0.58 – 0.59; HL: 0.48 – 0.50; AL: 0.80 – 0.92; GPW: 0.61 – 0.63; PL: 0.48 – 0.55; GEW: 0.73 – 0.78; EL: 2.70 – 2.85; TL: 3.66 – 3.90. Head (Figs 39.1 B, 39.2 B) wide (GHW / HL: 1.18 – 1.21); eyes large (longer than scape + pedicel), wide and convex; temples absent; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape moniliform, apically dilated, externally produced on one side; pedicel small moniliform; antennomere III more elongate and apically dilated; IV short and moniliform, apically dilated; V broader, apically dilated; VI same as V but larger; VII – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed; lateral mandibular tooth very short. Pronotum (Figs 39.1 B, 39.2 B) wide (GPW / PL: 1.15 – 1.27); anterior margin without prominent emargination, almost straight; anterolateral angles rounded, produced; lateral margins sinuate; posterior half much more contracted than anterior half; posterolateral angles obtusely rounded; pronotal lobe truncate without visible median notch; pronotal pad (Figs 39.1 C, 39.2 C) very wide and distinct, densely setose; pronotal pits shallow without well-developed grooves. Scutellar shield elliptical to subhexagonal, transverse, wider than long. Elytra (Figs 39.1 A, 39.2 A) long (EL / GEW: 3.65 – 3.70), slightly convex; vestiture long and dense; apical patch short and faint. Ventral surface. Prosternal process short, not extending to posterior of procoxae; hypomeral edges fused in most specimens; mesosternal process narrow and linear. Color feebly shiny, uniformly light brown / pale. Male Genitalia (Figs 39.2 D – F). (Tegmen length = 0.25 mm (n = 1 )) Lobe of basale moderately long, slender with blunt apex; shoulders obtusely angled to smooth with moderate slant; apicale and basale with distinct suture; parameres arcuate with apices thick and blunt, as if cut; penis much longer than tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE96E562A8DFA306CDE2C0D.taxon	distribution	Distribution (Fig. 64). Australia (Northern Territory, Queensland, Western Australia). Natural History. According to specimen label data, adults of H. australicus have been collected from light traps.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFEC6E5A2A8DF9E96DED2D74.taxon	description	(Figs 40.1, 40.2, 40.3, 65)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFEC6E5A2A8DF9E96DED2D74.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ Philippinen Mindoro, Mangarin [underside of label] Böttcher XI. 1917 / Zool. Mus. Berlin / [green label] ♂ / [blank red label] / [red label] HOLOTYPE ♂ Hemipeplus harkoneni KC & Pollock’, ZMHB. Twenty-four paratypes: 1 specimen, male, labeled same as holotype, in ZMHB; 2 specimens, sex unknown, labeled same as holotype, in ZMHB; 2 specimens, 1 male, 1 female, labeled: ‘ Philippinen, Mangarin. XI. 1917 leg. Böttcher’, in ZMHB; 1 specimen, male, labeled: ‘ Tarragona, Leyte I., Philippine Islds. VI: 12: 1945 / Col. & Pres. by C. L. Remington / at light’, in FMNH; 1 specimen, female, labeled: ‘ Los Banos Luzon, P. I. XI. 22 - 26.1947 / W. H. Lange Collector / NMNH’, in NMNH; 1 specimen, female, labeled: ‘ Luzon: Laguna: Mt. Makiling, 250 m. XI- 23 - 53 / J. L. Gressitt Collector / BPBM’, in BPBM; 2 specimens, 1 female, 1 sex unknown, labeled: ‘ Ligao, IX. 78. coconut / R. Mosquite coll. / Philippines Luzon I. / C. I. E. A 11473 / Hemipeplus sp. (unnamed in B. M.) det. R. B. Madge, 1979 [in one specimen only] / BMNH’, in BMNH; 1 specimen, male, labeled: ‘ San Pablo. Luzon, P. I., XI. 23.1947 / W. H. Lange Collector / NMNH’, in NMNH; 2 specimens, sex unknown, labeled: ‘ Mt. Makiling, Luzon, Baker / NMNH’, in NMNH; 1 specimen, sex unknown, labeled: ‘ San Jose, Leyte, P. I. I: 13: 1949 / Col. & pres. by Eugene Ray / at light / FMNH’, in FMNH (Fig. 40.2); 2 specimens, sex unknown, labeled: ‘ San Jose, Leyte, P. I. III- 1 - 4 - 1945 / Col. & pres. by Eugene Ray / at light / FMNH’, in FMNH; 2 specimens, sex unknown, labeled: ‘ San Jose, Leyte, P. I. II. 25.45 / Col. & pres. by Eugene Ray / at light / Hemipeplus det. Newton 1993 [in one specimen only] / FMNH’, in FMNH; 1 specimen, sex unknown, labeled: ‘ Tarragona, Leyte I., Philippine Islds. VI: 17: 1945 / Col. & pres. by C. L. Remington / at light / FMNH’, in FMNH; 1 specimen, sex unknown, labeled: ‘ Tarragona, Leyte I., Philippine Islds., V: 28: 1945. / Col. & pres. by C. L. Remington / at light / FMNH’, in FMNH; 1 specimen, sex unknown, labeled: ‘ Tarragona, Leyte I., Philippine Islds., VI: 24: 1945. / Col. & pres. by C. L. Remington / at light / FMNH’, in FMNH; 2 specimens, 1 male, 1 sex unknown (on same pin), labeled: ‘ [underlined in yellow] SARAWAK: / Biawak on nipah palm 17 - vi- 1968 Coll: S. Juses / Pres by Com Inst Ent BM 1971 - 1 / 9104 ’, in BMNH (Fig. 40.3); 1 specimen, female, labeled: ‘ [underlined in yellow] SARAWAK: / Biawak on nipah 18 - vi- 68 coll. Vincent A. / Pres by Com Inst Ent BM 1971 - 1 / 8985 ’, in BMNH. Derivation of Specific Epithet. This species (noun in the genitive case) is named in allusion to the first author’s late friend from Finland, Reijo Härkönen, who helped him in his research during its early phases.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFEC6E5A2A8DF9E96DED2D74.taxon	diagnosis	Diagnosis. Hemipeplus harkoneni sp. nov. can be distinguished by its large eyes, lack of temples, and pronotal pad narrower than in H. australicus (Figs 39.1 C, 39.2 C). Hemipeplus harkoneni sp. nov. has the following diagnostic features: temples absent; scape short, moniliform; eyes large (longer than scape + pedicel), wide and convex; pronotum subcordiform, wide and flat to more elongate with convex disc; anterior pronotal emargination clear; anterior half wider than posterior half; pronotal pad moderate; pronotal angles rounded; pronotal lobe truncate / indistinct without a clear median notch; body relatively more wide to elongate; elytra more or less flat; color uniformly pale to deeper brown; distribution: Malaysia (Sarawak), Philippines (Bangui, Leyte, Ligao, Luzon, Mangarin, San Jose, San Pablo).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFEC6E5A2A8DF9E96DED2D74.taxon	description	Description. Measurements (in mm) — GHW: 0.62 – 0.70; HL: 0.58 – 0.65; AL: 0.95 – 1.06; GPW: 0.66 – 0.73; PL: 0.54 – 0.66; GEW: 0.83 – 0.93; EL: 2.68 – 3.38; TL: 3.80 – 4.69. Head (Figs 40.1 B, 40.2 B, 40.3 B) wide (GHW / HL: 1.07 – 1.08); eyes large (longer than scape + pedicel), wide and convex; temples absent; occiput distinctly raised; antennae shorter than head and pronotum length combined; antennal scape short and moniliform; pedicel small moniliform; antennomere III more elongate and apically dilated; IV short and moniliform; V short and subcylindrical; VI – X subtriangular or bowl shaped, each larger than the next; XI subpyriform with apex narrowed after median notch; lateral mandibular tooth very small. Pronotum (Figs 40.1 B, 40.2 B, 40.3 B) wide (GPW / PL: 1.11 – 1.22), wider in some specimens than others; anterior margin slightly (Fig. 40.3 B) to more deeply (Figs 40.1 B, 40.2 B) emarginate; anterolateral angles rounded and produced; lateral margins sinuate; posterior half much more contracted; posterolateral angles rounded and obtuse; posterior lobe truncate without significant median notch; pronotal pad (Figs 40.1 C, 40.2 C, 40.3 C) present, more prominent in some specimens than others; pronotal pits shallow to deep, grooves present when deep. Scutellar shield elliptical to subhexagonal, transverse, not twice as wide as long. Elytra (Figs 40.1 A, 40.2 A, 40.3 A) elongate (EL / GEW: 3.29 – 3.63), more or less flat; vestiture short and dense; apical patch long, rufopiceous and prominent to faint light brown. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges fused (in holotype) to very thinly separated; mesosternal process narrow and linear. Color varies from uniformly testaceous to rufotestaceous with rufous head and pronotum. Male Genitalia (Figs 40. 1 D – F, 40. 3 D – F). (Tegmen length = 0.35 – 0.36 mm (n = 2 )) Lobe of basale long and slender with rounded apex; shoulders pointed upward, as in H. joerebeccae sp. nov. and H. antennatus sp. nov.; apicale and basale with distinct suture; parameres arcuate at apices with tips blunt; penis longer than tegmen. Variation. Several specimens from Luzon are more elongate with elytra dark, while several specimens from Ligao and Leyte are wider and light brown. Geographical Distribution (Fig. 65). Malaysia (Sarawak), Philippines (Bangui, Leyte, Ligao, Luzon, Mangarin, San Jose, San Pablo). FIGURE 40.1. Hemipeplus harkoneni sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. Natural History. According to specimen label data, H. harkoneni sp. nov. is associated with Cocos nuciferae L. or coconut (Arecaceae) and Nypa fruticans Wurmb or Nipah Palm (Arecaceae); adults have also been collected from light traps. FIGURE 40.2. Hemipeplus harkoneni sp. nov. (male). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax. FIGURE 40. 3. Hemipeplus harkoneni sp. nov. (from Malaysia). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE16E5C2A8DFF006AE62EE4.taxon	description	(Figs 41, 65) FIGURE 41. Hemipeplus bucculentus sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE16E5C2A8DFF006AE62EE4.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ LAOS: Vientiane Prov. Ban Van Eue, 15. V. 1966 / Native Collector RONDON- - BISHOP MUS. COLLECTION / [green label] BPBM / [blue label] ♀ / [red label] HOLOTYPE ♀ Hemipeplus bucculentus KC & Pollock’, in BPBM. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the species’ notably wide temples, resembling fat cheeks or a large mouth.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE16E5C2A8DFF006AE62EE4.taxon	diagnosis	Diagnosis. Hemipeplus bucculentus sp. nov. can be distinguished by its prominent round temples, pronotum produced below anterior axis, and dark color. Hemipeplus bucculentus sp. nov. has the following diagnostic features: temples prominent; scape moniliform, apically dilated, externally produced on one side; eyes small (shorter than scape + pedicel), shallow, and moderately convex; pronotum subcordiform with coarse texture; anterior emargination almost absent; anterior half wider than posterior half slightly below the anterior axis; pronotal pad absent; pronotal angles more or less rounded; pronotal lobe truncate with a hint of median notch; body long; elytra slightly convex; color dull, rufopiceous with darker pronotum; distribution: Laos (Vientiane Province).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE16E5C2A8DFF006AE62EE4.taxon	description	Description. Measurements (in mm) — GHW: 0.56; HL: 0.50; AL: 0.92; GPW: 0.53; PL: 0.47; GEW: 0.87; EL: 2.32; TL: 3.29. Head (Fig. 41 B) wide, (GHW / HL: 1.12); eyes small (only slightly shorter than scape + pedicel), shallow, and moderately convex; temples prominent, larger than 1 / 3 length of eye, broad and rounded; occiput with slight raised areas; antennae about subequal to head and pronotum length combined; antennal scape moniliform, apically dilated, externally produced on one side; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter and moniliform, apically dilated; V – VI elongate, apically dilated; VII more dilated than VI; VIII – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 41 B) wide (GPW / PL: 1.13); texture very rugose with wide punctures; anterior margin without prominent emargination; anterolateral angles narrowly rounded, produced slightly below the anterior axis; posterior half subparallel laterally, much more contracted than anterior half; posterolateral angles square, rounded, not produced; pronotal lobe truncate with a hint of median notch; pronotal pad absent (Fig. 41 C); pronotal pits and grooves around moderate. Scutellar shield elliptical, transverse, about twice as wide as long. Elytra (Fig. 41 A) long (EL / GEW: 2.67), slightly convex; vestiture long and moderately dense; apical patch long and piceous. Ventral surface. Prosternal process moderately large and wide, extending mid length of procoxae; hypomeral edges well separated; mesosternal process narrow and linear. Color dull, rufopiceous with darker pronotum; ventrally inner edges of pro- and mesosternum piceous including the median processes; tarsi, labial palpi, glossal sclerite, and galea paler, other parts rufous.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE16E5C2A8DFF006AE62EE4.taxon	discussion	Remarks. Hemipeplus bucculentus sp. nov. is the only known species of Hemipeplinae with pronotum laterally widened near the anterior midpoint and not at the anterolateral angles. Geographical Distribution (Fig. 65). Laos (Vientiane Province).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE66E5E2A8DFA816C0A2EE5.taxon	description	(Figs 42, 65)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE66E5E2A8DFA816C0A2EE5.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ MALAYSIA, Pahang, Cameron Highlands, Brinchang, Tr 11, 7 - 18 - 1992, 1457 m. + / Collectors: L. B. & C. W. O’Brien / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus rugosus KC & Pollock’, in FSCA. Allotype, female, labeled: ‘ MALAYSIA, Pahang, Cameron Highlands, TANAH RATA vill. env. Gunung Jasar (Mt.): 1470 - 1705 m, 04 ° 28.4 - 7 ’ N, 101 ° 21.6 - 22.1 ’ E Jiří Hájek leg. 18. iv. - 10. v. 2009 / coll. general National Museum, Prague, Czech Republic’, in NMP 6 V. Fourteen paratypes: 1 specimen, sex unknown, labeled same as holotype, in FSCA; 6 specimens, 1 male, 5 sex unknown, labeled: ‘ MALAYSIA, Pahang, Fraser’s Hill, May-bankL. Tr., 7 - 14.1992, C. W. & L. B. O’Brien’, in FSCA; 4 specimens, 1 male, 3 sex unknown, labeled: ‘ MALAYSIA, Pahang, Fraser’s Hill, May-bankL. Tr., 7 - 15 - 1992, C. W. & L. B. O’Brien’, in FSCA; 1 specimen, sex unknown, labeled: ‘ MALAYSIA, Pahang, Fraser’s Hill, May-bankL. Tr. 29. vii. 1992, LB and CW O’Brien leg’, in FSCA; 1 specimen, male, labeled: ‘ Philippinen Mindanao, Surigao leg. Böttcher [underside of label] 5.8.1916 ’, in ZMHB; 1 specimen, male, labeled: ‘ SUMATRA (W.), G. Singgalang S of BUKITTINGGI 12. III. 1991, 1900 m, Bocák and Bocáková lgt. ’, in NHMB. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the species’ rough pronotum, characterized by many distinct, wide punctures. FIGURE 42. Hemipeplus rugosus sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE66E5E2A8DFA816C0A2EE5.taxon	diagnosis	Diagnosis. Hemipeplus rugosus sp. nov. can be distinguished from the similar-appearing H. miyamotoi (Figs 43.1, 43.2) by its relatively larger eyes, pronotum with coarser texture, and less produced anterolateral pronotal angles. Hemipeplus rugosus sp. nov. has the following diagnostic features: temples prominent; scape short and moniliform; eyes moderately large (as long as scape + pedicel), moderately wide and moderately convex; pronotum subcordiform; anterior emargination moderate but distinct; anterior half wider than posterior half; pronotal pad not very distinct; anterior pronotal angles broadly rounded, posterior rounded to slightly obtuse angled; pronotal lobe truncate without median notch; body long; elytra slightly convex; color dull, rufopiceous with darker head and pronotum to uniformly rufopiceous; distribution: Indonesia (Sumatra), Malaysia (Pahang), Philippines (Surigao).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE66E5E2A8DFA816C0A2EE5.taxon	description	Description. Measurements (in mm) — GHW: 0.53 – 0.60; HL: 0.50 – 0.53; AL: 0.78 – 0.95; GPW: 0.53 – 0.63; PL: 0.45 – 0.53, GEW: 0.60 – 0.88; EL: 2.30 – 3.03; TL: 3.25 – 4.09. Head (Fig. 42 B) wide (GHW / HL: 1.06 – 1.13); eyes moderately large (as long as scape + pedicel), moderately wide and moderately convex; temples prominent, about 1 / 3 length of eye, rounded posteriorly; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape moniliform; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter and submoniliform, apically dilated; V – VI larger, apically dilated; VII – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent to very small. Pronotum (Fig. 42 B) wide (GPW / PL: 1.18 – 1.19) with wide punctures; anterior margin with moderate emargination; anterolateral angles broadly rounded; lateral margins sinuate with subparallel basal half; posterior half contracted; posterolateral angles obtuse, rounded to angled; pronotal lobe truncate without median notch; pronotal pad (Fig. 42 C) indistinct; pronotal pits shallow without prominent grooves. Scutellar shield angulate, subhexagonal, not twice as wide as long. Elytra (Fig. 42 A) long (EL / GEW: 3.44 – 3.83), slightly convex; vestiture long and dense; apical patch long and piceous, not visible when elytra are same color. Ventral surface. Prosternal process short, not extending to the posterior of procoxae; hypomeral edges well separated; mesosternal process narrow and linear. Color dull, rufopiceous with darker head and pronotum to uniformly rufopiceous; ventrally ununiformly rufopiceous with tarsi, labial palps, glossal sclerite, and galea lighter. Male Genitalia (Figs 42 D – F). (Tegmen length = 0.13 mm (n = 1 )) Smallest among all Hemipeplinae species; lobe of basale long, needle-like, sharp and pointed; shoulders rounded; apicale and basale with distinct suture; parameres arcuate with apices sharp and pointed; wide penis subequal to tegmen. Note: the genitalia of H. rugosus sp. nov. are very similar to those of H. miyamotoi, but the parameres are different when viewed laterally, with their apices projecting upward resembling a snake’s head (Fig. 42 F).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE66E5E2A8DFA816C0A2EE5.taxon	discussion	Remarks. Hemipeplus rugosus sp. nov. is inferred to be the adelphotaxon of H. miyamotoi, as evidenced by the morphology of the male genitalia. Geographical Distribution (Fig. 65). Indonesia (Sumatra), Malaysia (Pahang), Philippines (Surigao).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE46E612A8DFA816B8E2FE1.taxon	materials_examined	Type Material. Holotype, sex unknown, in KUEC (Japan, Shinmura-Yuwan, Amami-Oshima); not studied, see Kamiya (1961) for details. Other Material Examined. JAPAN. Kyushu. Miyazaki. Udo Miyazaki 29. vii. 1974, T. N & A. N. leg., Brit. Mus. 1976 – 441, (BMNH, 1 ♀). Yakushima Koidomari 1. viii. 1991, T. Ueno leg., (DAPC, 1); Ryukyu Islands. Amami. Asani, 24. iii. 1967, H. Nomura leg., (FSCA, 1). Amami Ōshima. Sato, 1. iii. 1989, T Ueno leg., (FSCA, 1); Naze city, 2 - iii- 1989, T. Ueno leg., (DAPC, 2 ♀) (Fig. 43.2); same locality, 9. iv. 1971, M. Sakai leg., (DAPC, 1 ♀); Hatauno 3. iv. 1991, T. Ueno leg., (DAPC, 1 ♂). Iriomote Ohtomi-rindo, 1. viii. 1988, T. Ueno leg., (DAPC, 1). Kanna. Okinawa, 11. viii. 1945, Eugene Ray leg., at light, (FMNH, 1); same locality, 14. viii. 1945, Eugene Ray leg., at light, (FMNH, 1); same locality, 16. viii. 1945, Eugene Ray leg., at light, (FMNH, 1 ♂, 1 sex unknown). Okinawa. Yona, Kunigami, 19. iii. 1988, T. Ueno leg., (DAPC, 1). Tokara Island. Nakanoshima, 4. viii. 1989, T. Ueno leg., (DAPC, 1). TAIWAN. Hualien County. Piluhsi (Piluchi) For. Res. Stn. 22. v. 1999, 2150 m, C. W. & L. B. O’Brien leg., (FSCA, 1 ♂, 24 sex unknown); same locality, S. E. Halbert, C. W. & L. B. O’Brien leg., (FSCA, 3); same locality, 23. v. 1999, S. E. Halbert, C. W. & L. B. O’Brien leg., (FSCA, 3); same locality, 23. v. 1999, 2150 – 2300 m, C. W. & L. B. O’Brien leg., (FSCA, 6) (Fig. 43.1); same locality, 24. v. 1999, C. W. & L. B. O’Brien leg., (FSCA, 5); same locality, 24. v. 1999, 2150 m, S. E. Halbert, C. W. & L. B. O’Brien leg., (FSCA, 4); same locality, 26. v. 1999, 2100 m, C. W. & L. B. O’Brien leg., (FSCA, 1). Kaohsiung. Kaohsiung Hsien, rd. from Tengchin to Chuyanshan 1400 m, 25. iv. 1990, A. Smetana (T 23) leg., (MHNG, 1). Nantou County. Puli, Nanshan River, ca. 400 m, 24. v. 1999, C. W. & L. B. O’Brien leg., (FSCA, 1). Tainan. Tsaoshan (Sozan), 4 – 5. viii. 1963, 200 – 300 m, J. L. Gressitt leg., on bamboo, (BPBM, 1). Taipei. Pihu, 49 km E of Taipei, on Highway 9 (to Yilan), 18. ii. 1972, T. C. Maa leg., (BPBM, 1). Yilan County. Tsuifeng Lake, 17. v. 1999, 2400 m, S. E. Halbert, C. W. & L. B. O’Brien leg., (FSCA, 1); 7 km mark on Taipingshan / Tsuifeng Lake Rd., 17. v. 1999, S. Halbert, C & L O’Brien leg., (FSCA, 1). FIGURE 43. 1. Hemipeplus miyamotoi Kamiya, 1961 (Taiwan). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. FIGURE 43. 2. Hemipeplus miyamotoi Kamiya, 1961 (Japan). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE46E612A8DFA816B8E2FE1.taxon	diagnosis	Diagnosis. Hemipeplus miyamotoi can be distinguished by its small eyes, round temples, and strongly produced anterolateral pronotal angles. Hemipeplus miyamotoi has the following diagnostic features: temples prominent and rounded; scape short and moniliform; eyes smallest among all Austro-Oriental Hemipeplinae (shorter than scape + pedicel), shallow and slightly convex; pronotum subcordiform; anterior emargination moderate; anterior half much wider than posterior half, distinctly produced; pronotal pad not distinct; pronotal angles rounded; pronotal lobe indistinct without median notch; body long; elytra slightly convex; color dull, variable: uniformly testaceous to rufotestaceous, head and pronotum rufopiceous to strongly piceous with lighter elytra; distribution: Japan (Kyushu, Ryukyu Islands), Taiwan. Re-description. Measurements (in mm) — GHW: 0.57 – 0.68; HL: 0.52 – 0.58; AL: 0.93 – 1.10; GPW: 0.62 – 0.73; PL: 0.48 – 0.60, GEW: 0.80 – 1.06; EL: 2.40 – 3.40; TL: 3.40 – 4.58. Head (Figs 43.1 B, 43.2 B) wide (GHW / HL: 1.10 – 1.17); eyes smallest among all Austro-Oriental Hemipeplinae (shorter than scape + pedicel), shallow and not very convex; temples prominent, about 1 / 3 length of eye, large and rounded; occiput with raised areas; antennae about subequal to or slightly shorter than head and pronotum length combined; antennal scape short, moniliform; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter and moniliform, apically dilated; V – VI larger, apically dilated; VII – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent to very short. Pronotum (Figs 43.1 B, 43.2 B) wide (GPW / PL: 1.22 – 1.29); anterior margin with moderate emargination; anterolateral angles narrowly rounded, strongly produced; lateral margins deeply sinuate anteriorly; posterior half contracted with margins subparallel; posterolateral angles obtuse, rounded; pronotal lobe truncate without median notch; pronotal pad (Figs 43.1 C, 43.2 C) indistinct; pronotal pits shallow to deep with or without visible grooves. Scutellar shield elliptical to subquadrate, transverse, about twice as wide as long. Elytra (Figs 43.1 A, 43.2 A) long (EL / GEW: 3.00 – 3.21), slightly convex; vestiture long and dense; apical patch long, faint to dark. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges well separated; mesosternal process narrow and linear. Color dull, variable: uniformly testaceous to rufotestaceous; head and pronotum rufopiceous to shiny piceous in some specimens with rufous mouthparts; ventrally equally variable; some specimens with darker head and pronotum have mesosternal process darker as well; palpi, glossal sclerite, galea paler in most specimens; some specimens are uniformly testaceous to unevenly rufopiceous ventrally. Male Genitalia (Figs 43. 1 D – F). (Tegmen length = 0.20 mm (n = 1 )) One of the smallest male genitalia among all Hemipeplinae species; lobe of basale short, slender, and pointed (slightly broader than in H. rugosus sp. nov.); shoulders slightly obtuse angled; apicale and basale with distinct suture; parameres arcuate with tips thin, sharp and pointed; penis wide, subequal to tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE46E612A8DFA816B8E2FE1.taxon	description	Variation. Specimens of H. miyamotoi exhibit large variation in color. The head and pronotum are piceous while the elytra are light brown in some specimens from Japan (Fig. 43.2 B), while some are dark brown with head and pronotum only slightly darker (Fig. 43.1 B); some are uniformly light brown / testaceous.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFE46E612A8DFA816B8E2FE1.taxon	discussion	Remarks. Hemipeplus miyamotoi has the smallest eyes among all Hemipeplinae species in the Austro-Oriental region relative to the length of its antennomeres. As mentioned above, similarity of male genitalia between H. rugosus sp. nov. and H. miyamotoi suggests a possible close phylogenetic relationship. Geographical Distribution (Fig. 65). Japan (Kyushu, Ryukyu Islands), Taiwan (new country record). Natural History. According to specimen label data, H. miyamotoi has been found in association with bamboo (Poaceae) and adults have been collected at light.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDB6E632A8DF985685E2F59.taxon	description	(Figs 44, 65)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDB6E632A8DF985685E2F59.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ Philippinen Mindanao, Surigao leg. Böttcher [underside of label] 5.8.1916 / Zool. Mus. Berlin / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus joerebeccae KC & Pollock’, in ZMHB. Thirty-six paratypes: 16 specimens, sex unknown, labeled same as holotype, in ZMHB; 4 specimens, sex unknown, labeled: ‘ Philippinen Mindanao, Surigao leg. Böttcher [underside of label] 28.8.1916 ’, in ZMHB; 1 specimen, sex unknown, labeled: ‘ Davao Mindanao Baker / 7 / 1 / 15 ’, in NMNH; 5 specimens, sex unknown, labeled: ‘ Philippinen Mindanao, Surigao leg. Böttcher [underside of label] 30.8.1916 ’, in ZMHB; 2 specimens, sex unknown, labeled: ‘ Philippinen Mindanao, Surigao leg. Böttcher [underside of label] 9.9.1916 ’, in ZMHB; 1 specimen, sex unknown, labeled: ‘ Philippinen Mindanao, Surigao leg. Böttcher [underside of label] 10.8.1916 ’, in ZMHB; 1 specimen, male, labeled: ‘ Philippinen, Dapa, Siargao / Dapa Siargao / F. Nevermann Staudinger vend. 11. X. 1930 / Nevermann Collection 1940 / Hemipeplus klematanicus [followed by something illegible] ’, in NMNH; 1 specimen, sex unknown, labeled: ‘ Philippinen, N-Luzon, Bangui. Jan. 1918, leg. Böttcher’, in ZMHB; 1 specimen, female, labeled: ‘ Philippinen, Mangarin. XI. 1917 leg. Böttcher’, in ZMHB; 1 specimen, sex unknown, labeled: ‘ P. I., NEGROS OR., Damaguete, 390 m, 26. VI. 1958 / Light Trap H. E. Milliron’, in BPBM; 3 specimens, female, labeled: ‘ Philippinen N. Luzon [something illegible] [underside of label] [something illegible] I. 1918 ’, in ZMHB. FIGURE 44. Hemipeplus joerebeccae sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. Derivation of Specific Epithet. The specific name (plural noun in the genitive case) is given in allusion to the first author’s friends Joe Schelling and Mary Rebecca Gracey from Albuquerque, New Mexico.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDB6E632A8DF985685E2F59.taxon	diagnosis	Diagnosis. Hemipeplus joerebeccae sp. nov. can be distinguished by its large eyes, short temples, pronotum narrower and color deeper than in H. luzonensis sp. nov. (Fig. 45). Hemipeplus joerebeccae sp. nov. has the following diagnostic features: temples extremely short, indistinguishable in some specimens, angulate to roundish; scape moniliform; eyes large (as long as scape + pedicel + antennomere III), wide and convex; pronotum subcordiform; anterior emargination distinct; anterior half wider than posterior half; pronotal pad distinct; pronotal angles rounded; pronotal lobe truncate without median notch; body long; elytra slightly convex; color dull, rufous to rufotestaceous, head and pronotum slightly darker; distribution: Philippines (Luzon, Mindanao (Surigao), Mindoro, Negros).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDB6E632A8DF985685E2F59.taxon	description	Description. Measurements (in mm) — GHW: 0.48 – 0.64; HL: 0.43 – 0.57; AL: 0.75 – 0.92; GPW: 0.50 – 0.72; PL: 0.45 – 0.60, GEW: 0.58 – 0.83; EL: 2.10 – 3.10; TL: 2.98 – 4.27. Head (Fig. 44 B) wide (GHW / HL: 1.12); eyes large (as long as scape + pedicel + antennomere III), wide and convex; temples extremely short, indistinguishable in some specimens, angulate to roundish; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape moniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter and moniliform, apically dilated; V larger, moniliform; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent to very small. Pronotum (Fig. 44 B) wide (GPW / PL: 1.11 – 1.20); anterior margin with distinct emargination; anterolateral angles rounded, produced; lateral margins sinuate on anterior half; posterior half contracted; posterolateral margins subparallel; posterolateral angles obtuse, rounded, not produced; pronotal lobe indistinct / truncate without median notch; pronotal pad (Fig. 44 C) distinct but narrow; pronotal pits moderately deep with moderate grooves. Scutellar shield subhexagonal, not twice as wide as long. Elytra (Fig. 44 A) long (EL / GEW: 3.62 – 3.73), slightly convex; vestiture very short but dense; apical patch long and only slightly darker than elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges very close together to fused; mesosternal process narrow and linear. Color dull, rufous to rufotestaceous, head and pronotum slightly darker. Male Genitalia (Figs 44 D – F). (Tegmen length = 0.35 mm (n = 1 )) This species has one of the most unique male genitalia structures among all Austro-Oriental Hemipeplinae species; lobe of basale very long (longest among all Austro-Oriental congeners), longer than rest of the tegmen combined, very thin, and apically rounded; shoulders acutely pointed upward; parameres broad with apical half arcuate, tips sharp and narrowed, triangular followed by notch in inner margins; penis very long, longer than tegmen. Variation. Some specimens of H. joerebeccae sp. nov. have very small temples, creating confusion with H. harkoneni sp. nov. which does not possess temples. Geographical Distribution (Fig. 65). Philippines (Luzon, Mindanao (Surigao), Mindoro, Negros). Natural History. According to specimen label data, adults of H. joerebeccae sp. nov. have been collected from light traps.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD96E652A8DFA5D681828DF.taxon	description	(Figs 45, 65)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD96E652A8DFA5D681828DF.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ P. I.: LUZON CAMARINES SUR Mt. Isarog, Pili, 800 - 900 m, 21. IV. ’ 65 / H. M. Torrevillas Light Trap BISHOP MUSEUM / [light green label] BPBM / [blue label] ♀ / [red label] HOLOTYPE ♀ Hemipeplus luzonensis KC & Pollock’, in BPBM. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the type locality of this species, i. e., Luzon, Philippines.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD96E652A8DFA5D681828DF.taxon	diagnosis	Diagnosis. Hemipeplus luzonensis sp. nov. can be distinguished from the similar-appearing H. joerebeccae sp. nov. (Fig. 44) by its relatively smaller eyes, wider pronotum, and paler color; it can be distinguished from the similar-appearing H. harkoneni sp. nov. by its short and angulate temples. Hemipeplus luzonensis sp. nov. has the following diagnostic features: temples very short and angulate; scape moniliform; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum subcordiform; anterior emargination deep and distinct; anterior half much wider than posterior half; pronotal pad narrow but distinct; anterior pronotal angles rounded and strongly produced; posterior angles obtuse, rounded; pronotal lobe very slight with faint median notch; body long; elytra more or less flat; color dull, uniformly rufotestaceous; distribution: Philippines (Luzon). FIGURE 45. Hemipeplus luzonensis sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD96E652A8DFA5D681828DF.taxon	description	Description. Measurements (in mm) — GHW: 0.74; HL: 0.71; AL: 1.12; GPW: 0.87; PL: 0.70; GEW: 1.00; EL: 3.70; TL: 5.11. Head (Fig. 45 B) wide (GHW / HL: 1.04); eyes large (longer than scape + pedicel), moderately wide and moderately convex; temples very short, angulate; occiput with strongly raised areas; antennae shorter than head and pronotum length combined; antennal scape moniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV small, submoniliform; V more elongate, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth potentially absent. Pronotum (Fig. 45 B) wide (GPW / PL: 1.24), subcordiform; anterior margin with deep and distinct emargination in the center; anterolateral angles rounded, strongly produced; lateral margins sinuate; posterior half contracted; posterolateral angles obtuse, rounded; pronotal lobe very faint with a hint of median notch; pronotal pad (Fig. 45 C) narrow, but well distinct; pronotal pits deep with moderate grooves. Scutellar shield subhexagonal, flat posterior margin, transverse, twice as wide as long. Elytra (Fig. 45 A) long (EL / GEW: 3.70), more or less flat; vestiture short and dense; apical patch long and slightly darker than elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges fused; mesosternal process narrow and linear. Color dull, uniformly rufotestaceous. Geographical Distribution (Fig. 65). Philippines (Luzon). Natural History. According to specimen label data, the single specimen of H. luzonensis sp. nov. was collected from a light trap.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDF6E672A8DFCDA6B3029E3.taxon	materials_examined	Type Material. Ochrosanis klematanica, Holotype, sex unknown, in MSNG (Malaysia, Sarawak); not studied, see Gestro (1873) for details. Other Material Examined. INDONESIA. Northern Sulawesi. Sulawesi Utara, Dumoga-Bone NP (now Bogani Nani Wartabone National Park),?. ii. 1985, Lowland Forest ca 200 m, Malaise trap, R. Ent. Soc. Lond. Project Wallace BM 1985 – 10, (BMNH, 1) (Fig. 46); same locality, 11. ii. 1985, Lowland Forest ca 200 m Line 1, carrion baited trap, R. Ent. Soc. Lond. Project Wallace BM 1985 – 10, (BMNH, 1); same locality, Tray 49, Fog 5 400 m, 11. ii. 1985 BMNH Plot C., R. Ent. Soc. Lond. Project Wallace BM 1985 – 10, (BMNH, 1); same locality, Tray 50, Fog 5 400 m, 11. ii. 1985 BMNH Plot C., R. Ent. Soc. Lond. Project Wallace BM 1985 – 10, (BMNH, 1 ♂); same locality, Tray 60, Fog 5 400 m, 11. ii. 1985 BMNH Plot C., R. Ent. Soc. Lond. Project Wallace BM 1985 – 10, (BMNH, 1); same locality, Tray 63, Fog 11 230 m, 10. iii. 1985 BMNH Plot A., R. Ent. Soc. Lond. Project Wallace BM 1985 – 10, (BMNH, 1); same locality, Plot A, ca 200 m lowland forest, flight interception trap,?. iv. 1985, (BMNH, 1); same locality, malaise trap, Fog 2 lowland forest edge ca 200 m, 9 – 16. v. 1985 BMNH Plot C., R. Ent. Soc. Lond. Project Wallace BM 1985 – 10, (BMNH, 1); same locality, site 2, ca 220 m, Rothamsted light trap,?. xi. 1985 coll. H. Barlow R. Ent. Soc. Lond. Project Wallace BM 1985 – 10 Project Wallace morphospecies 110.1: 8 of 9 exx 18 August 1995, (BMNH, 1). MALAYSIA. Penang. 13. x. 1913, G. E. Bryant leg., No. 8.13, G. Bryant Coll. 1919 — 147, Hemipeplus klematanicus (Gestro) det. D. A. Pollock, (BMNH, 1). Sarawak. Jalan Biawak, 18. vi. 1968, S. Kueh leg., on Nipah palm, Pres. by Com Inst. Ent. BM 1971 - I, 9158 (for female), 9045 (for unknown sex), (BMNH, 1 ♀, 1 sex unknown). Selangor. Klang, 30. xii. 1947, W. H. Lange leg., on Nypa fruticans, (NMNH, 2); same locality, 1. ii. 1948, W. H. Lange leg., on Nypa fruticans, (NMNH, 2). THAILAND. Phang Nga Province. Koh Ra, 20 m, MT 9 ° 10 ’ 47 ’’ N, 98 ° 16 ’ 0 ’’ E, 18 – 21. xii. 2001 G. Ballmer leg., Univ. Calif. Riverside, Ent. Res. Museum, UCRC ENT 73110, (UCRC, 1).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDF6E672A8DFCDA6B3029E3.taxon	diagnosis	Diagnosis. Hemipeplus klematanicus can be distinguished by its short, acutely pointed temples and acutely pointed posterior pronotal angles. Hemipeplus klematanicus has the following diagnostic features: temples short and acute angled; scape submoniliform; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum subcordiform; anterior emargination distinct; anterior half wider than posterior half; pronotal pad slight; anterior pronotal angles rounded, posterior angles sharply acute; pronotal lobe slightly present with faint median notch; body long; elytra slightly convex; color dull, rufotestaceous to rufopiceous, head and pronotum darker; distribution: Indonesia (Northern Sulawesi), Malaysia (Sarawak, Selangor), Thailand (Phang Nga). FIGURE 46. Hemipeplus klematanicus (Gestro, 1873) (Genitalia from a different specimen). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. Re-description. Measurements (in mm) — GHW: 0.50 – 0.64; HL: 0.48 – 0.62; AL: 0.85 – 1.05; GPW: 0.55 – 0.68; PL: 0.43 – 0.58; GEW: 0.73 – 0.90; EL: 2.38 – 3.25; TL: 3.29 – 4.45. Head (Fig. 46 B) wide (GHW / HL: 1.03 – 1.04); eyes large (longer than scape + pedicel), moderately wide and moderately convex; temples short and acutely angled; occiput with raised areas; antennae slightly shorter than head and pronotum length combined; antennal scape submoniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter and moniliform, apically dilated; V – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth very short. Pronotum (Fig. 46 B) wide (GPW / PL: 1.17 – 1.28); anterior margin with distinct emargination; anterolateral angles rounded, produced; lateral margins sinuate; posterior half slightly contracted; posterolateral angles acutely pointed; pronotal lobe slight with faint median notch; pronotal pad (Fig. 46 C) slight; pronotal pits moderately deep with moderate grooves. Scutellar shield transverse, subhexagonal, about twice as wide as long. Elytra (Fig. 46 A) long (EL / GEW: 3.26 – 3.61), slightly convex; vestiture long and dense; apical patch long and more or less darker than elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges very close together, only narrowly separated; mesosternal process narrow and linear. Color dull, rufotestaceous to rufopiceous, head and pronotum darker; ventrally unevenly rufous with edges of sternal plates darker; palpi and tarsi paler. Male Genitalia (Figs 46 D – F). (Tegmen length = 0.30 mm (n = 1 )) Lobe of basale long and slender; shoulders obtusely angled with moderate slant; parameres feebly arcuate at apices with tips thick and blunt; penis longer than tegmen with narrowed apex.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDF6E672A8DFCDA6B3029E3.taxon	description	Variation. The specimen of H. klematanicus from Thailand is much smaller than other specimens examined. Geographical Distribution (Fig. 65). Indonesia (Northern Sulawesi) (new country record), Malaysia (Sarawak, Selangor), Thailand (Phang Nga) (new country record). Natural History. According to specimen label data, H. klematanicus has been found in unopened fronds of Nypa fruticans (Arecaceae) and adults have been collected from malaise traps, flight interception trap, carrion baited trap in lowland forest, and light trap.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDD6E692A8DFB8368CF2BA8.taxon	description	(Figs 47, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDD6E692A8DFB8368CF2BA8.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ AUSTRALIA: Northern Terr., 25 kmWStuart Hwy on Van Arnham [Arnhem] Hwy., 25 - IX- 1992, D. H. Habeck on palm flowers / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus neoaustralicus KC & Pollock’, in FSCA. Paratype: 1 specimen, sex unknown, labeled same as holotype, in FSCA. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the species’ type locality, Australia.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDD6E692A8DFB8368CF2BA8.taxon	diagnosis	Diagnosis. Hemipeplus neoaustralicus sp. nov. is the only Australian species with temples and dark brown body. Hemipeplus neoaustralicus sp. nov. has the following diagnostic features: temples short and rounded which are rather extensions due to large eyes and short occiput; scape short and moniliform, apically dilated and externally produced on one side; eyes large (longer than scape + pedicel), wide and convex; pronotum subcordiform with flat disc; anterior emargination wide; anterior half much wider than posterior half, and produced; pronotal pad narrow but distinct; anterior pronotal angles narrowly rounded, posterior angles obtuse, angulate; pronotal lobe slight but wide and arcuate with indistinct median notch; body long and flat; color dull, uniformly dark brown; distribution: Australia (Northern Territory).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFDD6E692A8DFB8368CF2BA8.taxon	description	Description. Measurements (in mm) — GHW: 0.78 – 0.88; HL: 0.61 – 0.70; AL: 1.01 – 1.12; GPW: 0.88 – 0.95; PL: 0.65 – 0.72, GEW: 1.03 – 1.23; EL: 3.15 – 3.47; TL: 4.41 – 4.89. Head (Fig. 47 B) wide (GHW / HL: 1.26 – 1.28); eyes large (longer than scape + pedicel), wide and convex; temples short and rounded which are rather extensions due to large eyes and short occiput; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape moniliform, apically dilated, externally produced on one side; pedicel small moniliform; antennomere III more elongate and apically dilated; IV – V shorter and moniliform, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 47 B) wide (GPW / PL: 1.32 – 1.35) with flat disc; anterior margin with moderately wide emargination; anterolateral angles narrowly rounded, strongly produced; lateral margins sinuate; posterior half contracted; posterolateral angles obtuse, somewhat angulate and produced; pronotal lobe small but wide and arcuate with faint median notch; pronotal pad (Fig. 47 C) distinct and narrow; pronotal pits deep with prominent elliptical grooves. Scutellar shield subpentagonal, transverse, less than twice as wide as long. Elytra (Fig. 47 A) long (EL / GEW: 2.82 – 3.06) and flat; vestiture short and dense; apical patch wide, darker than elytra. Ventral surface. Prosternal process moderate, extending to mid length of procoxae; hypomeral edges well separated; mesosternal process narrow and linear. Color dull, uniformly dark brown; ventrally hypomera, mesanepisterna, and mesepimera slightly darker. FIGURE 47. Hemipeplus neoaustralicus sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. Male Genitalia (Figs 47 D – F). (Tegmen length = 0.27 mm (n = 1 )) Lobe of basale long and slim with pointed apex; shoulders obtusely angled, moderately slanted; apicale and basale with distinct suture; parameres strongly arcuate with tips blunt; penis longer than tegmen. The genitalia are similar to those of other Australian species but the parameres are significantly expanded laterally, or arcuate. Geographical Distribution (Fig. 64). Australia (Northern Territory). Natural History. According to specimen label data, H. neoaustralicus sp. nov. was found associated with palm flowers (Arecaceae).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD36E6A2A8DFE4D6DED2D44.taxon	description	(Figs 48, 65)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD36E6A2A8DFE4D6DED2D44.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ [oval label with green color on upper side, blank] / [underlined in yellow] Sarawak, [/ underlined in yellow] Borneo, Wallace. / Pascoe Coll. 93 – 60. / Hemipeplus [something illegible] / Hemipeplus klematanicus, Gestro Determined from description. G. J. A. / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus pseudoklematanicus KC & Pollock’, in BMNH. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the species’ close morphological resemblance to Hemipeplus klematanicus.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD36E6A2A8DFE4D6DED2D44.taxon	diagnosis	Diagnosis. Hemipeplus pseudoklematanicus sp. nov. can be distinguished from the similar-appearing H. klematanicus (Fig. 46) by its rounded, instead of acutely pointed, temples, wider pronotum, and paler color. Hemipeplus pseudoklematanicus sp. nov. has the following diagnostic features: temples short and rounded; scape moniliform, apically dilated; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum subcordiform; anterior emargination distinct; anterior half slightly wider than posterior half; pronotal pad distinct; anterior pronotal angles rounded, posterior angles obtuse; pronotal lobe slightly present without median notch; body long; elytra slightly convex; color dull, rufotestaceous, not nearly as dark as H. klematanicus, pronotum lighter; distribution: Malaysia (Sarawak).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD36E6A2A8DFE4D6DED2D44.taxon	description	Description. Measurements (in mm) — GHW: 0.65; HL: 0.65; AL: 1.23; GPW: 0.75; PL: 0.60; GEW: 0.93; EL: 2.84; TL: 4.09. Head (Fig. 48 B) about as long as wide (GHW / HL: 1); eyes large (longer than scape + pedicel), moderately wide and moderately convex; temples short and rounded; occiput with strongly raised areas; antennae subequal to head and pronotum length combined; antennal scape moniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter and moniliform, apically dilated; V moniliform, larger, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth unobservable in the specimen. Pronotum (Fig. 48 B) wide (GPW / PL: 1.25); anterior margin with distinct emargination; anterolateral angles rounded, produced; lateral margins moderately sinuate; posterior half contracted; posterolateral angles obtuse; pronotal lobe slight without median notch; pronotal pad distinct (Fig. 48 C); pronotal pits deep with moderate grooves. Scutellar shield transverse, subpentagonal, about twice as wide as long. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges not very close together unlike in H. klematanicus, and well separated; mesosternal process narrow and linear. Elytra (Fig. 48 A) long (EL / GEW: 3.05), slightly convex; vestiture short and dense; apical patch broad and rufous. Color dull, rufotestaceous, pronotum lighter. Male Genitalia (Figs 48 D – F). (Tegmen length = 0.29 mm) Lobe of basale short and apically pointed; shoulders obtusely angled with moderate slant; parameres arcuate apically with corners of apices darker which is because of an additional lobe that extends behind them (visible on lateral view) (Fig. 48 F); apices thick and blunt; penis longer than tegmen, shrunk at the halfway point and apically.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD36E6A2A8DFE4D6DED2D44.taxon	discussion	Remarks. Hemipeplus pseudoklematanicus sp. nov. superficially resembles H. klematanicus (Fig. 46); both species are sympatric in Malaysia (Sarawak). Geographical Distribution (Fig. 65). Malaysia (Sarawak). FIGURE 48. Hemipeplus pseudoklematanicus sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD16E6C2A8DFF006A372F11.taxon	description	(Figs 49, 65) FIGURE 49. Hemipeplus philippinensis sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD16E6C2A8DFF006A372F11.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ P. I., PALAWAN, Eran Pt., 8 km SW Tarumpitao Pt. 31. XII. 59 - 4. I. 60 / L. W. Quate Collector / [light green label] BPBM / [blue label] ♀ / [red label] HOLOTYPE ♀ Hemipeplus philippinensis KC & Pollock’, in BPBM. One paratype [without antennae]; female, same label as in holotype, in BPBM. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the type locality of this species, i. e., Philippines.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD16E6C2A8DFF006A372F11.taxon	diagnosis	Diagnosis. Hemipeplus philippinensis sp. nov. can be distinguished by its short, angulate temples and subquadriform pronotum. Hemipeplus philippinensis resembles H. nuciferae (Fig. 52) and H. palawanensis sp. nov. (Fig. 53); it is sympatric with the latter. However, the pronotum in H. philippinensis is subquadriform while it is subcordiform in H. nuciferae which also has much larger and more convex eyes (Fig. 52 B). It differs from H. palawanensis sp. nov. (Fig. 53 B) in having short and angulate temples instead of short and rounded temples, larger eyes, and pronotum less quadriform. Hemipeplus philippinensis sp. nov. has the following diagnostic features: temples short and strongly angulate; scape moniliform, apically dilated; eyes large (longer than scape + pedicel), shallow and less convex; pronotum about subquadriform; anterior emargination shallow but distinct, moderately deep in the center; anterior half wider than posterior half; pronotal pad narrow but distinct; anterior pronotal angles rounded, moderately produced; posterior angles obtuse, rounded; pronotal lobe slight with faint median notch; body long; elytra slightly convex; color dull, about uniformly rufotestaceous; distribution: Philippines (Palawan).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD16E6C2A8DFF006A372F11.taxon	description	Description. Measurements (in mm) — GHW: 0.52 – 0.57; HL: 0.48 – 0.50; AL: 0.88; GPW: 0.55 – 0.65.; PL: 0.45 – 0.55; GEW: 0.64 – 0.77; EL: 2.48 – 2.93; TL: 3.41 – 3.98. Head (Fig. 49 B) wide (GHW / HL: 1.08 – 1.14); eyes large (longer than scape + pedicel), shallow and less convex; temples short, acutely pointed; occiput with strongly raised areas; antennae shorter than head and pronotum length combined; antennal scape moniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV small, moniliform; V – VI larger, subequal, moniliform, apically dilated; VII – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 49 B) wide (GPW / PL: 1.18 – 1.22); anterior margin with shallow but distinct emargination in the center; anterolateral angles rounded, moderately produced; lateral margins sinuate; posterior half slightly more contracted than anterior half; posterolateral angles obtuse, rounded; pronotal lobe slight with median notch; pronotal pad (Fig. 49 C) narrow, but well distinct; pronotal pits moderately deep with moderate grooves. Scutellar shield elliptical, transverse, not twice as wide as long. Elytra (Fig. 49 A) long (EL / GEW: 3.80 – 3.88), slightly convex; vestiture moderately long and dense; apical patch long but faint. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges narrowly separated, narrower than in the sympatric H. palawanensis sp. nov.; mesosternal process narrow and linear. Color dull, about uniformly rufotestaceous. Variation. Variation exists in pronotal pad which is more prominent in the holotype than paratype. Geographical Distribution (Fig. 65). Philippines (Palawan).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD66E6E2A8DFA15686829E9.taxon	description	(Figs 50, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD66E6E2A8DFA15686829E9.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ NEW GUINEA: NE. Madang. 5 m. Oct. 22, 1958 / J. L. Gressitt Collector / Palm / [light green label] BPBM / [red label] HOLOTYPE ♂ Hemipeplus exhaustus KC & Pollock’, in BPBM. One paratype, female, labeled: ‘ NEW GUINEA: NE. Madang, 5 m. Oct. 28, 1958 / J. L. Gressitt Collector / Palm’, in BPBM. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the sheer exhaustion the first author had while describing tens of obscure species of Hemipeplinae.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD66E6E2A8DFA15686829E9.taxon	diagnosis	Diagnosis. Hemipeplus exhaustus sp. nov. can be distinguished by its large eyes, short, rounded temples, and rufotestaceous color. It resembles H. enarotaliensis sp. nov. (Fig. 51) (similar genitalia) but is distinctly smaller in body size with eyes distinctly larger resembling those of H. neoguineensis sp. nov. (Fig. 37 B) (similar genitalia); however, unlike H. neoguineensis sp. nov., H. exhaustus sp. nov. has temples. Hemipeplus exhaustus sp. nov. has the following diagnostic features: temples short and roundish; scape submoniliform, slightly elongate, apically dilated; eyes large (as long as scape + pedicel + antennomere III), wide and convex; pronotum subcordiform; anterior emargination shallow but distinct; anterior half much wider than posterior half; pronotal pad narrow, distinct; pronotal angles rounded; pronotal lobe truncate without median notch; body long; elytra slightly convex; color dull, uniformly rufotestaceous; distribution: Papua New Guinea (Madang). FIGURE 50. Hemipeplus exhaustus sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD66E6E2A8DFA15686829E9.taxon	description	Description. Measurements (in mm) — GHW: 0.53 – 0.55; HL: 0.50 – 0.53; AL: 0.79 – 0.80; GPW: 0.55; PL: 0.45 – 0.48, GEW: 0.70; EL: 2.26; TL: 3.21 – 3.27. Head (Fig. 50 B) wide (GHW / HL: 1.04 – 1.06); eyes large (as long as scape + pedicel + antennomere III), wide and convex; temples short and rounded; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape submoniliform, slightly elongate, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter and moniliform, apically dilated; V – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 50 B) wide (GPW / PL: 1.15 – 1.22); anterior margin with shallow but distinct emargination; anterolateral angles rounded, produced; lateral margins sinuate; posterior half sharply contracted; posterolateral angles obtuse, rounded; pronotal lobe truncate without median notch; pronotal pad (Fig. 50 C) distinct; pronotal pits moderately deep with moderate grooves. Scutellar shield subhexagonal, transverse, about twice as wide as long. Elytra (Fig. 50 A) long (EL / GEW: 3.23), slightly convex; vestiture short and dense; apical patch long and slightly darker than the elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges fused; mesosternal process narrow and linear. Color dull, uniformly rufotestaceous. Male Genitalia (Figs 50 D – F). (Tegmen length = 0.25 mm (n = 1 )) Lobe of basale short and slender with rounded apex; shoulders obtusely angled with moderate slant; apicale and basale with distinct suture; parameres arcuate with tips blunt; penis longer than tegmen. The genitalia are similar to those of H. australicus.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD66E6E2A8DFA15686829E9.taxon	discussion	Remarks. Hemipeplus exhaustus sp. nov., similar to H. enarotaliensis sp. nov. and H. neoguineensis sp. nov., is closely related to the Australian species in terms of male genitalia morphology, suggesting possible common ancestry. Geographical Distribution (Fig. 64). Papua New Guinea (Madang). Natural History. According to specimen label data, H. exhaustus sp. nov. is associated with palms (Arecaceae).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD46E702A8DFB8D6A3E28A1.taxon	description	(Figs 51, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD46E702A8DFB8D6A3E28A1.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ Enarotali / J. & M. Sedlacek Collectors BISHOP / [light green label] BPBM / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus enarotaliensis KC & Pollock’, in BPBM. Allotype, female, labeled: ‘ NEW GUINEA: (NW) Wisselmeren Enarotadi [Enarotali] 1800 - 1900 m, 1 – 6 [corrected with an ink]. VIII. ’ 62 / J. Sedlacek Collector BISHOP’, in BPBM. Six paratypes: 4 specimens, sex unknown, labeled same as allotype; 1 specimen, male, labeled: ‘ NEW GUINEA: (NW) Wisselmeren Enarotadi [Enarotali] 1800 – 1900 m, 1 - 9. VIII. ’ 62, J. Sedlacek Collector BISHOP’, in BPBM; 1 specimen, sex unknown, labeled: ‘ NEW GUINEA: (NW) Wisselmeren Enarotadi [Enarotali] 1850 m, 5 – 9. VIII. 1962, J. M. Sedlacek Collector BISHOP’, in BPBM. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the species’ type locality, i. e., Enarotali, Indonesia.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD46E702A8DFB8D6A3E28A1.taxon	diagnosis	Diagnosis. Hemipeplus enarotaliensis sp. nov. can be distinguished from the similar-appearing H. nuciferae (Fig. 52) by its less emarginate anterior pronotal margin and rufous color. Hemipeplus enarotaliensis sp. nov. has the following diagnostic features: temples short and rounded; scape submoniliform, apically dilated; eyes large (longer than scape + pedicel), moderately wide and moderately convex; pronotum subcordiform; anterior emargination indistinct; anterior half much wider than posterior half; pronotal pad narrow, distinct; pronotal angles rounded; pronotal lobe truncate without median notch; body long; elytra slightly convex; color dull, rufous head and pronotum with rufous to rufotestaceous elytra; distribution: Indonesia (Enarotali).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD46E702A8DFB8D6A3E28A1.taxon	description	Description. Measurements (in mm) — GHW: 0.58 – 0.65; HL: 0.50 – 0.58; AL: 0.97 – 1.00; GPW: 0.55 – 0.65; PL: 0.50 – 0.58, GEW: 0.70 – 0.83; EL: 2.85 – 3.31; TL: 3.85 – 4.47. Head (Fig. 51 B) wide (GHW / HL: 1.12 – 1.16); eyes large (longer than scape + pedicel), moderately wide and moderately convex; temples short and rounded; occiput with raised areas; antennae slightly shorter than head and pronotum length combined; antennal scape submoniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter and moniliform, apically dilated; V – VI larger, apically dilated; VII – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; FIGURE 51. Hemipeplus enarotaliensis sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. lateral mandibular tooth absent. Pronotum (Fig. 51 B) wide (GPW / PL: 1.10 – 1.12); anterior margin with indistinct emargination; anterolateral angles rounded, produced; lateral margins sinuate; posterior half contracted; posterolateral angles obtuse, rounded, not produced; pronotal lobe truncate without median notch; pronotal pad (Fig. 51 C) distinct but narrow; pronotal pits moderately deep with moderate grooves. Scutellar shield elliptical, transverse, about twice as wide as long. Elytra (Fig. 51 A) long (EL / GEW: 3.99 – 4.07), slightly convex; vestiture short and dense; apical patch long and slightly darker than the elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges fused to very narrowly separated; mesosternal process narrow and linear. Color dull, rufous head and pronotum with rufous to rufotestaceous elytra; ventrally uniformly rufous to rufopiceous. Male Genitalia (Figs 51 D – F). (Tegmen length = 0.33 mm (n = 1 )) Lobe of basale short and pointed; shoulders about round and smooth, moderately slanted; apicale and basale with distinct suture; parameres arcuate with tips blunt, as if cut; penis longer than tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFD46E702A8DFB8D6A3E28A1.taxon	discussion	Remarks. The male genitalia of H. enarotaliensis sp. nov. resemble those of Australian species, suggesting possible common ancestry. Geographical Distribution (Fig. 64). Indonesia (Enarotali).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCA6E722A8DFD456B6E2841.taxon	materials_examined	Type Material. Lectotype (here designated; Article 74 (ICZN 1999 )), male, labeled: ‘ [round label with red margin] Type / DUTCH E. Indies, Sangi. 1928. A. Ryne. On unfolded leaves of coconut. / Pres. By. Imp. Bur. Ent. Brit. Mus. 1926 — 376 / Hemipeplus Coll. 4087 Det. G. E. Bryant / Hemipeplus nuciferae, type arrow / [red label] LECTOTYPE Hemipeplus nuciferae Arrow Designated by D. A. Pollock 1997 / [with a barcode] NHMUK 014663489 / [green label] ♂ ’, in BMNH. Paralectotype, sex unknown, labeled: ‘ [round label with yellow margin] Co-type / DUTCH, E. Indies, Sangi. 1928. A. Ryne. On unfolded leaves of coconut. / Pres. By. Imp. Bur. Ent. Brit. Mus. 1926 — 376 / Hemipeplus sp. [something illegible] marginipennis, LEC Det. G. E. Bryant / Hemipeplus nuciferae co-type arrow’, in BMNH. Other Material Examined. INDONESIA. Northern Sulawesi. Dumoga-Bone NP (now Bogani Nani Wartabone National Park),? 19. vii. 1985, Tray 35, Fog 15, 400 m, BMNH, Plot C, E. Ent. Soc. London. Project Wallace, BM 1985 – 10, (BMNH, 1 ♀, 1 ♂). South East Sulawesi. Kolaka,?. xi. 1930, R. Awibowo leg., on coconut leaves, (BMNH, 2).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCA6E722A8DFD456B6E2841.taxon	diagnosis	Diagnosis. Hemipeplus nuciferae can be distinguished from the similar-appearing H. enarotaliensis sp. nov. (Fig. 51) by its wider, more emarginate anterior pronotal margin and paler color. Hemipeplus nuciferae has the following diagnostic features: temples short and angulate; scape short, moniliform; eyes large (longer than scape + pedicel), wide and convex; pronotum subcordiform; anterior emargination shallow but distinct; anterior half produced, much wider than posterior half; pronotal pad distinct; pronotal angles rounded; pronotal lobe truncate without median notch; body long; elytra slightly convex; color dull, rufotestaceous; distribution: Indonesia (Sulawesi). Re-description. Measurements (in mm) — GHW: 0.54 – 0.60; HL: 0.47 – 0.57; AL: 0.80 – 0.90; GPW: 0.58 – 0.64; PL: 0.47 – 0.52; GEW: 0.67 – 0.73; EL: 2.63 – 2.72; TL: 3.57 – 3.81. Head (Fig. 52 B) wide (GHW / HL: 1.05 – 1.15); eyes large (longer than scape + pedicel), wide and convex; temples short and angulate; occiput with raised areas; antennae shorter than head and pronotum length combined; antennal scape moniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter and moniliform, apically dilated; V larger, moniliform; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 52 B) wide (GPW / PL: 1.23); anterior margin with shallow emargination; anterolateral angles rounded, produced; posterior half contracted; posterolateral margins subparallel; posterolateral angles obtuse, rounded, not produced; pronotal lobe indistinct / truncate without median notch; pronotal pad (Fig. 52 C) distinct; pronotal pits moderately deep with moderate grooves. Scutellar shield transverse, elliptical to FIGURE 52. Hemipeplus nuciferae Arrow, 1930 (Lectotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view. subpentagonal, less than twice as wide as long. Elytra (Fig. 52 A) long (EL / GEW: 3.73 – 3.93), slightly convex; vestiture short and dense; apical patch long and broad, rufopiceous. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges fused; mesosternal process narrow and linear. Color dull, uniformly rufotestaceous, head and pronotum darker in one specimen; ventrally uniformly rufotestaceous with palpi, glossal sclerite, galea, and tarsi paler. Male Genitalia (Figs 52 D – F). (Tegmen length = 0.33 mm (n = 1 )) Lobe of basale moderately long, apically dilated and narrowed at the tip; shoulders slanted, obtusely angled; parameres sinuate with apical half slightly bulged, tips moderately sharp and narrowed, subtriangular; penis longer than tegmen.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCA6E722A8DFD456B6E2841.taxon	description	Variation. One specimen from Sulawesi has a darker head and pronotum and slightly darker elytra than the other specimens examined. Geographical Distribution (Fig. 65). Indonesia (Sulawesi). Natural History. According to specimen label data and Arrow (1930), H. nuciferae was found associated with young unexpanded coconut leaves.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC86E732A8DFD256DCB2DBE.taxon	description	(Figs 53, 65)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC86E732A8DFD256DCB2DBE.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ P. I., PALAWAN, Eran Pt., 8 km SW Tarumpitao Pt. 31. XII. ‘ 59 - 4. I. ’ 60 / L. W. Quate Collector / [light green label] BPBM / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus palawanensis KC & Pollock’, in BPBM. One paratype, male, same label as in holotype, in BPBM. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the type locality of this species, i. e., Palawan, Philippines.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC86E732A8DFD256DCB2DBE.taxon	diagnosis	Diagnosis. Hemipeplus palawanensis sp. nov. can be distinguished from the similar-appearing H. philippinensis sp. nov. (Fig. 49) by its relatively smaller eyes and rounded, instead of angulate, temples. Hemipeplus palawanensis sp. nov. has the following diagnostic features: temples short and rounded; scape submoniliform; eyes moderately large (as long as scape + pedicel), moderately wide and moderately convex; pronotum subquadriform; anterior emargination distinct, moderately deep in the center; anterior half only slightly wider than posterior half; pronotal pad narrow but distinct; anterior pronotal angles rounded, not much produced; posterior angles rectangular, rounded; pronotal lobe short with median notch; body long, flat; color dull, uniformly rufotestaceous; distribution: Philippines (Palawan).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC86E732A8DFD256DCB2DBE.taxon	description	Description. Measurements (in mm) — GHW: 0.57 – 0.61; HL: 0.47 – 0.53; AL: 0.77 – 0.92; GPW: 0.60 – 0.67; PL: 0.48 – 0.53; GEW: 0.77 – 0.80; EL: 2.42 – 2.78; TL: 3.37 – 3.84. Head (Fig. 53 B) wide (GHW / HL: 1.15 – 1.21); eyes moderately large (as long as scape + pedicel), shallow, and less convex; temples short and rounded; occiput with strongly raised areas; antennae shorter than head and pronotum length combined; antennal scape submoniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV small, moniliform; V moniliform, larger; VI larger, moniliform, apically dilated; VII – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 53 B) wide (GPW / PL: 1.25 – 1.26), subquadriform with flat disc; anterior margin with distinct emargination in the center; anterolateral angles rounded, slightly produced; lateral margins weakly sinuate; posterior half slightly more contracted than anterior half; posterolateral angles rectangular, rounded; pronotal lobe short with median notch; pronotal pad (Fig. 53 C) narrow, but distinct; pronotal pits shallow without prominent grooves. Scutellar shield elliptical, transverse, not twice as wide as long. Elytra (Fig. 53 A) long (EL / GEW: 3.14 – 3.48) with flat disc; vestiture short and dense; apical patch long but faint. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges narrowly separated but slightly wider than in sympatric H. philippinensis sp. nov.; mesosternal process narrow and linear. Color dull, uniformly rufotestaceous. Male Genitalia (Figs 53 D – F). (Tegmen length = 0.28 mm (n = 1 )) Lobe of basale short and blunt, apically bulged; shoulders obtusely angled with moderate slant; apicale and basale with distinct suture; parameres arcuate at the apices; penis slightly longer than tegmen. Geographical Distribution (Fig. 65). Philippines (Palawan). FIGURE 53. Hemipeplus palawanensis sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCE6E762A8DFF006B6E2CF5.taxon	description	(Figs 54.1, 54.2, 65) FIGURE 54.1. Hemipeplus sinensis sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. FIGURE 54. 2. Hemipeplus sinensis sp. nov. (Allotype) A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCE6E762A8DFF006B6E2CF5.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ CHINA, E GUIZHOU prov. FODINGSHAN, Ganshi 25 km S of Shiquian, 1300 m 5 – 9 Jun 1997, Bolm lgt. / [green label] ♂ / [orange label] BASEL / [red label] HOLOTYPE ♂ Hemipeplus sinensis KC & Pollock’, in NHMB. Allotype, female, labeled: ‘ CHINA: Hainan isl. [MF 17 a] Limushan Mts., 3 km NNW of first administr center, at a dam 19 ° 10 ’ N 109 ° 44 ’ E [at light] 560 m; 6. v. 2011. Fikáček lgt. / coll. general National Museum Prague, Czech Republic’, in NMP 6 V. Two paratypes: 1 specimen, male, labeled: ‘ SICHUAN, 4 – 20. v. 1989. 29 ° 34 ’ N 103 ° 24 ’ E, EMEI mt. 1000 m, Vít Kubáň leg. / coll. general National Museum Prague, Czech Republic’, in NMP 6 V; 1 specimen, female, labeled: ‘ FUKIEN [Fujian], S. China Shaowu: KiHsienKai T. C. Maa / X. 1944 ’, in BPBM. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the species’ type locality, i. e., China.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCE6E762A8DFF006B6E2CF5.taxon	diagnosis	Diagnosis. Hemipeplus sinensis sp. nov. can be distinguished from the similar-appearing H. labuanensis sp. nov. (Fig. 55) by its relatively more produced anterolateral pronotal angles, narrower pronotal pad, and inner margins of parameres without prominent expansions. Hemipeplus sinensis sp. nov. has the following diagnostic features: temples short and angulate; scape moniliform; eyes moderately large (just as long as scape + pedicel), shallow and less convex to moderately wide and moderately convex; pronotum subcordiform; anterior emargination broad and distinct; anterior half wider than posterior half; pronotal pad distinct; anterior pronotal angles rounded, projected rather upward than sideward; posterior angles obtuse, rounded; pronotal lobe almost absent without median notch; body long; elytral disc slightly convex; color dull, almost uniformly testaceous; distribution: China (Fukien, Guizhou, Hainan, Sichuan).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCE6E762A8DFF006B6E2CF5.taxon	description	Description. Measurements (in mm) — GHW: 0.55 – 0.67; HL: 0.53 – 0.62; AL: 0.93 – 1.14; GPW: 0.58 – 0.77; PL: 0.48 – 0.55; GEW: 0.83 – 0.97; EL: 2.68 – 2.93; TL: 3.69 – 4.10. Head (Figs 54.1 B, 54.2 B) wide (GHW / HL: 1.04 – 1.08); eyes moderately large (just as long as scape + pedicel), shallow and less convex to moderately wide and moderately convex; temples short and angulate; occiput with strongly raised areas; antennae shorter than head and pronotum length combined; antennal scape moniliform; pedicel small moniliform; antennomere III more elongate and apically dilated; IV smaller, moniliform to submoniliform; V slightly larger, moniliform to submoniliform, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth very small. Pronotum (Figs 54.1 B, 54.2 B) wide (GPW / PL: 1.21 – 1.40), subcordiform; anterior margin with broad and distinct emargination; anterolateral angles rounded, produced upward instead of sideward; lateral margins moderately sinuate; posterior half contracted; posterolateral angles obtuse, rounded; pronotal lobe almost absent without median notch; pronotal pad (Figs 54.1 C, 54.2 C) distinct; pronotal pits shallow without proper grooves to deep with moderately developed grooves. Scutellar shield subhexagonal, flat posterior margin, transverse, about twice as wide as long. Elytra (Figs 54.1 A, 54.2 A) long (EL / GEW: 3.02 – 3.23) with slightly convex disc; vestiture moderately long and dense; apical patch very faint. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges well separated; mesosternal process narrow and linear. Color dull, uniformly brown; ventrally palpi paler. Male Genitalia (Figs 54. 1 D – F). (Tegmen length = 0.25 mm (n = 1 )) Lobe of basale very short and blunt; shoulders obtusely angled, slanted; apicale and basale with distinct suture; parameres very short, slightly arcuate at the apices; penis much longer than tegmen, basally wide. Variation. Females (one from Hainan (Fig. 54.2) and one from Fukien) have much wider and flatter pronota than males. It is possible that the specimens with wider pronotum may represent a different species; this might be elucidated using DNA analysis.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCE6E762A8DFF006B6E2CF5.taxon	discussion	Remarks. Among its Oriental congeners, H. sinensis sp. nov. most closely resembles H. labuanensis sp. nov. (Malaysia) in its external appearance owing to anterolateral margins of pronotum produced upward. Geographical Distribution (Fig. 65). China (Fukien, Guizhou, Hainan, Sichuan). Hemipeplus sinensis sp. nov. appears to be widespread in China.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCC6E782A8DF8B168432F10.taxon	description	(Figs 55, 65)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCC6E782A8DF8B168432F10.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ BRITISH N. BORNEO Labuan Island Nov. 28 – 29, 1959 [last digit corrected in ink] / T. C. Maa Collector BISHOP / [green label] ♂ / [light green label] BPBM / [red label] HOLOTYPE ♂ Hemipeplus labuanensis KC & Pollock’, in BPBM. Allotype, female, labeled: ‘ BORNEO (Brit. N) Labuan I., 15 m. Oct. 24, 1957 / Nr. air strip / grass / J. L. Gressitt Collector BISHOP Museum’, in BPBM. One paratype, female, labeled same as allotype, in BPBM. FIGURE 55. Hemipeplus labuanensis sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar). Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the species’ type locality, i. e., Labuan, Malaysia.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCC6E782A8DF8B168432F10.taxon	diagnosis	Diagnosis. Hemipeplus labuanensis sp. nov. can be distinguished from the similar-appearing H. sinensis sp. nov. (Fig. 54.1, 54.2) by its less produced anterolateral pronotal angles, wider pronotal pad, and inner margins of parameres with prominent expansions. Hemipeplus labuanensis sp. nov. has the following diagnostic features: temples tiny and roundish; scape moniliform, apically dilated; eyes moderately large (just as long as scape + pedicel), moderately wide and moderately convex; pronotum subcordiform; anterior emargination broad, deep, and distinct; anterior half wider than posterior half; pronotal pad wide and distinct; anterior pronotal angles rounded, produced rather upward than sideward; posterior angles obtuse, rounded; pronotal lobe almost absent without median notch; body long; elytra slightly convex; color dull, uniformly testaceous; distribution: Malaysia (Labuan).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFCC6E782A8DF8B168432F10.taxon	description	Description. Measurements (in mm) — GHW: 0.55 – 0.62; HL: 0.53 – 0.58; AL: 0.86; GPW: 0.63 – 0.68; PL: 0.53 – 0.57; GEW: 0.76 – 0.87; EL: 2.80 – 3.15; TL: 3.86 – 4.30. Head (Fig. 55 B) wide (GHW / HL: 1.04 – 1.07); eyes moderately large (just as long as scape + pedicel), moderately wide and moderately convex; temples very short and roundish; occiput with strongly raised areas; antennae shorter than head and pronotum length combined; antennal scape moniliform, apically dilated and externally produced on one side; pedicel small moniliform; antennomere III more elongate and apically dilated; IV smaller, moniliform; V slightly larger, moniliform, apically dilated; VI smaller, moniliform, apically dilated; VII – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 55 B) wide (GPW / PL: 1.19), subcordiform; anterior margin with broad, deep, and distinct emargination; anterolateral angles rounded, produced upright; lateral margins moderately sinuate; posterior half contracted; posterolateral angles obtuse to square, rounded; pronotal lobe almost absent without median notch; pronotal pad (Fig. 55 C) wide and distinctly setose; pronotal pits shallow without proper grooves. Scutellar shield subhexagonal, transverse, less than twice as wide as long. Elytra (Fig. 55 A) long (EL / GEW: 3.62 – 3.68), slightly convex; vestiture moderately long and dense; apical patch short and broad to moderately long, faint. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges well separated; mesosternal process narrow and linear. Color dull, uniformly testaceous. Male Genitalia (Figs 55 D – F). (Tegmen length = ca 0.25 mm, not measured) Lobe of basale moderately long, slender; shoulders obtusely angled, moderately slanted; apicale and basale with distinct suture; parameres very short, slightly arcuate at the apices, tips sharp; inner margins of parameres with prominent expansions; penis wide, longer than tegmen. Geographical Distribution (Fig. 65). Malaysia (Labuan). Natural History. According to specimen label data, H. labuanensis sp. nov. is associated with grasses (Poaceae).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC26E7A2A8DFA156AC22E31.taxon	description	(Figs 56, 65)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC26E7A2A8DFA156AC22E31.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ Philippinen Mindanao, Surigao leg. Böttcher [underside of label] 30.8. 1916 / Zool. Mus. Berlin / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus nigerrimus KC & Pollock’, in ZMHB. Two paratypes: 1 specimen, sex unknown, labeled: ‘ Philippinen Mindanao, Surigao leg. Böttcher [underside of label] 30.8.1916 / klematanicus Gestro [handwritten in red ink] ’, in ZMHB; 1 specimen, sex unknown, labeled: ‘ Philippinen Mindanao, Surigao leg. Böttcher [underside of label] 28.8.1916 ’, in ZMHB. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the holotype’s darker body color, distinguishing it from H. klematanicus and prompting a detailed examination of these darker specimens.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC26E7A2A8DFA156AC22E31.taxon	diagnosis	Diagnosis. Hemipeplus nigerrimus sp. nov. can be distinguished from the similar-appearing H. klematanicus (Fig. 46) by its relatively smaller eyes, less acutely pointed posterior pronotal angles, and darker color. Hemipeplus nigerrimus sp. nov. has the following diagnostic features: temples short and angulate; scape moniliform, apically dilated; eyes moderately large (as long as scape + pedicel), moderately wide and moderately convex; pronotum subcordiform; anterior emargination distinct; anterior half slightly wider than posterior half; pronotal pad narrow, moderately distinct; anterior pronotal angles rounded, posterior angles slightly obtuse to square, more angled than rounded (never pointed acutely as in H. klematanicus); pronotal lobe indistinct with faint median notch; body long; elytra convex; color dull, uniformly rufopiceous to darker head and pronotum; distribution: Philippines (Mindanao). FIGURE 56. Hemipeplus nigerrimus sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view with spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC26E7A2A8DFA156AC22E31.taxon	description	Description. Measurements (in mm) — GHW: 0.50 – 0.56; HL: 0.45 – 0.52; AL: 0.85 – 1.01; GPW: 0.55 – 0.63; PL: 0.47 – 0.52; GEW: 0.60 – 0.66; EL: 2.63 – 2.96; TL: 3.55 – 4.00. Head (Fig. 56 B) wide (GHW / HL: 1.08 – 1.11); eyes moderately large (as long as scape + pedicel), moderately wide and moderately convex; temples short and angulate; occiput with strongly raised areas; antennae about subequal to head and pronotum length combined; antennal scape moniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV shorter, moniliform; V – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 56 B) wide (GPW / PL: 1.17 – 1.21); anterior margin with distinct emargination; anterolateral angles rounded, produced; lateral margins sinuate; posterior half much more contracted than anterior half; posterolateral angles slightly obtuse to square, more angled than rounded but not pointed (Fig. 56 B); pronotal lobe indistinct with faint median notch; pronotal pad (Fig. 56 C) narrow, moderately distinct; pronotal pits moderately deep with moderately developed grooves. Scutellar shield elliptical, transverse, less than twice as wide as long. Elytra (Fig. 56 A) long (EL / GEW: 4.38 – 4.48), convex; vestiture short and dense; apical patch broad and slightly darker than elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges narrowly separated; mesosternal process narrow and linear. Color dull, uniformly rufopiceous to head and pronotum darker than other dorsal parts. Male Genitalia (Figs 56 D – F). (Tegmen length = 0.35 mm (n = 1 )) This is the only species in the Austro-Oriental region in which the apices of parameres are not arcuate or bulged, and are straight; lobe of basale long, slender with blunt apex; shoulders obtusely angled with moderate slant; apicale and basale with distinct suture; parameres with bases thicker than apices; apices straight without distinct curves; penis much longer than tegmen, medially widened. Variation. Specimens vary in the intensity of darkness of the body color.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC26E7A2A8DFA156AC22E31.taxon	discussion	Remarks. Hemipeplus nigerrimus sp. nov. is similar to H. klematanicus (Fig. 46) (particularly the rufopiceous forms of H. klematanicus); however, the genitalia are distinct between the two species (Figs 46 D – F, 56 D – F). Geographical Distribution (Fig. 65). Philippines (Mindanao).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC06E7C2A8DFBF5687C28D1.taxon	description	(Figs 57, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC06E7C2A8DFBF5687C28D1.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ NEW BRITAIN Keravat 135 m. Nov. 20 – 25, ‘ 59 / T. C. Maa Collector / [light green label] BPBM / [dark green label] ♂ / [red label] HOLOTYPE Hemipeplus craigi ♀ [typo for ♂] KC & Pollock’, in BPBM. Allotype, female, labeled: ‘ NEW BRITAIN Gazelle Pen Kerawat 60 m. IX- 11 - ’ 55 / J. L. Gressitt Collector’, in BPBM. Nine paratypes: 1 specimen, sex, unknown, labeled: ‘ NEW BRITAIN: Gazelle Pen Kerawat 60 m. 29. viii. 1955, Gressitt leg. ’, in BPBM; 1 specimen, female, labeled: ‘ NEW GUINEA: (NE) Bulolo 1005 m. Aug. 25, 1956 / E. J. Ford, Jr. Collector’, in BPBM; 3 specimens, 1 male, 2 female, labeled: ‘ NEW IRELAND (SW) Ridge above “ Camp Bishop ” 15 km up Kait R. 250 - 500 m. VI- 11 - 1956 / J. L. Gressitt Collector’, in BPBM; 1 specimen, female, labeled: ‘ NEW IRELAND (SW) Ridge above “ Camp Bishop ” 15 km. up Kait R. 250 - 450 m. VII- 9 - 1956, / J. L. Gressitt Collector’, in BPBM; 1 specimen, sex unknown, labeled: ‘ NEW BRITAIN Gazelle Pen. Bainings: St. Paul’s 350 m. Sept. 8. 1955 / J. L. Gressitt Collector / definitely not nuciferae’, in BPBM; 1 specimen, sex unknown, labeled: ‘ PNG. Keravat, New Britian, VII. X. 1973 J. Stibick / temples Det. D. A. Pollock’, in CNCI; 1 specimen, sex unknown, labeled: ‘ DUTCH NEW GUINEA: Cyclops Mts., Sabron. 930 ft. vi. 1936. L. E. Cheesman. B. M. 1936 - 271. / Hemipeplus’, in BMNH. Derivation of Specific Epithet. The specific name (noun in the genitive case) is given in allusion to the first author’s longtime friend from the USA, Craig Winzelberg.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC06E7C2A8DFBF5687C28D1.taxon	diagnosis	Diagnosis. Hemipeplus craigi sp. nov. can be distinguished from the similar-appearing H. saymyname sp. nov. (Fig. 58) by its uniformly rufous color and parameres of male genitalia much broader, penis and lobe of basale shorter. Hemipeplus craigi sp. nov. has the following diagnostic features: temples short and roundish to angulate; scape submoniliform; eyes moderately large (just as long as scape + pedicel), moderately wide and moderately convex; pronotum subcordiform; anterior emargination moderate; anterior half wider than posterior half; pronotal pad narrow but distinct; anterior pronotal angles rounded and produced; posterior angles square, rounded; pronotal lobe very slight with median notch; body long; elytra slightly convex; color dull, uniformly rufous; distribution: Indonesia (Papua: Jayapura), Papua New Guinea (East Papua: Bulolo; New Britain). FIGURE 57. Hemipeplus craigi sp. nov. (Holotype; Figs E, F from paratype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax with spiculum gastrale; D. Male genitalia dorsal view; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC06E7C2A8DFBF5687C28D1.taxon	description	Description. Measurements (in mm) — GHW: 0.58 – 0.66; HL: 0.56 – 0.60; AL: 0.98 – 1.08; GPW: 0.58 – 0.72; PL: 0.50 – 0.58; GEW: 0.73 – 0.82; EL: 2.58 – 3.43; TL: 3.64 – 4.61. Head (Fig. 57 B) wide (GHW / HL: 1.04 – 1.10); eyes moderately large (just as long as scape + pedicel), moderately wide and moderately convex; temples short and roundish; occiput with raised areas; antennae about subequal to or shorter than head and pronotum length combined; antennal scape submoniliform, apically dilated and externally produced on one side; pedicel small moniliform; antennomere III more elongate and apically dilated; IV smaller, submoniliform; V more elongate, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 57 B) wide (GPW / PL: 1.16 – 1.24), subcordiform; anterior margin with moderate emargination; anterolateral angles rounded, produced; lateral margins sinuate; posterior half contracted; posterolateral angles obtuse to square, rounded; pronotal lobe very faint with median notch; pronotal pad (Fig. 57 C) narrow, but well distinct; pronotal pits moderately deep with moderate grooves. Scutellar shield transverse, subhexagonal, about twice as wide as long. Elytra (Fig. 57 A) long (EL / GEW: 3.53 – 4.18), slightly convex; vestiture moderately long and dense; apical patch long but very faint. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges narrowly to well separated; mesosternal process narrow and linear. Color dull, uniformly rufous; ventrally uniformly rufotestaceous with palpi lighter to uniformly rufous. Male Genitalia (Figs 57 D – F). (Tegmen length = 0.32 mm (n = 1 )) Lobe of basale short and moderately slender; shoulders obtuse, rounded, moderately slanted; apicale and basale with distinct suture; parameres slightly arcuate near the apices, tips sharp and moderately pointed; penis slightly longer than tegmen. Geographical Distribution (Fig. 64). Indonesia (Papua: Jayapura), Papua New Guinea (East Papua: Bulolo; New Britain).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC66E7E2A8DFCD56A092B45.taxon	description	(Figs 58, 65)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC66E7E2A8DFCD56A092B45.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ [E. Malaysia] 16 miles point from Keningau Sabah, Is. Borneo 9. iii. 1993, T Ueno leg. / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus saymyname KC & Pollock’, in BPBM. Ten paratypes: 9 specimens, 1 female, 8 sex unknown, labeled same as holotype, in DAPC; 1 specimen, male, labeled: ‘ SABAH: Mt. Kinabalu 1550 - 1650 m, 23. V. 1987 Burckhardt-Löbl’, in MHNG. Derivation of Specific Epithet. The specific name (noun in apposition) is given in allusion to the iconic phrase ‘ Say my name’ from the popular TV series ‘ Breaking Bad’.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC66E7E2A8DFCD56A092B45.taxon	diagnosis	Diagnosis. Hemipeplus saymyname sp. nov. can be distinguished from the similar-appearing H. craigi sp. nov. (Fig. 57) by its elytra usually suffused in light piceous shade, parameres of male genitalia much thinner, penis and lobe of basale longer. Hemipeplus saymyname sp. nov. has the following diagnostic features: temples short and roundish; scape short and moniliform; eyes moderately large (just as long as scape + pedicel), moderately wide and moderately convex; pronotum subcordiform; anterior emargination moderate but distinct; anterior half wider than posterior half; pronotal pad moderately present; anterior pronotal angles rounded and produced; posterior angles obtuse, rounded; pronotal lobe indistinct with faint median notch; body long; elytra slightly convex; color dull, rufous with most of the elytra suffused in piceous shade to uniformly rufotestaceous; distribution: Malaysia (Sabah).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC66E7E2A8DFCD56A092B45.taxon	description	Description. Measurements (in mm) — GHW: 0.55 – 0.62; HL: 0.52 – 0.60; AL: 0.85 – 0.95; GPW: 0.55 – 0.63; PL: 0.43 – 0.53; GEW: 0.70 – 0.87; EL: 2.70 – 3.34; TL: 3.65 – 4.47. Head (Fig. 58 B) wide (GHW / HL: 1.03 – 1.06); eyes moderately large (just as long as scape + pedicel), moderately wide and moderately convex; temples short and roundish; occiput with strongly raised areas; antennae shorter than head and pronotum length combined; antennal scape short, moniliform, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV small, moniliform; V larger, moniliform, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 58 B) wide (GPW / PL: 1.19 – 1.28), subcordiform; anterior margin with moderate emargination; anterolateral angles rounded, produced; lateral margins sinuate; posterior half contracted; posterolateral angles obtuse to square, rounded; pronotal lobe very faint with a hint of median notch; pronotal pad (Fig. 58 C) narrow, but distinct; pronotal pits shallow without grooves. Scutellar shield transverse, subhexagonal, about twice as wide as long. Elytra (Fig. 58 A) long (EL / GEW: 3.84 – 3.86), slightly convex; vestiture moderately long and dense; apical patch long and piceous, appears faint when in piceous elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges very narrowly separated to less so; mesosternal process narrow and linear. Color rufous with most of the elytra usually suffused in piceous shade; antennomeres III – basal XI usually darker; color uniformly rufotestaceous in the specimen from Mt. Kinabalu; ventrally inner edges of prosternum and hypomeral edges darker; abdominal sternites slightly darker; rest is uniformly rufous – rufotestaceous; palpi and legs lighter in some specimens. FIGURE 58. Hemipeplus saymyname sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. Male Genitalia (Figs 58 D – F). (Tegmen length = 0.28 mm) Lobe of basale long and slender; shoulders obtusely angled, moderately slanted; apicale and basale with distinct suture; parameres slender, slightly arcuate near the apices, tips not sharp; penis wide, much longer than tegmen with narrowed apex. Variation. As given in the description above, one specimen of H. saymyname sp. nov. has a uniformly brown body while all others have elytra with a piceous shade. Geographical Distribution (Fig. 65). Malaysia (Sabah).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC46E7F2A8DFE216B8B2DF3.taxon	description	(Figs 59, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC46E7F2A8DFE216B8B2DF3.taxon	materials_examined	Type Material. Holotype, female, labeled: ‘ Irian Jaya: Jayawi-jaya, Okloma-Emdoman, 28.9.1993 1400 – 1700 m leg. A: RIEDEL / SMNS / [blue label] ♀ / [red label] HOLOTYPE ♀ Hemipeplus bromineus KC & Pollock’, in SMNS. One paratype, female, labeled same as holotype, in SMNS. Derivation of Specific Epithet. The specific name (adjective in the nominative case, in gender agreement with substantive) is given in allusion to the species’ bromine-like color.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC46E7F2A8DFE216B8B2DF3.taxon	diagnosis	Diagnosis. Hemipeplus bromineus sp. nov. can be distinguished from the similar-appearing obscure form of H. heisenbergi sp. nov. (Fig. 60.3) by its barely emarginate anterior pronotal margin, lack of notch on its indistinct pronotal lobe, and more distinct pronotal pad. Hemipeplus bromineus sp. nov. also resembles H. rugosus sp. nov. (Fig. 42) which has larger temples, less prominent pronotal pad, and much deeper emargination on the anterior pronotal margin. Hemipeplus bromineus sp. nov. has the following diagnostic features: temples very short and roundish but prominent in contrast to its small eyes; scape submoniliform, slightly elongate; eyes small (smaller than scape + pedicel), shallow and less convex; pronotum subcordiform; anterior emargination shallow, not prominent; anterior half wider than posterior half; pronotal pad narrow but distinct; anterior pronotal angles rounded, slightly produced; posterior angles square, rounded; pronotal lobe indistinct without median notch; body long; color dull, rufopiceous / bromine-brown, some parts slightly lighter than others (more piceous on head, pronotum, suture and apices of elytra); distribution: Indonesia (Papua).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFFC46E7F2A8DFE216B8B2DF3.taxon	description	Description. Measurements (in mm) — GHW: 0.62 – 0.63; HL: 0.62 – 0.63; AL: 1.10 – 1.12; GPW: 0.63 – 0.64; PL: 0.55 – 0.55; GEW: 0.85; EL: 3.28 – 3.40; TL: 4.45 – 4.58. Head (Fig. 59 B) as long as wide (GHW / HL: 1.00) eyes small (smaller than scape + pedicel), shallow and less convex; temples very short and roundish; occiput with strongly raised areas; antennae shorter than head and pronotum length combined; antennal scape submoniliform, slightly elongate, apically dilated; pedicel small moniliform; antennomere III more elongate and apically dilated; IV smaller, submoniliform; V – VI larger, submoniliform, apically dilated; VII – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth not observable in the specimens. Pronotum (Fig. 59 B) wide (GPW / PL: 1.15 – 1.16), subcordiform; anterior margin with slight emargination; anterolateral angles rounded, moderately produced; posterior half contracted, not produced; posterolateral margins subparallel; posterolateral angles obtuse to square, rounded; pronotal lobe almost absent without median notch; pronotal pad (Fig. 59 C) narrow, but distinct; pronotal pits shallow without proper grooves. Scutellar shield subhexagonal, transverse, about twice as wide as long. Elytra (Fig. 59 A) long (EL / GEW: 3.86 – 4.00) with disc slightly convex; vestiture moderately long and dense; apical patch long and slightly darker than the elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges well separated; mesosternal process narrow and linear. Color dull, rufopiceous / bromine-brown (more piceous on head, pronotum, suture of elytra and apices); ventrally palpi and tarsi paler. Geographical Distribution (Fig. 64). Indonesia (Papua). FIGURE 59. Hemipeplus bromineus sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF3A6E832A8DFF006D7C2B8D.taxon	description	(Figs 60.1, 60.2, 64)	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF3A6E832A8DFF006D7C2B8D.taxon	materials_examined	Type Material. Holotype, male, labeled: ‘ NEW GUINEA (NE) Nondugl 1600 m. July 9, 1955 / J. L. Gressitt Collector / [light green label] BPBM / [green label] ♂ / [red label] HOLOTYPE ♂ Hemipeplus heisenbergi KC & Pollock’, in BPBM. Allotype: female, labeled: ‘ NEW GUINEA: NE. Moife, 2100 m. 15 km NW of Okapa Oct. 11 - 13, 1959 / T. C. Maa Collector / Swept in clearing’, in BPBM. Twenty-four paratypes; 2 specimens, 1 male, 1 female, labeled same as holotype, in BPBM; 1 specimen, sex unknown, labeled same as allotype, in BPBM; 1 specimen, male, labeled: ‘ WEST NEW GUINEA Bokondini, 40 km N of Baliem Val. ca. 1300 m 16 - 23. XI. 1961 / L. W. Quate Collector’, in BPBM; 1 specimen, male, labeled: ‘ NEW GUINEA (NE) Goroka, 1550 m. July 9 [corrected in ink], 1955 / J. L. Gressitt Collector’, in BPBM; 1 specimen, male, labeled: ‘ NEW GUINEA (NE) Goroka, 1550 m. July 19, 1955 / J. L. Gressitt Collector / ON SUGARCANE’, in BPBM; 4 specimens, 3 females, 1 sex unknown, labeled: ‘ NEW GUINEA (NE) Goroka, 1550 m. July 25, 1955, / J. L. Gressitt Collector / Saccharin [for one] ’, in BPBM; 1 specimen, male, labeled: ‘ NEW GUINEA (NE) Goroka, 1550 m. July 26, 1955 / J. L. Gressitt Collector / on sugarcane’, in BPBM; 1 specimen, male, labeled: ‘ NEW GUINEA: NE Okapa, Purosa 1700 - 2000 m, 18. I. 1966 / J. Sedlacek Collector BISHOP MUS. ’, in BPBM; 1 specimen, sex unknown, labeled: ‘ New Guinea (NE) Mt. Piora 6 ° 45 ’ S 146 ° E 1900 m, 12. VI. 6 6 / 6460 / J. L. Gressitt Collector’, in BPBM; 1 specimen, sex unknown, labeled: ‘ NEW GUINEA (NE) Miramar, Asano V: 1800 m. VI- 27 - ’ 55 / J. L. Gressitt Collector’, in BPBM; 1 specimen, sex unknown, labeled: ‘ NEW GUINEA: NE. Tapo (= Tapu) 1650 m. 3 km NW of Kainantu, X- 22 - 1959 / T. C. Maa Collector’, in BPBM; 1 specimen, sex unknown, labeled: ‘ Palmer R. at Black R. Papua June 23 - 30,1936. Archbold Exped. ’, in AMNH; 1 specimen, sex unknown, labeled: ‘ NEW GUINEA (NE): Mt. Hagen area 1650 m., VI- 28 - ’ 57 / D. Elmo Hardy Collector Bishop Museum’, in BPBM; 1 specimen, sex unknown, labeled: ‘ NE NEW GUINEA: Akivitana Riv. 1550 m., 1 [not sure about the first digit] 0. I. 65 / J. & M. Sedlacek Collectors BISHOP MUSEUM’, in BPBM; 1 specimen, female, labeled: ‘ PNG / EH Prov. Umg. Kainantu Onerunka VI. 79 [written on top in ink] ’, in MHNG; 1 specimen, male, labeled: ‘ PNG / EH Prov. Umg. Kainantu Onerunka, 22 IV 79 [written on top in ink] ’, in MHNG; 1 specimen, female, labeled: ‘ NEW GUINEA / NE / Baiyer River Sanctuary, 1 - 5. IX. 1969. / / No. NGB-U. / leg. Dr. J. Balogh / Hemipeplus sp. D. G. H. Halstead det., 1977 / Hemipeplus sp. det. Halstead 1977 ’, in HNHM; 1 specimen, female, labeled: ‘ [something illegible] / J. & M. Sedlacek Collectors BISHOP MUSEUM’, in BPBM; 1 specimen, sex unknown, labeled: ‘ NEW GUINEA (Neth.) Wisselmeren: Kamo-Dobei div. 1700 m. Aug. 13. 1955. / J. L. Gressitt Collector’, in BPBM; 1 specimen, sex unknown, labeled: ‘ TERR. PAPUA & NEW GUINEA [underlined in red]: Alyrua 27. ix. 1957. J. Smart / Brit-Mus. 1957 - 693 / [with a barcode] NHMUK 015009784 ’, in BMNH. Derivation of Specific Epithet. The specific name (noun in the genitive case) is given in allusion to Walter White’s alias ‘ Heisenberg’, the iconic protagonist of the acclaimed TV series ‘ Breaking Bad’.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF3A6E832A8DFF006D7C2B8D.taxon	diagnosis	Diagnosis. Hemipeplus heisenbergi sp. nov. can be distinguished from the similar-appearing H. bromineus sp. nov. (Fig. 59) by its distinctly emarginate anterior pronotal margin and less prominent pronotal pad. Hemipeplus heisenbergi sp. nov. also slightly resembles H. miyamotoi (Fig. 43.1) from Japan and Taiwan, but its eyes are relatively larger and temples, smaller than the latter. Hemipeplus heisenbergi sp. nov. has the following diagnostic features: temples very short and roundish but prominent in contrast to its small eyes; scape moniliform to submoniliform (slightly elongate in some specimens), apically dilated; eyes small (only slightly longer than the scape), shallow and less convex; pronotum subcordiform; anterior emargination deep, and prominent; anterior half wider than posterior half; pronotal pad not very prominent; anterior pronotal angles rounded, produced; posterior angles square, rounded; pronotal lobe indistinct, but with median notch; body long; color dull, rufous to rufopiceous, head and pronotum darker; distribution: New Guinea.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF3A6E832A8DFF006D7C2B8D.taxon	description	Description. Measurements (in mm) — GHW: 0.66 – 0.73; HL: 0.60 – 0.70; AL: 1.12 – 1.30; GPW: 0.72 – 0.89; PL: 0.57 – 0.68; GEW: 0.87 – 1.03; EL: 3.00 – 3.66; TL: 4.17 – 5.04. Head (Figs 60.1 B, 60.2 B) wide (GHW / HL: 1.04 – 1.10) eyes small (only slightly longer than the scape), shallow and less convex; temples very short and roundish; occiput with strongly raised areas; antennae slightly shorter or about subequal to head and pronotum length combined; antennal scape moniliform to submoniliform, apically dilated, externally produced on one side; pedicel small moniliform; antennomere III more elongate and apically dilated; IV smaller, submoniliform; V larger, submoniliform, apically dilated; VI – X subtriangular or bowl shaped, FIGURE 60. 1. Hemipeplus heisenbergi sp. nov. (Holotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Figs 60.1 B, 60.2 B) wide (GPW / PL: 1.26 – 1.30), subcordiform; anterior margin with deep emargination; anterolateral angles rounded, produced; lateral margins sinuate; posterior half contracted; posterolateral angles slightly obtuse, rounded; pronotal lobe almost absent but with median notch; pronotal pad (Figs 60.1 C, 60.2 C) not very distinct; pronotal pits deep to moderately deep with moderately developed grooves. Scutellar shield transverse, subpentagonal, twice to less than twice as wide as long. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges narrowly to well separated; mesosternal process narrow and linear. Elytra (Figs 60.1 A, 60.2 A) long (EL / GEW: 3.45 – 3.55) with almost flat disc; vestiture moderately short and dense; apical patch long and wide, rufous to piceous, less visible when the elytra are piceous. Color dull, rufous (Fig. 60.1) to rufopiceous (Fig. 60.2), head and pronotum darker; ventrally variable, uniformly rufotestaceous to rufopiceous with edges of pro- and mesosternum piceous; some specimens ventrally rufous with inner edges of prosternum darker; palpi, glossal sclerite, galea, and tarsi paler in some, not in others. FIGURE 60. 2. Hemipeplus heisenbergi sp. nov. (Allotype). A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax. Male Genitalia (Figs 60. 1 D – F). (Tegmen length = 0.33 mm (n = 1 )) Lobe of basale thick, short, and wide with dilated apex; shoulders obtuse and slanted; apicale and basale with distinct suture; parameres entirely arcuate, tips moderately sharp; penis longer than tegmen with pointed apex. Variation. Apart from color variation, some specimens have a much narrower pronotum, more prominent pronotal pad, and slightly larger eyes. The hypomeral edges are more narrowly separated in some specimens than others. Geographical Distribution (Fig. 64). Widespread in New Guinea from west to east. Natural History. According to specimen label data, H. heisenbergi sp. nov. is associated with sugarcane (Poaceae). The allotype label reads that it was collected by sweeping in a clearing.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF396E872A8DFE696ACF2D84.taxon	materials_examined	Material Examined. 1 specimen, male, labeled: ‘ INDONESIA: E. New Guinea, Papua Prov., Star Mountains, Oksibil N, 4 ° 54 ’ 01 ’’ S, 140 ° 38 ’ 48 ’’ E, 1350 - 1380 m, 10. III. 2018, gardens & secondary lower montane rainforest on limestone, leg. D. Telnov’, in BMNH (Fig. 60.3).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF396E872A8DFE696ACF2D84.taxon	diagnosis	Diagnosis. Hemipeplus heisenbergi sp. nov. “ obscure form ” can be distinguished from the typical form of H. heisenbergi by its slenderer appearance. Hemipeplus heisenbergi sp. nov. “ obscure form ” has the following diagnostic features: temples very short and roundish but prominent in contrast to its small eyes; scape moniliform, apically dilated; eyes small (only slightly longer than the scape), shallow and less convex; pronotum subcordiform; anterior emargination deep, and prominent; anterior half wider than posterior half; pronotal pad indistinct; pronotal angles rounded; pronotal lobe indistinct, but with median notch; body long; elytra slightly convex; color dull, uniformly piceous with appendages rufopiceous; distribution: New Guinea (Papua: Star Mountains).	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF396E872A8DFE696ACF2D84.taxon	description	Description. Measurements (in mm) — GHW: 0.65; HL: 0.63; AL: 1.20; GPW: 0.65; PL: 0.57; GEW: 0.87; EL: 3.17; TL: 4.37. Head (Fig. 60.3 B) wide (GHW / HL: 1.03) eyes small (only slightly longer than the scape), shallow and less convex; temples very short and roundish; occiput with strongly raised areas; antennae subequal to head and pronotum length combined; antennal scape moniliform, apically dilated (not externally produced on one side); pedicel small moniliform; antennomere III more elongate and apically dilated; IV smaller, submoniliform; V slightly larger, submoniliform, apically dilated; VI – X subtriangular or bowl shaped, each wider than the next; XI subpyriform with apex narrowed after the notch; lateral mandibular tooth absent. Pronotum (Fig. 60.3 B) wide (GPW / PL: 1.14), subcordiform; anterior margin with distinct emargination; anterolateral angles rounded, slightly produced; lateral margins sinuate; posterior half contracted; posterolateral angles slightly obtuse, rounded; pronotal lobe almost absent but with faint median notch; pronotal pad (Fig. 60.3 C) indistinct; pronotal pits shallow without well-developed grooves, disc very rough with wide punctures and grooves. Scutellar shield transverse, elliptical, about twice as wide as long. Elytra (Fig. 60.3 A) long (EL / GEW: 3.64), slightly convex; vestiture long and dense; apical patch present but not well visible owing to piceous elytra. Ventral surface. Prosternal process short, not extending posterior of procoxae; hypomeral edges narrowly separated; mesosternal process narrow and linear. Color dull, uniformly piceous with appendages rufopiceous. Male Genitalia (Figs 60. 3 D – F). (Tegmen length = 0.33 mm (n = 1 )) Lobe of basale short, slender with blunt apex; shoulders obtusely angled, slightly slanted; apicale and basale with distinct suture; parameres entirely arcuate, tips blunt; penis longer than tegmen with pointed apex.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
E14D87AEFF396E872A8DFE696ACF2D84.taxon	discussion	Remarks. Hemipeplus heisenbergi sp. nov. “ obscure form ” resembles typical H. heisenbergi sp. nov. in many aspects but is still morphologically distinct in terms of pronotum size, body color, punctures on body surface, and long vestiture. It is listed separately from H. heisenbergi sp. nov. because it might prove to be a distinct species if further research is done with more specimens. Alternatively, it might only be an aberration since the basic characters, such as the size of eyes, temples, and pronotal pad, are about the same. The male genitalia are similar (even the length). For now, it is treated merely as a form of H. heisenbergi sp. nov. Although not explicitly stated on the specimen data label, they appear to be sympatric since the typical form is widespread in New Guinea. Geographical Distribution (Fig. 64). Indonesia (Papua: Star Mountains). Natural History. According to specimen label data, the single specimen of H. heisenbergi sp. nov. “ obscure form ” was collected from gardens and secondary lower montane rainforest on limestone. FIGURE 60. 3. Hemipeplus heisenbergi sp. nov. “ obscure form ”. A. Dorsal habitus (with scale bar); B. Dorsal head and prothorax; C. Lateral view of head and prothorax; D. Male genitalia dorsal view (left paramere damaged) without spiculum gastrale; E. Tegmen dorsal view (with scale bar); F. Tegmen lateral view with one paramere removed. FIGURE 61. Distribution Map of North and Central American Hemipeplinae species. FIGURE 62. Distribution Map of South American Hemipeplinae species. FIGURE 63. Distribution Map of Afrotropical Hemipeplinae species. Repositories of primary types designated in this study: Neotype: H. hemipterus: FSCA Lectotype: H. australicus: BMNH; H. nuciferae: BMNH Holotypes of spp. nov.: Eurypeplus quadrimaculatus: MNHN; H. anishae: BMNH; H. antennatus: BPBM; H. belizensis: BMNH; H. brasiliensis: BMNH; H. bromineus: SMNS; H. bucculentus: BPBM; H. craigi: BPBM; H. enarotaliensis: BPBM; H. exhaustus: BPBM; H. gressitti: BPBM; H. harkoneni: ZMHB; H. heisenbergi: BPBM; H. joerebeccae: ZMHB; H. labuanensis: BPBM; H. luzonensis: BPBM; H. manusicus: BPBM; H. monteithi: UQIC; H. neoaustralicus: FSCA; H. neoguineensis: BPBM; H. nigerrimus: ZMHB; H. palawanensis: BPBM; H. philippinensis: BPBM; H. pollocki: ASU; H. pseudoklematanicus: BMNH; H. pseudoquadricollis: FSCA; H. rugosus: FSCA; H. saymyname: BPBM; H. sinensis: NHMB FIGURE 64. Distribution Map of Australian and Oceanian Hemipeplinae species. FIGURE 65. Distribution Map of Oriental Hemipeplinae species.	en	Kc, Sajan, Pollock, Darren A. (2025): Review of the Hemipeplinae (Coleoptera: Mycteridae) fauna of the world with descriptions of twenty-nine new species. Zootaxa 5574 (1): 1-140, DOI: 10.11646/zootaxa.5574.1.1, URL: https://doi.org/10.11646/zootaxa.5574.1.1
