taxonID	type	description	language	source
E13FD4141724D3154C8AFBA07CCEFA89.taxon	type_taxon	Type species. Archicupes jacobsoni Rohdendorf, 1961. Species included. Type species? A. reichardti Rohdendorf, 1961 (could be indeed a member of the genus Kaltanicupes rather than of Archicupes; Kirejtshuk, 2020), and Archicupes storozhenkoi Yan & Strelnikova, sp. nov.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141724D3164C8AFA8478E7FEC0.taxon	description	(Fig. 2)	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141724D3164C8AFA8478E7FEC0.taxon	materials_examined	Material. Holotype — PIN, No. 5890 / 8. Part and counterpart of isolated elytron, left. Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141724D3164C8AFA8478E7FEC0.taxon	diagnosis	Diagnosis. Epipleural margin with small cells, forming two parallel rows near the humerus, at midlength become a singular row. Elytron with four visible veins, vein 3 reaching at least at elytral distal fourth, vein 6 is stemming from 4 at the level of humerus, enclosing the vein 5, all reaching beyond elytral midlength. Field III is widest at the elytral base.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141724D3164C8AFA8478E7FEC0.taxon	description	Description. Elytron with maximum width at distal third, apex pointed, asymmetrical, not elongated into thin, tail-like extension. Cells rather large, rounded, organized irregularly, without maculae. Cells of different shapes, mostly round, or rounded-oval, some deformed and concave, usually smaller at the elytral base. Cuticular surface between cells glabrous. Singular row of cells between first vein and epipleural carina, field III is extended at the base and has up to 4 cells, then reduced to two rows. Number of cells varies from single to double in fields IV and V. There are 3 – 4 cells row in the field VI. Cells somewhat organize uniformly at elytron`s apical fourth. Measurements (in mm). Elytron length 5.8, width 1.8.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141724D3164C8AFA8478E7FEC0.taxon	etymology	Etymology. After prominent entomologist Sergei Yu. Storozhenko.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141724D3164C8AFA8478E7FEC0.taxon	discussion	Remarks. Species below were attributed to Permocupedidae for having more than three veins, lattice elytra cell pattern and small overall size. The specimen was attributed to the genus Archicupes for having two rows of cells in a first field and three central veins having a common base, also for the lack of vein closest to sutural margin (A 2 sensu Ponomarenko 1969). Comparison. Differs from most other species in having elytral apex not extended into tail-like structure, also in three rather than two central veins having a common base and field III being widest at the base.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141727D3164C8AFA257877F9C5.taxon	type_taxon	Type species. Cytocupes angustus Rodendorf, 1961 Species included. Type species and Cytocupes magnus Yan & Strelnikova, sp. nov.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141727D3174C8AF94D7FA9FD99.taxon	description	(Fig. 3)	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141727D3174C8AF94D7FA9FD99.taxon	materials_examined	Material. Holotype — PIN, No. 5890 / 10. Isolated elytron, left. Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141727D3174C8AF94D7FA9FD99.taxon	diagnosis	Diagnosis. Veins 4 and 5 are very closely spaced. Veins 2 and 3 are merging at the level of elytral distal 1 / 5. Veins 4 and 5 half as long as the elytron. Double rows of cells nearly touching each other’s borders, in the field III, IV and VI.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141727D3174C8AF94D7FA9FD99.taxon	description	Description. Elongated elytron, 4 times as long as wide, carinated, apex symmetrical, rounded. Epipleural and sutural margins lacks cells or punctures. Humeral bulge weakly defined. Veins 2 and 3 are merging at the level of elytral distal 1 / 5. Veins 4 and 5 are close to each other, half as long as the elytron. Fields III, V and VI has straight double row of cells, field IV have 1 – 2 rows of cells. Field IV are expanded at the base and has up to 3 cells. Cells roundish, rarely oval-shaped, without encrusted maculae. Cells larger near humeral area, size decrease very slightly towards the apex. Measurements (in mm). Elytron length 11.6, width 2.9.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141727D3174C8AF94D7FA9FD99.taxon	etymology	Etymology. After Latin “ large ”, due to considerable elytron size.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141727D3174C8AF94D7FA9FD99.taxon	discussion	Remarks. The specimen was attributed to the genus Cytocupes due to having veins 2 and 3 merging near elytral apex; having short vein 4 (CuA sensu Ponomarenko 1969) near elytral base; lack of cells on epipleural margin. Comparison. Differs from other congener species in large size; having elytral apex not elongated into tail-like extension; field IV being wider than V, having 3 cells in it instead of 6.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141726D3174C8AF97878D6F8A8.taxon	type_taxon	Type species. Tatarocupes granulatus Ponomarenko, 2004 Species included. Type species and Tatarocupes brevicellularis Yan & Strelnikova, sp. nov.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141726D3104C8AF8A07961FD55.taxon	description	(Fig. 4)	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141726D3104C8AF8A07961FD55.taxon	materials_examined	Material. Holotype — PIN, No. 5890 / 13. Isolated elytron, left. Paratypes: PIN, No 5890 / 12, part and counterpart, and PIN, No 5890 / 15, partially preserved isolated elytra. Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141726D3104C8AF8A07961FD55.taxon	diagnosis	Diagnosis. Elytron of middle size, carinated, bearing at least 5 veins and one scutellar striole. Cells roundish, or rarely slightly oval-shaped, reinforced with dark maculae. Elytral surface covered with dense tubercles. Vein 2 drows close to epipleural carina near humerus; Vein 3 very short, as long as combined length of 4 cells. Cells between veins are placed unevenly, without forming well-defined longitudinal rows.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141726D3104C8AF8A07961FD55.taxon	description	Description. Epipleural carina widening at basal third, then narrowing towards elytral apex. Vein 2 drows located close to epipleural carina near humerus then widens posteriorly; Vein 3 very short, as long as combined length of 4 cells, veins 3 and 4 are close together. Epipleuron without cells, in the field II up to 3 rows of cells, field III — 3 in central part and 4 in basal part, IV — 1 row of cells. There are more than 34 cells in a row. Measurements (in mm). Holotype — Elytron length 8.6, width 3; paratype PIN, No. 5890 / 15 — length of the preserved part of the elytron 6.9, width 2.6.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141726D3104C8AF8A07961FD55.taxon	etymology	Etymology. After Latin “ brevis ” — small, and “ cellularis ” — due to lattice pattern formed by cells.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141726D3104C8AF8A07961FD55.taxon	discussion	Remarks. The specimen was attributed to the genus Tatarocupes for having at least 6 rows of cells between two veins, closest to sutural margin. Comparison. Differs in having a single row of cells between veins 3 and 4 (at least near the elytral base).	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141721D3114C8AF94C7D4DFEC1.taxon	type_taxon	Type species. Alisacupes dunaevi Yan & Strelnikova, sp. nov.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141721D3114C8AF94C7D4DFEC1.taxon	diagnosis	Diagnosis. Elytron with 4 veins, at least two merging at the elytral apex. Epipleuron covered with tubercles. Composition. Monotypic.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141721D3114C8AF94C7D4DFEC1.taxon	description	Description. Large elytra, elongated, apex slightly pointed, assymetrical. Epipleural flange wide, at least as wide as several cell`s width combined. Elytral cells roundish, walls reinforced with maculae, cuticular surface between cells covered with tubercles. Сells are surrounded by macula.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141721D3114C8AF94C7D4DFEC1.taxon	etymology	Etymology. After Alisa quarry, source locality of the described material. Gender: masculine.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141721D3114C8AF94C7D4DFEC1.taxon	discussion	Remarks. Species below was attributed to Asiocoleidae for having four veins, with three of them being on the elytral disc; large size of the elytron. Comparison. Differs in moderately large size; in having tubercles at proximal half of epipleron`s length. Similar to Loculitricoleus but differs from it in larger size, tubercles sculpture on elytron and by four main veins instead of three.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141720D3114C8AFE557E2DFC1E.taxon	description	(Fig. 5)	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141720D3114C8AFE557E2DFC1E.taxon	materials_examined	Material. Holotype — PIN, No. 5890 / 16. Part and counterpart of isolated elytron, left. Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141720D3114C8AFE557E2DFC1E.taxon	diagnosis	Diagnosis. As for the genus.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141720D3114C8AFE557E2DFC1E.taxon	description	Description. Elytron with 4 veins, at least 2 nd and 3 rd are merging at elytral apex. Epipleural flange as wide as combined width of two cells near elytral base, slightly widens near the apex. Number of cell rows in the fields: II — 2, field III — 2 in central part and 3 near elytral basal, field IV — 4, field V — 3, field VI — 1. There are up to 43 cells in a row. Measurements (in mm): eytron length 13.6, elytron width 4.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141720D3114C8AFE557E2DFC1E.taxon	etymology	Etymology. After prominent naturalist, teacher and herpetologist E. A. Dunaev.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141720D3114C8AFE557E2DFC1E.taxon	discussion	Remarks. The specimen was attributed to the genus Tatarocupes for having at least 6 rows of cells between two veins, closest to sutural margin.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3124C8AFE957EB2FCB5.taxon	type_taxon	Type species. Uskatocoleus zalesskyi Rohdendorf, 1961 Species included. In total, 19 species: type species, U. artus Ponomarenko, 2015, U. avitus Rohdendorf, 1961, U. convexus Rohdendorf, 1961, U. convexus Ponomarenko, 2013, U. eurypygus Ponomarenko, 2011, U. foveolatus Rohdendorf, 1961, U. hirsutus Ponomarenko, 2015, U. iljinskiensis Rohdendorf, 1961, U. kaltanicus Rohdendorf, 1961, U. kushejakovicus Rohdendorf, 1961, U. lutugini Rohdendorf, 1961, U. micron Ponomarenko, 2013, U. minimus Rohdendorf, 1961, U. minor Ponomarenko, 2015, U. rhynchophorus Ponomarenko, 2011, U. sukhonensis Ponomarenko, 2013, U. surijokovicus Rohdendorf, 1961, and U. uralensis Ponomarenko, 2013.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3124C8AFE957EB2FCB5.taxon	discussion	Remarks. U. convexus Rohdendorf, 1961 and U. convexus Ponomarenko, 2013 are homonyms, therefore, in this article a new species name is proposed for the junior homonym — U. isadicus nom. nov. in honor of the location of Isady, from where this species was described.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3134C8AFCA17F15FF5D.taxon	description	(Fig. 6)	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3134C8AFCA17F15FF5D.taxon	materials_examined	Material. Holotype — PIN, No. 5890 / 1, part and counterpart, a pair of interlocked elytra with mesothorax (mesonotum and mesoscutellum), abdominal apex and mesotarsus — partially preserved. Paratypes: PIN, No. 5890 / 5 and PIN, No. 5890 / 7, parts and counterparts of isolated elytra, right and left respectively. Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3134C8AFCA17F15FF5D.taxon	diagnosis	Diagnosis. Scutellum large, semicircular, only slightly shorter than 1 / 2 of elytral tongue`s (schiza) length; scutellar incision on the sutural margin near elytral base, forms a weak protrusion. Humerus forms almost right angle. Elytra`s undersides bear very long elytral tongue, as long as 1 / 5 of elytral overall length, slightly outwardly curved. Elytron`s apices flattened.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3134C8AFCA17F15FF5D.taxon	description	Description. Elongated elytron with round bases, proximal 2 / 3 of the length — parallel-sided, then gradually narrowing posteriorly, distal third oblique, width to length ratio 1: 5. Elytral surface uniformly covered with punctures, inner structures (columellae also visible on the elytral surface as bigger punctures). Humeral bulges indistinct. Elytron`s tongues located at the midlength. Elytron edged, which is slightly thicker around humerus, at basal ¼ and at the rounded apex. Remnants of M, Cu veinal systems and bases of A are present. Mesocoxae transverse, slightly oblique, contiguous. X-shaped metathoracic scutoscutellar suture is partially visible. Apical abdominal ventrite subtrapezoid, almost as long as penultimate sternite, abdominal length makes less than a half of elytral length. Measurements (in mm). Holotype — elytral length 7.1 (measured without elytron`s basal attachment protrusions), maximum width 1.6; paratype PIN, No. 5890 / 5 — length 9.5, width 2.7; paratype PIN, No. 5890 / 7 — length 9.1, width 2.8.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3134C8AFCA17F15FF5D.taxon	etymology	Etymology. After Tura village, nearest settlement to Alisa quarry, where types specimens were found.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3134C8AFCA17F15FF5D.taxon	discussion	Remarks. Attributed to Uskatocoleus for having finely sculptured elytral surface, well defined humeral bulge and straight interlocking longitudinal protrusion (“ schiza ”). As an elytra-only family, Schizocoleidae possess very few diagnostic characters, specimens are mainly divided into taxa by locality and age. Cuticle on PIN, No. 5890 / 1 could been rather thin, since remnants of hindwing venation could be observed. Comparison. Similar to Uskatocoleus zalesskyi Rohdendorf, 1961 in having long elytral tongue, but differs in lacking of fine longitudinal stria and having twice the length. Similar to U. angustior Rohdendorf, 1961 in having long elytral tongue, but differs in twice the length, having concave elytral tongue and parallel-sided 2 / 3 of basal elytral length. Differs from Rossocoleus verrucosus Rohdendorf, 1961, R. latus Rohdendorf, 1961 and R. latior Rohdendorf, 1961 in twice the length. Differs from Rhombocoleites uralensis Ponomarenko, 2013 in having longer elytral tongue and apical abdominal sternite half as long.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173DD30C4C8AFE497F2AFBFD.taxon	description	(Fig. 7)	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173DD30C4C8AFE497F2AFBFD.taxon	materials_examined	Material. Holotype — PIN, No. 5890 / 2, part and counterpart of left elytron, base and apex partially damaged. Paratypes — PIN, No. 5890 / 3, 6 and 9 parts and counterparts of left elytra, partially damaged. Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173DD30C4C8AFE497F2AFBFD.taxon	diagnosis	Diagnosis. Very large elytron, narrowing anteriorly and posteriorly with maximum width at midlength.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173DD30C4C8AFE497F2AFBFD.taxon	description	Description. At least three times as long as wide, maximum width at two thirds of its length; carinated; slightly curved near distal third. Locking tongue straight about ¼ of elytron`s length, located at 1 / 3 of elytron`s width, closer to epipleural margin at elytron`s basal half. Specimens 5890 / 2 and 6 have remnants of R and M veinal systems visible. Measurements (in mm). Holotype’s elytral length 14.5, width 4.5; paratype`s PIN 5890 / 3 length 12.3, width 4; paratype`s PIN 5890 / 6 length 11.7, width 4.7; paratype`s PIN 5890 / 9 length 9.5, width 2.7.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173DD30C4C8AFE497F2AFBFD.taxon	etymology	Etymology. After Latin “ maximus ” due to elytron`s large size, largest among Schizocoleus.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173DD30C4C8AFE497F2AFBFD.taxon	discussion	Remarks. Elytral cuticle was probably rather thin, this allowed folded wings to be partially seen after fossilization, similar to Uskatocoleus turensis sp. nov. Species below attributed to Schizocoleus mainly because of its large size and lack of wrinkled or scaly cuticular sculpture. Comparison. Differs from all congeners by large size.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173DD30C4C8AFF6978D9FEC1.taxon	type_taxon	Type species. Schizocoleus kuznetskiensis Rohdendorf, 1961, Species included. In total, 9 species: type species, S. depressus Ponomarenko, 2013, S. glabrus Ponomarenko, 2004, S. longus Ponomarenko, 2004, S. major Ponomarenko, 2013, S. minimus Ponomarenko, 2013, S. robustus Ponomarenko, 2013, S. septentrionalis Ponomarenko, 2008, and S. triassicus Ponomarenko, 2016	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173CD30D4C8AFF697FAAFD55.taxon	description	(Fig. 8)	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173CD30D4C8AFF697FAAFD55.taxon	materials_examined	Material. Holotype — PIN, No. 5890 / 14, right elytron. Paratype — PIN, No. 5890 / 4, part and counterpart of left elytron. Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173CD30D4C8AFF697FAAFD55.taxon	diagnosis	Diagnosis. Asymmetrical base with flat flange going obliquely towards epipleural margin; interlocking longitudinal protrusion (“ schiza ”) makes 1 / 3 of elytral length or slightly less.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173CD30D4C8AFF697FAAFD55.taxon	description	Description. At least three times as long as wide, basal 2 / 3 of the length — parallel sided, then gradually narrowing towards asymmetrical, pointed apex; finely carinated. Interlocking device located at the midlength level, closer to the epipleural margin, distanced from the elytral edge for 1 / 3 of maximum elytral width. Measurements (in mm). Holotype’s elytral length 8.5, width 2.8; paratype`s PIN, No. 5890 / 4 length 7.2, width 2.4	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173CD30D4C8AFF697FAAFD55.taxon	etymology	Etymology. After Latin “ frumentum ” meaning “ seed ” or “ grain ”, due to elytron`s shape. Comparison. Differs from all congeners in having an assymetrical base with flat flange going obliquely towards epipleural margin and rather long interlocking longitudinal protrusion.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173FD30E4C8AFF697808FD69.taxon	description	(Fig. 9)	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173FD30E4C8AFF697808FD69.taxon	materials_examined	Material. Holotype — PIN, No. 5890 / 11, part and counterpart of right elytron. Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173FD30E4C8AFF697808FD69.taxon	diagnosis	Diagnosis. Asymmetrical base with weak scutellar incision; epipleuron wide around humerus at 1 / 5 of elytron`s proximal lenth, then abruptly narrows towards the apex; interlocking longitudinal protrusion narrow (“ schiza ”), long, makes 3.5 of elytral length, lies closely to elytron`s midwidth.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173FD30E4C8AFF697808FD69.taxon	description	Description. Elytron carinated; weakly narrows from the base; apex asymmetrical, pointed. Epipleural margin obliquely narrows toward the apex at elytron`s distal third; sutural margin weakly curved. Humeral bulge weakly defined. Measurements (in mm). Elytral length 6.4, width 2.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173FD30E4C8AFF697808FD69.taxon	etymology	Etymology. After Latin “ subtilis ” meaning “ small ”, due to elytron`s moderate size. Comparison. Asymmetrical base with weak scutellar incision. Interlocking longitudinal protrusion narrow (“ schiza ”), long, makes 3.5 of elytral length, lies closely to elytron`s midwidth.	en	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
