identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E5727F10FF94FFA614A3FE77513D8BBA.text	E5727F10FF94FFA614A3FE77513D8BBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus (Ser.) Rchb., Deut. Bot. Herb. - Buch	<div><p>Bolanthus (Ser.) Rchb., Deut. Bot. Herb.-Buch: 205 (1841).</p><p>Type (see Hernández-Ledesma et al. 2015: 324): — Bolanthus hirsutus (Labill.) Barkoudah, Willdenowia 45 (3): 324 (2015) ≡ Saponaria hirsuta Labill., Dec. Syr. 4: 9 (1812) ≡ Gypsophila hirsuta (Labill.) Spreng., Syst. 2: 373 (1826).</p><p>Description:— Low prostrate or cushion-shaped perennials. Leaves small, linear, inflorescence dichasial, subcapitate. Flowers solitary or 2–25 in ± dense clusters. Bracts leafy, linear to linear-lanceolate, ± adpressed to the calyx. Calyx tubular-turbinate, with 5 projecting ribs and scarious commissures, without calcium oxalate druses. Petals white or bright pink, often with purple veins, with long linear delicately winged claw and small limb. Stamens 10, anthers not exserted. Styles 2. Ovary with 8–20 ovules. Capsule oblong-ovoid, dehiscing by 4 teeth. Seeds comma-shaped, with projecting radicle (Huber-Morath 1967).</p><p>Bolanthus minuartioides (Jaub. &amp; Spach) Hub.-Mor. ≡ Heterochroa minuartioides Jaub. &amp; Spach, Pl. Or. 1: 26 (1843) ≡ Jordania minuartioides (Jaub. &amp; Spach) Boiss., Diagn. Ser. 1(8): 94 (1849) ≡ Gypsophila minuartioides (Jaub. &amp; Spach) Boiss., Fl. Or. 1: 558 (1867) ≡ Acanthophyllum minuartioides (Jaub. &amp; Spach) Bark., Wentia 9: 183 (1962).</p><p>Lectotype (designated here): — [Turkey B2 Manisa] in collibus aridis prope Selendi, Jaubert s.n. (P05059990! [photo]).</p><p>Occurrence of tubercle in calyx</p><p>Calyx shape</p><p>Size of calyx (mm)</p><p>Prominence and number of calyx vein</p><p>Calyx hairiness</p><p>Calyx colour</p><p>Shape of calyx teeth (CT)</p><p>Size of CT (mm)</p><p>CT hairiness</p><p>CT venation</p><p>CT margin</p><p>CT apex</p><p>Petal shape</p><p>Petal length (mm)</p><p>Shape of petal (PL)</p><p>Size of PL (mm)</p><p>Ratio length PL × petal</p><p>How much of the PL is outside the calyx Occurrence of spot in PL</p><p>PL hairiness (abaxial and adaxial surfaces)</p><p>PL colour (abaxial and adaxial surfaces) Number of PL teeth</p><p>Shape of PL teeth</p><p>Ratio length PL × teeth</p><p>Size of claw (mm)</p><p>Ratio claw width × collar width</p><p>Ratio lenght fruit × calyx</p><p>Occurrence of anthophore</p><p>Size of bracteola</p><p>Bracteola base (membranaceous or not)</p><p>Position of bracteoles (surround/not surround the calyx and pass through/not pass through) Filament length</p><p>Stylus length</p><p>Description (Figs. 1–5): — Perennial herbs. Stems prostrate, 1–8, 5–15 cm, base 0.8–1.2 mm diameter, loose branched, lower long eglandular hairs, upper eglandular and rarely mixed with glandular hairs, nodes 5–9, internodes 5–10 mm, eglandular hairs. Leaves subulate-setaceous, 2.5–5 × 0.4–0.6 mm, glabrous or puberulent hairs, prominent 3-nerved, greenish, ± adpressed to stem; terminal acute; sheaths membranous, 0.8–2 mm, puberulent hairs. Inflorescence in apex and side tight clusters sphere shaped, dense eglandular, rarely mixed with glandular hairs; flowers sessile, in each clusters 8–30. Bracts subulate, 4–5.5 × 0.7–0.9 mm, as long as inflorescence or passed slightly, dense eglandular, rarely mixed with glandular hairs; margin membranous, ciliate, ciliates 0.1–0.15 mm; terminal acute. Not petiolate. Calyx tubulate, dense eglandular, rarely mixed with glandular hairs, 4–5 × 1.2–1.5 mm; calyx teeth narrowly triangular, 1.8–2.5 mm tall, margins ciliate, terminal acute. Petals oblong, 5–7 × 0.8–1 mm, 1.2–1.5 times as much as sepals, white; terminal emarginate; base cuneate. Stamens 10, filament 3–3.5 mm tall. Styles 5–10 mm. Capsule oblong, 3.5–4 × 0.8–1 mm, 2–4-seeded, usually as much as sepals. Seeds comma shaped 0.8–1 × 0.6–0.7 mm, brown, side ridge surface tuberculate, granular.</p><p>Phenology:— June–July, 700–2080 m, rocky slopes. General distribution:— Endemic. East Mediterranean and Irano-Turanian element. Distribution for Türkiye (Fig. 5):— Central Anatolia, Aegean and Mediterraneanregion. Typification of the name Heterochroa minuartioides:— The protologue of Heterochroa minuartioides (Jaubert</p><p>&amp; Spach 1843: 26) consists of a detailed diagnosis and the provenance (“In Phrygiae locis petrosis, propè Selendi et</p><p>Taouchanleu, legit Jaubert”). No holotype was reported. As a consequence a lectotype should be designated according to the Art. 9 of ICN. We have selected the specimen at P (barcode P05059990) as the lectotype for H. minuartioides . This specimen was chosen because it matchs the protologue (both provenance and morphology) and corresponds to the current concept of the species by showing prostrate or decumbent habit, long-eglandular-hairy stem, entirely white without purple veins petals, tubular calyx, and short pedicel (Madhani et al. 2018, Koç et al. 2019).</p><p>Notes:— The name Heterochroa minuartioides was transferred to different genera during the time, i.e. Jordania Boiss. (by Boissier, 1849; see Bornmüller 1908), Gypsophilla L. (again by Boissier 1867), Acanthophyllum C.A. Mey. (by Barkoudah 1962) and, finally, to Bolanthus (by Huber-Morath 1967). The taxon differed from the other taxa in having loose, creeping stem structure, dense inflorescence, long sepals, white petals and being loosely branched (see the diagnostic key presented above). Also it has been recommended to be placed in the LC category particularly due to its widespread and stable population, with no significant decline based on the criterion A, B and C (not met).Although, area of occupancy (AOO) and extent of occurrence (EOO) were not quantitatively calculated, qualitative assessments confirm a broad distribution and a large, stable population. (IUCN 2017).</p><p>Specimens examined: — TÜRKİYE. B2 Kütahya: EskiŞehir-Kütahya 45 km, limestone rocks, 900 m, 13 June 1954, Demiriz (ISTF-13292); UŞak: Banaz, Murat mountain, stony meadow, 1100 m, 19 June 1978, Çırpıcı (ISTF– 31967); UŞak: around Kula, UŞak way, 10 km from UŞak way’s right, Ürün village, 720 m, 21 June 1993, Seçmen (EGE-18734); Kütahya: Gediz, Murat mountain, from the ski center to the left, way of go to summit, 2050 m, 02August 2015, Koç 2131 &amp; Hamzaoğlu (Bozok Univ. Herb.); Kütahya: between Aslantepe and Çavdarhisar, calcareous peaks, 1090 m, 24 June 2015, Koç 2006 &amp; Hamzaoğlu (Bozok Univ. Herb.); B3 Afyon: Dinar, Karakuzu, 05 July 1949, Attila (ISTF-8639); EskiŞehir: Haymana-Sivrihisar 93 km, 13 km ago from Sivrihisar, mountain steppe, 1120 m, 30 May 1956, Demiriz (ISTF-15420); EskiŞehir: SakaryabaŞı, 19 June 1980, Seçmen, Leblebici-Bekat (EGE-16804); Afyon: Sandıklı, around spas, 950 m, 08 June 1983, Gemici (EGE-25461); Afyonkarahisar: İscehisar, around Seydiler, 1150 m, 30 June 2012, Hamzaoğlu 6394 &amp; Koç (Bozok Univ. Herb.); Konya: Derebucak, entry of Çamlık town, sandy areas, 1300 m, 30 May 2005, Hamzaoğlu 3640 &amp; Koç (Bozok Üniv. Herb.); Konya: Derebucak, Çamlık town, environment of picnic area, 1370 m, 31 June 2010, Koç 1055 &amp; Hamzaoğlu (Bozok Univ. Herb.); Konya: between Kulu and Cihanbeyli, steppe, 1110 m, 15 June 2009, Koç 175, Hamzaoğlu &amp; Budak (Bozok Univ. Herb.); B4 Konya: between Bozkır and Akören, limestone rocks, 1105 m, 18 June 2013, Hamzaoğlu 6765 &amp; Koç (Bozok Univ. Herb.); C2 Denizli: Babadağ: Vakıfköy, sand mines, open sunny area, 1100 m, 25 May 1998, Oluk (EGE–34880); Antalya: 16 km from Korkuteli to Elmalı, limestone rocky, 27 June 1979, Seçmen, Leblebici &amp; Bekat (EGE-16805); Antalya: Elmadag, 1300 m, 28 May 1962, Sorger (ISTO-15048); Antalya: between Kızılcadağ and Korkuteli, 7 km from Korkuteli, meadows, 1080 m, 23 June 1980, Özhatay and Özhatay &amp; Tuzlacı (ISTE-45002); Antalya: east up of city, Susamlık hill, 1050 m, 04 June 1965, Dem, Emberker, Dölek, Ümit &amp; Tut (ISTF-20507); YeŞilova exit lakeside, 1150 m, 06 July 1983, Bekat &amp; Leblebici (EGE-18863); C3 Antalya: Çobanisa–Korkuteli, 3 km from Çobanisa, Öküzgözü location, 1200 m, 09 June 1996, Özhatay and Özhatay &amp; Duman (ISTE-72345); Burdur: Çeltikçibeli, forest forestation, pine forests under, 25 May 1996, Baytop &amp; Çubukçu (ISTE-9532); Isparta: Between Keçiborlu and Dinar, 6 km from Dinar, meadows, 1000 m, 31 May 1965, Dem, Emberker, Attila &amp; Aslanerer (ISTF-20122); Burdur: way of Karamanlı-Tefenni, 1 km, Juniperus, Quercus forest, stony meadows, 1150 m, 01 June 1965, Dem, Emberker, Aslanerer &amp; Ümit (ISTF-20201); Burdur: way of Burdur –Tefenni, Karaçalı south, Asılık hill, stony meadows southeastern, 1000 m, 01 June 1965, Dem, Emberker, Aslanerer, Ümit &amp; Tut (ISTF-20190); Antalya: Çeltikçi– Burdur, Pinus brutia forest, 1000 m, 04 June 1965, Dem, Emberker, Dölek, Ümit &amp; Tut (ISTF-20497); Antalya: Çalbalı Dağ, SE part of Bakırlı Dağ, meadows and deforested rock slopes, chiefly limestone with patches of serpentine, 1550–1650 m, 18 July 1984, Görk, Hartvig &amp; Strid (EGE-31426); Isparta: Senirkent, 900 m, 09 June 1983, Bekat (EGE-37007); Isparta: Eğridir, Kasnak forest calcareous land, mixed forest, 1100 m, 03 July 1974, Peşmen &amp; Güner 1665 (HUB-03417); Konya: Derebacak, PınarbaŞı, Kaklık location, stony places, 1400 m, 02 July 2002, Demirelma (KNYA-22688); Isparta: Keçiborlu, northwest of town, oak clearance stony, 1250 m, 16 June 2013, Hamzaoğlu 6748 &amp; Koç, (Bozok Univ. Herb.); Isparta: between Keçiborlu and Uluborlu, 1700 m, 08 July 2014, Budak 3144, Hamzaoğlu &amp; Koç, (Bozok Univ. Herb.); C4 İçel: Ermenek, 10 June 1950, Attila (ISTF-10428); Konya: Ermenek, Kazancı township, 900–1100 m, 20 June 1983, Sümbül (HUB-03419); Konya: Obruk, Kızıltepe, 1250 m, 15 June 1982, Dural (KNYA-14503); Konya: southwestern of Çal mountain, J. excelsa clearance, 1600 m, 30 May 1989, Evce, Serin &amp; Tatlı (KNYA-1233); Konya: way of BeyŞehir, Keçili stream location, 1450 m, 09 June 1979, Dural (KNYA-1231); Karaman: Kızılyaka township, road circumference, inside of Quercus forest, 1400 m, 28 July 1984, Serin (KNYA-1236); C5 Konya: Ereğli, Aydos mountain, Delimahmutlu, 2 km east, juniper forest, limestone rock, 1650 m, 26 June 1976, Erik 1636 (HUB-03418).</p><p>Bolanthus spergulifolius (Jaub. &amp; Spach) Hub.-Mor., Notes Roy. Bot. Gard. Edinburgh 28: 23 (1967) ≡ Heterochroa spergulifolia Jaub. &amp; Spach, Fl. Or. 1: 28 (1843) ≡ Jordania spergulifolia (Jaub. &amp; Spach) Boiss. &amp; Heldr., Diagn. Ser. 1(8): 94 (1849) ≡ Gypsophila spergulifolia (Jaub. &amp; Spach) Boiss., Fl. Or. 1: 559 (1867) = Gypsophila jaubertiana Boiss., Fl. Or. Suppl. 89 (1888) ≡ Acanthophyllum spergulifolium (Jaub. &amp; Spach) Bark., In Wentia 9: 182 (1962)</p><p>Lectotype (designated here):— [Turkey B2 UŞak] In siccis Phrygiae, Jaubert s. n. (P04981509! [photo]).</p><p>Description (Figs. 2–5): — Perennial herbs. Stems prostrate, 1–8, 9–18 cm, base 0.8–1.2 mm diameter, loose branched, complately dense glandular hairs, nodes 5–9, internodes 10–20 mm, dense glandular hairs. Leaves subulate-setaceous, 5–7 × 0.5–0.7 mm, glandular hairs, prominent 1-nerved, greenish, not adpressed to stem; terminal acute; sheaths membranous, 1–1.3 mm, glandular hairs. Inflorescence in apex and side loose clusters, dense glandular hairs; flowers pedicellate, in each clusters 5–9. Bracts, 4–5 × 0.7–0.9 mm, almost as long as inflorescence, dense glandular hairs; margin membranous, ciliate, ciliates 0.1–0.15 mm; terminal acute. Pedicels 1–3 mm, dense glandular hairs. Calyx tubulate, dense glandular hairs, 4.5–5.5 × 1.2–1.4 mm; calyx teeth narrowly triangular, 1–1.5 mm tall, margins ciliate, terminal acute. Petals oblanceolate, 6–7 × 0.8–1.2 mm, 1.5 times as much as sepals, limb pink, petal nerves dark ping; terminal emarginate; base cuneate. Stamens 10, filament 3.5–4.5 mm tall. Styles 2, 4–5 mm tall. Capsule oblong, 4–4.5 × 1–1.4 mm, 2–4-seeded, usually as much as sepals. Seeds comma shaped 0.8–1.1 × 0.6–0.8 mm, brown, side and ridge surface tuberculate, granular.</p><p>Phenology: June–July, 1150–2100 m, serpentine stony places.</p><p>General Distribution: Endemic. East Mediterranean element.</p><p>Distribution for Türkiye (Fig. 5): The Mediterranean region.</p><p>Typification of the name Heterochroa spergulifolia:— The protologue of Heterochroa spergulifolia (Jaubert &amp; Spach 1843: 28) consists of a detailed diagnosis and the provenance (“In Phrygiâ, unà cum Heterochroa minuartioidi, legit Jaubert”). No holotype was reported.As a consequence a lectotype should be designated according to the Art. 9 of ICN. We have selected the specimen at P (barcode P04981509) as the lectotype for H. spergulifolia . This specimen was chosen because it matches the protologue (both provenance and morphology) and corresponds to the current concept of the species by showing many branched, densely glandular and eglandular stem, subulate leaves, white with purple veins petals, and dichasial subcapitate subsesile inflorescence (Koç &amp; Hamzaoğlu 2015).</p><p>Notes:— The taxon was firstly defined with the name Heterochroa spergulifolia by Jaubert and Spach in 1843 (Boissier 1867). Later on, it was transferred to the genus Jordania by Boissier in 1849 (Bornmüller 1908). The taxon was transferred to the genus Gypsophilla again by Boissier in 1867, and it was transferred to the genus Acanthophyllum by Barkoudah (Boissier 1867, Barkoudah 1962). Finally, the taxon was transferred to the genus Bolanthus by Huber-Morath in 1967 (Huber-Morath 1967). Also, the taxon Gypsophila jaubertiana published by Boissier was defined as synonym species of the Bolanthus spergulifolius in 1888 (Boissier 1867). The taxon differed from the other taxa in terms of having loose, creeping stem structure, loose inflorescence, long, pink petals and relatively long internodes. Also it has been recommended to be placed in the VU B1a category (IUCN 2017).</p><p>Specimens examined: — TÜRKİYE. B2 Kütahya: Gediz, Murat mountain, way of spas, serpentine soils, grassy plains, 1495 m, 25 June 2015, Koç 2043 &amp; Hamzaoğlu (Bozok Univ. Herb.) ; B5 Niğde: Melendiz mountains, Asmasız village, 2100 m, 30 July 1982, Ocakverdi (KNYA-1237) .</p></div>	https://treatment.plazi.org/id/E5727F10FF94FFA614A3FE77513D8BBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF9DFFA714A3FD7E57528CE4.text	E5727F10FF9DFFA714A3FD7E57528CE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus cherlerioides (Bornm.) Barkoudah, Wentia	<div><p>Bolanthus cherlerioides (Bornm.) Barkoudah, Wentia 9: 170 (1962) ≡ Gypsophila cherlerioides Bornm., Mitt. Bot.</p><p>Ges. 23: 2 (1908)</p><p>Lectotype (designated by YıldızbaŞ &amp; Koç 2018: 199): — [Turkey B3 Konya] Sultandagh, in jugis alpinis supra AkŞehir, 1600–1800 m, 21 June 1899, Bornmüller 4399 (E00135537, photo!; isolectotype G00418268, photo!).</p><p>Description (Figs. 3–5): — Perennial herbs. Stems dense compact cushions, ascending-erect, 50–300, 1–3 cm, base 0.5–0.7 mm diameter, compact branched, complately glandular hairs, nodes 3–5, internodes 0–1 mm, dense imbricate. Leaves setaceous, 3–4.5 × 0.3–0.5 mm, glandular hairs, prominent 3—nerved, greenish, rotundate; terminal acute; sheats membranous, 0.3–0.5 mm, glandular hairs. Inflorescence in apex, glandular hairs; flowers sessile, 1—flowers. Bracts setaceous, 3–4.5 × 0.3–0.5 mm, almost as much as inflorescence, glandular hairs; margin membranous, ciliate, ciliates 0.1–0.15 mm; terminal acute. Not pedicellate. Calyx tubulate, dense glandular hairs, 4–6 × 1.2–1.4 mm; calyx teeth narrowly triangular, 1.5–2.3 mm tall, margins ciliate, terminal acute. Petals oblanceolate, 4–7 × 0.8–1.2 mm, almost up to sepals, limb pink, petal nerves dark pink; terminal acute; base cuneate. Stamens 10, filament 3–4 mm tall. Styles 2, 4–6 mm tall. Capsule oblong, 3.5–5.5 × 1–1.5 mm, 2–4-seeded, usually as much as sepals. Seeds comma shaped 0.8–0.9 × 0.6–0.8 mm, black, side and ridge surface tuberculate, granular.</p><p>Phenology: July–August, 1550–2635 m, limestone places.</p><p>General Distribution: Endemic. East Mediterranean and Irano–Turanian element.</p><p>Distribution for Türkiye (Fig. 5): Central Anatolia and Mediterranea region.</p><p>Notes:— The taxon was firstly defined with the name Gypsophila cherlerioides by Bornmüller in 1908 (Bornmüller 1908). Later on, it was transferred to the Genus Bolanthus by Barkoudah in 1962 (Barkoudah 1962). The taxon differed from the other taxa in terms of being densely tufted, forming imbricate leaves(no visible internodes), being single flowered and having petals and sepals with similar sizes. It was stated that the holotype specimen of the taxon had been kept in the B herbarium before it was damaged (Huber-Morath 1967). Therefore the isotype specimen kept in the E herbarium was selected as lectotype (YıldızbaŞ &amp; Koç 2018). Also it has been recommended to be placed in the VU B1-a category (IUCN 2017).</p><p>In recent years, a plant spreading in the Mersin Sertavulus Pass, Bolanthus sertavulus Özçelik, has been described by Özçelik (2024). This new species shares many similarities with B. cherlerioides but differs in several features. It is characterized by its dense, erect, and long glandular hairs on above-ground parts, a shorter and thicker stem (1–2 cm), shorter and stiffer leaves, and a height of 5–10 cm. Therefore, although this study recognizes it as a potentially new species, definitive separation from B. cherlerioides or acceptance as a synonym needs genetic analysis and detailed ecological studies.</p><p>Specimens examined: — TÜRKİYE. B3 Afyon: Çay, Sultandağları, Gelincikana hill south, 2000 m, 30 July 1976, Dökmeci (ISTE-35677); Konya: Sultandağları, Çakıllar tops, Sarıpınar location, 1800 m, 15 July 1980, Çakırer (ISTE-45488); Konya: AkŞehir, Sultandağları, Tekke tops, Karagür basin, around Hanoğlu stream, 1950 m, 28 September 1976, Dökmeci, (ISTE-35997); Konya: Sultandağları, Cankurtaran tops, 1900 m, 29 September 1976, Dökmeci (ISTE-36068); Konya: AkŞehir, Sultandağı Örkenez gate, 1600 m, Seçmen (EGE-32509); Konya: AkŞehir, Şarkikaraağaç exit, 09 July 1989, Erik 4048, Sümbül &amp; Venter (HUB-03413); Konya: Sultan mountains, Doğanhisar, west of Fırınlı village, Gökgedik hill, 1850 m, 26 June 1978, Ocakverdi (KNYA); Konya: between AkŞehir and Gelendost, 1570 m, 15 June 2009, Koç 217, Hamzaoğlu &amp; Budak (Bozok Univ. Herb.); Konya: between AkŞehir and Şarkikaraağaç, around Yellibel gate, massive slopes, 1550 m, 14 July 2011, Hamzaoğlu 6136 &amp; Koç, (Bozok Univ. Herb.); C3 Isparta: Davras mountain, [Ascent from the Kır Yayla], N–facing cirque and summit region above the yayla i.e. the main ridge beetween the OğlaktaŞı hill and Ulparçukur hill [= main summit, 2635 m] –Lat. 37˚45ˈ77”N, long. 030˚44ˈ 64”E [cirque]; alt. c. 2000–2550 m, Limestone and dolomite, 2100 m, 28 June 2000, Eren &amp; Parolly (ISTE-100796).</p></div>	https://treatment.plazi.org/id/E5727F10FF9DFFA714A3FD7E57528CE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF9CFFA714A3FBBB504F8764.text	E5727F10FF9CFFA714A3FBBB504F8764.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus thymoides Hub. - Mor., Bauhinia	<div><p>Bolanthus thymoides Hub.-Mor., Bauhinia 2(2): 187, t. 3 (1963)</p><p>Type: — [Turkey C 2 Burdur] d. Tefenni, Passhöhe südlich ob Dirmil, auf Serpentinschutt, 1600–1660 m, 28 June 1948, Huber-Morath 8447 (holotype G 00006008, photo!).</p><p>Description (Figs. 6–9): — Perennial herbs. Stems cushions, ascending-erect, 5–50, 3–8 cm, base 0.8–1 mm diameter, compact branched, complately glandular hairs, nodes 3–6, internodes 2–5 mm, glandular hairs. Leaves linear-setaceous, 3–6 × 0.5–0.7 mm, glandular hairs, prominent 3-nerved, greenish, not adpressed to stem; terminal acute; sheaths membranous, 0.2–0.3 mm, glandular hairs. Inflorescence in apex, glandular hairs; flowers sessile, 5–10-flowers. Bracts linear-setaceous, 4–5.5 × 0.5–0.7 mm, almost as much as inflorescence or passed 1.2 times, glandular hairs; margin membranous, ciliate, ciliates 0.1–0.15 mm; terminal acute. Not pedicellate. Calyx tubulate, dense glandular hairs, 4–5 × 1–1.3 mm; calyx teeth narrowly triangular, 1–1.3 mm tall, margins ciliate, terminal acute. Petals linear-oblong, 5–6 × 0.7–0.9 mm, almost up to sepals or up to 1.2 times, limb white, petal nerves dark pink; terninal truncate; base cuneate. Stamens 10, filament 3–4 mm tall. Styles 2, 3–4 mm tall. Capsule oblong, 3.5–4.5 × 1–1.3 mm, 2–4-seeded, usually as much as sepals. Seeds comma shaped 0.8–0.9 × 0.6–0.8 mm, brown, side and ridge surface tuberculate, granular.</p><p>Phenology: June–July, 605–2000 m, serpentine stony slopes.</p><p>General Distribution: Endemic. East Mediterranean element.</p><p>Distribution for Türkiye (Fig. 9): Aegean and Mediterranean region.</p><p>Notes:— The taxon was firstly defined with the name Bolanthus thymoides by Huber-Morath in 1963 (Huber-Morath 1963). The taxon has a wide distribution area in Türkiye. The taxon differed from the other taxa in terms of having skewed, ascendant, tick stems branching from the base, tough leaves, short internodes and dense inflorescence with many flowers. Also it has been recommended to be placed in the LC category (IUCN 2017).</p><p>Specimens examined: — TÜRKİYE. C2 Muğla: Sandras mountain, northwestern of Çiçekbaba hill, Serçe crevice, 1850 m, 14 July 1979, Özhatay (ISTO-23928); Burdur: Altınyayla Dirmil Gate, Esenli plateau, roadside, 1610 m, 06 June 1996, Özhataya and Özhatay &amp; Duman (ISTE-72219); Burdur: Dirmil, hills from Dirmil above, 1750–1800 m, 08 July 1993, Bekat &amp; Leblebici (EGE-18864); Denizli: Altınyayla, Dirmil gate down, the open of steppe, 605 m, 20 June 2002, Dönmez 10922 &amp; Dönmez (HUB); Burdur: Çamlı plateau, Dirmil gate protected Pinus nigra and Juniperus sp. forest serpentine master rock, 1650 m, 30 June 1996, Aytaç 7405 (GAZI); Burdur: YeŞilova, Salda lake entry, 1300–1400 m, 09 July 2014, Budak 3179 (Bozok Univ. Herb.); Burdur: between YeŞilova and Salda village, EŞeler mountain, serpentine, stony places, 2000 m, 18 August 2014, Koç 1848 &amp; Hamzaoğlu (Bozok Univ. Herb.); C3 Burdur:Altınyayla, Dirmil gate, 1585 m, 27 July 1996, Özhatay &amp; Özhatay (ISTE-72866); Antalya: Elmalı, Çağlıkara, 1540 m, 28 June 1975, Çetik (KNYA-1238).</p></div>	https://treatment.plazi.org/id/E5727F10FF9CFFA714A3FBBB504F8764	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF9EFFA314A3FDC456878DCA.text	E5727F10FF9EFFA314A3FDC456878DCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus frankenioides (Boiss.) Barkoudah., Wentia	<div><p>Bolanthus frankenioides (Boiss.) Barkoudah., Wentia 9: 169 (1962) ≡ Gypsophila frankenioides Boiss., Diagn. Ser.</p><p>1(1): 10 (1843)</p><p>Lectotype [designated by Barkoudah (1962: 169) as “ holotype ”, here corrected according to the Art. 9.10 of ICN]: — [Turkey C2 Denizli] in regione alpina Cariae, Cadmus orientalis supra Khonas [Honaz Da.] ad nives, June 1842, Boissier s. n. (G00544521! [photo]; isolectotype: P01903171!).</p><p>Description:— Perennial herbs. Stems prostrate, 5v50, 8-15 cm, base 0.7–1 mm diameter, loose branched, complately glandular hairs, nodes 3–6, internodes 5–15 mm, glandular hairs. Leaves linear-oblanceolate, 3–5 × 0.5–0.7 mm, glandular hairs, sparsely glandular hairs or glabrous prominent 3-nerved, greenish, not adpressed to stem; terminal acute; sheaths membranous, 0.2–0.3 mm, glandular hairs. Inflorescence in apex and sides, glandular hairs; flowers pedicellate, 3–15-flowers. Bracts linear-oblanceolate, 4–5.5 × 0.5–0.7 mm, almost as much as inflorescence; margin membranous, ciliate, ciliates 0.1–0.15 mm; terminal acute. Pedicels 1–4 mm, glandular hairs. Calyx tubulate, glandular hairs, 3–3.5 × 0.9–1.1 mm; calyx teeth narrowly triangular, 1–1.3 mm tall, margins ciliate, terminal acute. Petals oblanceolate, 4–5 × 0.6–1 mm, up to 1.5 times sepals, lamina pink, petal petal nerves dark pink; terminal cute; base cuneate. Stamens 10, filament 3–4 mm tall. Styles 2, 3-4 mm tall. Capsule oblong, 2.5–3 × 0.9–1.1 mm, 2–4-seeded, usually as much as sepals. Seeds comma shaped 1–1.1 × 0.8–0.9 mm, brown, side and ridge surface tuberlculate, granular.</p><p>Pedicels 2–4 mm, as long as calyx...................................................................................................................................... var. frankenioides Pedicels 1–2 mm, shorter than calyx ..................................................................................................................................... var. fasciculatus</p><p>Typification of the name Gypsophila frakenioides:— The protologue of Gypsophila frankenioides (Boissier 1843: 10) consists of a detailed diagnosis and the provenance (“Hab. in pratis ad nives deliquescentes in parte superiori montis Cadmi supra Colossam ubi legi Junio 1842 ”). Barkoudah (1962: 169) stated that the holotype and isotype are at G and P, respectively. Also Madhani et al. (2024: 1245) indicated a holotype at G (G00544521), as well as a possible isotype (G00544571). However, no holotype was explicitly reported in the protologue by Boissier (1843: 10). By checking the herbarium G, we noted that Barkoudah annotated G00544521 as “ holotype ”, whereas G00544571 was annotated by him as “ isotype ”. According to the Art. 9.10 of ICN, Barkoudah’s indication of the holotype (which precedes Madhani et al. ’s proposal) and his annotation on G00544521, are errors to be corrected to lectotype. So, G00544521 is the lectotype and P01903171 is the isolectotype for G. frankenioides . Both these specimens matches the protologue and corresponds to the current concept of the species by showing prostrate stems, linear leaves with glandular hairy, oblanceolate pink petals, and inflorescence in apex and sides (YıldızbaŞ 2017).</p></div>	https://treatment.plazi.org/id/E5727F10FF9EFFA314A3FDC456878DCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF98FFA314A3FAD1513B8A66.text	E5727F10FF98FFA314A3FAD1513B8A66.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus frankenioides var. frankenioides var. frankenioides	<div><p>Bolanthus frankenioides var. frankenioides</p><p>Phenology: June–July, 1490–2060 m, usually around snow beds or flat landed areas.</p><p>General Distribution: Endemic. East Mediterranean Element.</p><p>Distribution for Türkiye: The Mediterranean region. (Figs. 7–9).</p><p>Notes:— The Bolanthus frankenioides var. frankenioides, originally described as Gypsophila frankenioides by Boissier in 1843 (Boissier 1843), was transferred to the genus Bolanthus by Barkoudah in 1962 (Barkoudah 1962). The taxon has a wide distribution area in Türkiye. The taxon differed from the other taxa in terms of having loose, creeping stem structure, reverse lanceolate leaves, relatively long internodes and short calyx. Also it has been recommended to be placed in the VU B1a category (IUCN 2017). Its distribution in Türkiye was given in Figure 9.</p><p>Specimens examined: — TÜRKİYE. C2 Antalya: Akdag, YeŞilgöl, 1800 m, 24 June 1967, Sorger (ISTO-15049) ; Burdur: Dirmil plateau way, the hills on Dirmil, 1750–1800 m, 08 July 1993, Bekat &amp; Leblebici (EGE-18865) ; Denizli: Babadağ, Evrentepe subalpine meadow, 1850 m, 09 August 1996, Oluk (EGE-34913) ; Antalya: Elmalı, Beydağı, Küçüksöğle village, Serkiz area above the plateau, Kırkınar crevice location, 2240 m, 28 July 2012, Hamzaoğlu 6550 &amp; Koç (Bozok Univ. Herb.) ; Antalya: Elmalı, Gömbe, SubaŞı plateau, Şirenli plains, 2060 m, 30 July 2012, Hamzaoğlu 6579 &amp; Koç (Bozok Univ. Herb.) ; Burdur: Altınyayla southwest, around Akpınar plateau, limestone slopes, 1865 m, 04 August 2015, Koç 2167 &amp; Hamzaoğlu (Bozok Univ. Herb.) ; Burdur: going from Altınyayla to Kemer, 1490 m, 25 June 2015, Koç 2062 &amp; Hamzaoğlu (Bozok Univ. Herb.) .</p></div>	https://treatment.plazi.org/id/E5727F10FF98FFA314A3FAD1513B8A66	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF98FFA014A3FD3D52E48CC0.text	E5727F10FF98FFA014A3FD3D52E48CC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus frankenioides var. fasciculatus (Boiss. & Heldr.) Barkoudah, Wentia	<div><p>Bolanthus frankenioides var. fasciculatus (Boiss. &amp; Heldr.) Barkoudah, Wentia 9: 170 (1962) ≡ Gypsophila frankenioides Boiss. var. fasciculata Boiss. &amp; Heldr., Diagn. Ser. 1(8): 58 (1849)</p><p>Lectotype [designated by Barkoudah (1962: 170) as “ isotype ”, here corrected according to the Art. 9.10 of ICN): —[Turkey C3 Isparta] in pascuis siccis montis Stavros Pisidiae supra Isbarta, 10 Aug 1845, Heldreich 1017 (G00544524, photo!; ısotype FI, n.v.).</p><p>= Bolanthus stenopetalus Hartvig &amp; Strid, In Bot. Jahrb. Syst. 108: 318 (1987)!, syn. nov.</p><p>Type: — [ Turkey C2 Muğla]. Sandras Dağ, W. Side of the summit area, 2100–2200 m, snowbed meadows and rocky slopes, serpentine, lat. 37˚04’ N, long. 28˚50’ E, 7 July 1984, Hartvig, Seçmen &amp; Strid 23375 (holotype C; isotypes B, E! EGE, G00226570!, photo!) .</p><p>Typification of the name Gypsophila frakenioides var. fasciculata:— Barkoudah (1962: 170) stated “Type: Turkey: Wilayet Antalya, Mt. Davros-dagh, Heldreich s. n. 1895, isotype (FI)”. However, no holotype or isotype was reported in the protologue. According to the Art. 9.10 of ICN, Barkoudah’s indication of the isotype is an error to be corrected to lectotype. So, G 00544524 is the lectotype and specimen at FI (see Madhani et al. 2024: 1245) isolectotype for G. frakenioides var. fasciculata .</p><p>Phenology: June–July, 1550–2200 m, usually around snow beds or flat landed areas.</p><p>General Distribution: Endemic. East Mediterranean Element.</p><p>Distribution for Türkiye: The Mediterranean region. (Figs. 8–9).</p><p>Notes:— Gypsophila frankenioides var. fasciculata was firstly described by Boissier and Heldrich (in Boissier 1849: 58) and later transferred to the genus Bolanthus by Barkoudah (1962). The taxon exhibits quite a few similarity with the Bolanthus frankenioides . The taxon differes from the other taxa in terms of its pedicel length and trichome density. Due to their overlapping distribution areas, these features were determined as varying morphological characters and thus it was deemed suitable to consider this taxon as variety. Also it has been recommended to be placed in the VU B1a category (IUCN 2017). Its distribution in Türkiye was given in Figure 9.</p><p>Bolanthus stenopetalus was firstly described by Hartvig &amp; Strid (1987: 318). The location of the type of this taxon is The Sandras Mountain of Muğla Province and at this location, this type is known to be found in the taxon Bolanthus frankenioides var. fasciculatus (Huber-Morath 1967) . The type specimen of the taxon and the specimens collected during the field studies at Sandras mountain of Muğla province were detaily examined. The paper published by Hartvig and Strid and the detailed descriptions were gathered and as a result, the features that were defined as distinctive characters including ciliated and long leaves, flowers with short pedicels, pink and reverse lanceolate petals and ovariums with 8-10 ovules were identified as the features of the taxon Bolanthus frankenioides var. fasciculatus . For this reason, the taxon was considered as synonym.</p><p>Specimens examined: — TÜRKİYE. C2 Muğla Köyceğiz, Sandras mountain, BeŞparmak hill rocky stony soil, 1900 m, 12 July 1978, Özhatay &amp; Özhatay (ISTO-32912) ; Muğla Köyceğiz, eastern slopes of Çiçekbaba hill, 1900 m, 14 July 1978, Özthatay &amp; Özhatay (ISTO-20482) ; Muğla: Sandras mountain, Çiçekbaba hill, serpentine cliff, 2200 m, 26 July 1977, Özhatay No: 1312 (ISTO-19455) ; Muğla: Sandras mountain, BeŞparmak hill, 1900 m, 12 July 1978, Özhatay &amp; Özhatay (ISTE-40513) ; Muğla / Antalya: Akdağ, pebbled slopes, limestone, 1700–2000 m, 08 July 1992, Gemici, Özel &amp; Görk (EGE-26644) ; Muğla: Sandras mountain, SW of the summit, Marshy places by a creek in opening of Pinus nigra forest, serpentine, 1750m, 06 July 1984, Hartvig, Seçmen &amp; Strid (EGE-27895) ; Antalya: Elmalı, Akdağ, 14 July 1968, Pamukçuoğlu (HUB-03415) ; Muğla: Köyceğiz, Yayla village, top of Gökçeova lake, Sandras mountain, serpentine rocky, 2030 m, 05 August 2015, Koç 2188 &amp; Hamzaoğlu (Bozok Univ. Herb.) ; C4 Konya: SeydiŞehir, south of Tınaz mountain, the open of SaydaŞ forest, 1550 m, 20 June 1982, Ocakverdi (KNYA-1216) .</p></div>	https://treatment.plazi.org/id/E5727F10FF98FFA014A3FD3D52E48CC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF9BFFA014A3FBD7577B8878.text	E5727F10FF9BFFA014A3FBD7577B8878.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus huber	<div><p>Bolanthus huber -morathii Simon, Bauhinia 6(1): 83 (1977).</p><p>Type: [ Turkey A2 Bursa] South side of the Uludağ, at the road between Soğukpinar and Keles, 3 km after Soğukpinar, ca. 900 m, 03 August 1976 (holotype P; isotype GAZI!) .</p><p>Description (Figs. 10–13): — Perennial herbs. Stems prostrate, 1–8, 9–18 cm, base 0.8–1.2 mm diameter, loose branched, glabrous or puberulent, nodes 5–9, internodes 5–10 mm. Leaves setaceous, 4–6 × 0.5–0.6 mm, glabrous, prominent 1-nerved, greenish, not adpressed to stem; terminal acute; sheaths membranous, 1–1.3 mm, glabrous or puberulent. Inflorescence and side loose clusters, dense glandular hairs; flowers pedicellate, in each clusters 5–9. Bracts subulate, 4–5 × 0.7–0.9 mm, up to ½ inflorescence, dense glandular hairs; margin membranous, ciliate, ciliates 0.1-0.15 mm; terminal acute. Pedicels 1–3 mm, dense glandular hairs. Calyx tubulate, dense glandular hairs, 4–5 × 1.2–1.4 mm; Calyx teeth narrowly triangular, 1–1.5 mm tall, margins ciliate, terminal acute. Petals oblanceolate, 6–7 × 0.8–1.2 mm, up to 1.5 times sepals, lamina pink, petal nerves dark pink; terminal emarginate; base cuneate. Stamens 10, filament 3.5–4.5 mm tall. Styles 2, 4–5 mm tall. Capsule oblong, 4–4.5 × 1–1.4 mm, 2–4-seeded, usually as much as sepals. Seeds comma shaped 0.8–1.1 × 0.6–0.8 mm, brown, side and ridge surface tuberculate, granular.</p><p>Phenology: June–July, 900–1300 m, serpentine places.</p><p>General Distribution: Endemic. East Mediterranean and Irano–Turanian element.</p><p>Distribution for Türkiye: Central Anatolia and Mediterranean region.</p><p>Notes:— The taxon was firstly defined with the name Bolanthus huber-morathii by Simon in 1977 (Simon 1977). The location of the type of this taxon is the serpentinized areas of the Bursa-Keles province. The taxon differes from the other taxa of the Bolanthus in terms of having hairless, creeping stems, setaceous shaped leaves, long internodes and pedicellate flowers. Also it has been recommended to be placed in the VU B1a category (IUCN 2017). Its distribution in Türkiye was given in Figure 13.</p><p>Specimens examined: — TÜRKİYE. A2 Bursa: between Soğukpınar and Keles, Soğukpınar exit, 930 m, 19 July 2016, Koç 2352 &amp; Hamzaoğlu (Bozok Univ. Herb.); B2 Kütahya: TavŞanlı, southwest of Arifler village, Pinus nigra, Quercus and Juniperus communities, 1300 m, 23 July 2014, Kocakaya clad 226 &amp; Koç (Bozok Univ. Herb.).</p></div>	https://treatment.plazi.org/id/E5727F10FF9BFFA014A3FBD7577B8878	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF9BFFBE14A3FF3F53FB8895.text	E5727F10FF9BFFBE14A3FF3F53FB8895.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus mevlanaea Aytaç, Ann. Bot. Fenn.	<div><p>Bolanthus mevlanaea Aytaç, Ann. Bot. Fenn. 41(3): 218 (2004).</p><p>Type:— [ Turkey C3 Antalya] Between Akseki and Bozkır, 45 km, Golcuk, 1780 m, steppe, 16 July 1997, Aytac 7733 (holotype GAZI!; isotypes: GAZI!, ANK, E, HUB) .</p><p>Description (Figs. 11–13): — Perennial herbs. Stems 20–50 cm wide compact branched ascending or erect, 3–5 cm, base 0.8–1.2 mm diameter, short puberulent haris, internodes 2–5 mm. Leaves setaceous, 2–5 × 0.5v 0.6 mm, glabrous, prominent 1-nerved, greenish, not adpressed to stem; terminal acute; sheaths membranous, 1–1.3 mm. Inflorescence in the terminal clusters, dense glandular hairs; flowers sessile, in each clusters 5–10. Calyx tubulate, dense glandular hairs, 4–5 × 1.2–1.4 mm; calyx teeth narrowly triangular, 1.5–2 mm tall, margins ciliate, terminal acute. Petals linear-oblong, 5.5–6 × 0.8–1.2 mm, up to 1.5 times sepals, limb white, petal nerves dark pink; terminal emarginate; base cuneate. Stamens 10, anthers 2–3 mm tall. Styles 2. Capsule oblong, 3–4 × 1–1.4 mm, 2–4-seeded, usually as much as sepals. Seeds comma shaped 0.8–1.1 × 0.6–0.8 mm, brown, side and ridge surface tuberculate, granular.</p><p>Phenology: July, 1780 m, stony places.</p><p>General Distribution: Endemic. East Mediterranean element.</p><p>Distribution for Türkiye: The Mediterranean region.</p><p>Notes:— The taxon was firstly defined with the name Bolanthus mevlanae by Aytaç in 2004 (Aytaç 2004). The type specimens of the taxon was detaily examined. The paper published by Aytaç and the detailed descriptions were gathered and as a result, the features that were defined as distinctive characters by the researcher including short internodes, sepals 4–5 mm long, absence of pedicels and dense inflorescence were determined to be the characters shared with Bolanthus thymoides . However, it differs from this taxon in terms of having hairless, setaceous shaped leaves. Also it has been recommended to be placed in the VU B1-a category (IUCN 2017). Its distribution in Türkiye was given in Figure 13.</p><p>Specimens examined: — TÜRKİYE. C3 Antalya: Between Akseki and Bozkır, 45 km, Golcuk, 1780 m, steppee, 16 July 1997, Aytaç 7733 (GAZI) ; Konya: south of BeyŞehir lake, around Üstünler, oak bottoms, 1150 m, Küçüködük, 30 June 1986 (KNYA-1234) .</p></div>	https://treatment.plazi.org/id/E5727F10FF9BFFBE14A3FF3F53FB8895	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF87FFBC14A3FB0E53408832.text	E5727F10FF87FFBC14A3FB0E53408832.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus turcicus Koc & Hamzaoglu, PhytoKeys	<div><p>Bolanthus turcicus Koç &amp; Hamzaoğlu, PhytoKeys 52: 82 (2015).</p><p>Type:— [ Turkey B4 Aksaray] Aksaray province, Hasan Mountain above Karkın town, 1950 m, volcanic stony slopes and alpine steppe, 18 July 2014, Hamzaoğlu 7110 &amp; Koç (holotype GAZI!; isotypes: GAZI!, ANK!, Dept of Bozok Univ. !) .</p><p>Description (Figs. 12–13): — Perennial herbs. Stems prostrate, 1–8, 9–18 cm, base 0.8–1.2 mm diameter, loose branched, dense mixed eglandular and glandular hairs, nodes 5–9, internodes 5–10 mm. Leaves linear, 5–8 × 0.5–0.7 mm, dense mixed eglandular and glandular hairs, prominent 3-nerved, greenish, not adpressed to stem; terminal acute; sheaths membrabous, 1–1.3 mm, dense mixed eglandular and glandular hairs. Inflorescence in apex and side loose clusters, dense mixed eglandular and glandular hairs; flowers pedicellate, in each clusters 5–10. Bracts subulate, 3–5 × 0.7–0.9 mm, almost as long as inflorescence, dense mixed eglandular and glandular hairs; margins membranous, ciliate, ciliates 0.1–0.15 mm; terminal acute. Pedicels 1–2 mm, dense mixed eglandular and glandular hairs. Calyx tubulate, dense mixed eglandular and glandular hairs, 3.5–4.5 × 0.8–1.2 mm; calyx teeth narrowly triangular, 1–1.5 mm tall, margins ciliate, terminal acute. Petals oblanceolate, 3.3–4.5 × 0.8–1.2 mm, almost as long as sepals, limb white, petal nerves dark pink; terminal emarginate; base cuneate. Stamens 10, filament 3.5–4.5 mm tall. Styles 2, 4–5 mm tall. Capsule oblong, 4–4.5 × 1–1.4 mm, 2–4-seeded, usually as much as sepals. Seeds comma shaped 0.8–1.1 × 0.6–0.8 mm, brown, side and ridge surface, granular.</p><p>Phenology: June–July, 1950–2760 m, volcanic stony slopes.</p><p>General Distribution: Endemic. Irano-Turanian element.</p><p>Distribution for Türkiye: Central Anatolia region.</p><p>Notes:— The taxon was firstly defined with the name Bolanthus turcicus by Koç and Hamzaoğlu in 2015 (Koç &amp; Hamzaoğlu 2015). Unlike the other taxa of Bolanthus which are distributed generally in the Mediterranean region, the taxon is distributed in the Central Anatolian Region and grows on volcanic soils. The taxon differes from the other taxa of the Bolanthus in terms of having long internodes, loose, creeping stem structure, petals rearly exceesing the sepals and bracts surrounding the calyx. Also it has been recommended to be placed in the EN B1a category (IUCN 2017). Its distribution in Türkiye was given in Figure 13.</p><p>Specimens examined: — TÜRKİYE. B4 Aksaray: above Karkın village, Hasan mountain, around Karbeyaz hotel, subalpine step, 38˚09ˈ39” N, 34˚09ˈ 53” E, 1950m, 18 July 2014, Hamzaoğlu 7110 &amp; Koç (Bozok Univ. Herb.) ; Aksaray: Karkın town, above Karbeyaz hotel, south of Hasan mountain, 2760 m, 07 August 2015, Koç 2226 &amp; Hamzaoğlu (Bozok Univ. Herb.) .</p></div>	https://treatment.plazi.org/id/E5727F10FF87FFBC14A3FB0E53408832	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF87FFBD14A3FFC650398CC0.text	E5727F10FF87FFBD14A3FFC650398CC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus sandrasicus Hamzaoglu & Koc, Nordic	<div><p>Bolanthus sandrasicus Hamzaoğlu &amp; Koç, Nordic Journal of Botany 35: 563–568 (2017).</p><p>Type:— [ Turkey C2 Muğla] Köyceğiz, South of Kartal Lake, 1960 m a.s.l, subalpine serpentine flats, 17 Jul 2016, Koç 2343 &amp; Hamzaoğlu (holotype GAZI!; isotypes: GAZI!, ANK!, Bozok Univ. Herb.!) .</p><p>Description (Figs. 14–16): — Perennial herb forming lax cushions. Stems ascending-erect, 2–3 cm long, 0.8–1.2 mm in diameter at base, with mixed eglandular and glandular hairs, 3–6-noded; internodes short, 1–2 mm long. Leaves linear-setaceous, 3–6 × 0.5–0.7 mm, glandular-hairy, 3-nerved, greenish, rigid; apex acute; sheath membranous, 0.2–0.3 mm long, glandular-hairy. Inflorescence congested in small sessile or subsessile terminal or subterminal clusters, 4–10- flowered, densely glandular-hairy; flowers sessile. Bracts linear-setaceous, 3.0–4.5 × 0.5–0.7 mm, as long as calyx; apex acute. Calyx tubular, densely glandular-hairy, 2.5–4.0 × 0.8–1.2 mm, with 5 projecting ribs; teeth lanceolate, 1.0– 1.3 mm long, with acute apex, purplish. Petals oblanceolate, 4.0–5.5 × 0.8–1.0 mm, 1.5 times longer than calyx, purple; apex rounded; base cuneate. Stamens 10; filaments 2.0– 3.5 mm long. Styles 2.2–2.5 mm long. Capsule oblong, 2.5–3.5 × 0.8–1.2 mm, 2–4-seeded, usually as long as calyx. Seeds brown, comma-shaped, with prominent radicle, 0.9–1.1 × 0.6–0.8 mm; ventral and dorsal surfaces with irregular polygonal cells.</p><p>Phenology: May–July, 1960 m, serpentine areas.</p><p>General Distribution: Endemic. East Mediterranean element.</p><p>Distribution for Türkiye: Aegean and Mediterranean region.</p><p>Notes:— The taxon was firstly defined with the name Bolanthus sandrasicus by Koç, Hamzaoğlu and Büyük in 2017 (Koç et al. 2017). It is similar to B. thymoides Hub.-Mor. but differs by stems 2–3 cm long (vs. 3–8 cm long), inter-nodes 1–2 mm (vs. 2–5 mm), calyx 2.5–4.0 mm long, teeth purplish (vs. 4–5 mm long, greenish), petals oblanceolate, 4.0– 5.5 mm long, purple (vs. linear-oblong, 5–6 mm long, white with purple veins), capsule 2.5–3.5 mm long (vs. 3.5–4.5 mm long), and seeds with ventral and dorsal surfaces having irregular polygonal cells (vs. regular rectangular cells). The taxon grows on the subalpine serpentine areas. Also, due to overgrazing, the habitat of this speciesis is under threat, it has been recommended to be placed in the EN B1a category (IUCN 2017). Its distribution in Türkiye was given in Figure 16.</p></div>	https://treatment.plazi.org/id/E5727F10FF87FFBD14A3FFC650398CC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF80FFBB14A3FABC56E488D4.text	E5727F10FF80FFBB14A3FABC56E488D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus aziz-sancarii Koc & Hamzaoglu, Systematic Botany	<div><p>Bolanthus aziz-sancarii Koç &amp; Hamzaoğlu, Systematic Botany 44(1): 189–196 (2019).</p><p>Type:— [ Turkey B3 Afyonkarahisar]. Afyonkarahisar: between Bayat and Iscehisar, 1500m, fissures of tuffaceous volcanic rocks, 02 Jul 2010, Koç 1209 &amp; Hamzaoğlu (holotype GAZI!; isotypes: GAZI!, ANK!, HUB!) .</p><p>Description (Figs. 15–16): — Perennial herb. Stems forming lax cushions, 4.5–15 cm in diam, ascendant-erect, base 0.8–1.2 mm diam, entirely covered with eglandular and glandular hairs. Stems short, 8–12-noded, internodes 4–8 mm, glandular-hairy. Leaves linear-setaceous, 4–5.5 × 0.5–0.7 mm, glandular-hairy, 3—nerved, greenish, rigid, apex acute; sheath membranous, 0.2–0.3 mm, glandular-hairy. Inflorescence terminal, densely glandular-hairy, 1–3-flowered, flowers pedicellate. Bracts lanceolate, 5.5–7 × 0.9–1.2 mm, as long as calyx; edge with ciliae 0.1–0.15 mm long; apex acute. Pedicels 1–2 mm long, densely glandular-hairy. Calyx tubular, densely glandularhairy, 5.5–7 × 1.3–1.7 mm, with 5 projecting ribs; teeth lanceolate, 1–1.5 mm long, apex acute. Petals linear-oblong, 7–8.5 × 0.8–1.2 mm, 1.5 times longer than calyx, entirely white, without purple veins; apex emarginate; base cuneate. Stamens 10, filaments 4–6 mm. Styles 2, 4–6 mm long. Capsule oblong, 5–6.5 × 1.3–1.7 mm, 2–4-seeded, usually as long as calyx. Seeds comma-shaped, with prominent radicle, 0.9–1.1 × 0.6–0.8 mm, brown.</p><p>Phenology: May–July, 1500 m, tuffaceous volcanic rocks.</p><p>General Distribution: Endemic. Irano Turanian element.</p><p>Distribution for Türkiye: Aegean region.</p><p>Notes:— The taxon was firstly defined with the name Bolanthus aziz-sancarii by Koç, Hamzaoğlu and Büyük in 2019 (Koç et al. 2019). It was collected in Afyonkarahisar province is similar to B. thymoides Hub.-Mor. in having stems completely covered with eglandular and glandular hairs, lax cushions, leaves that are 3–6 mm long, and linear-oblong petals. However, it differs from this taxon by having an imbricate leaf arrangement, calyx 5.5–7 mm long, petals 7–8.5 mm long, entirely white petals, and an inflorescence with 1–3 pedicellate flowers (versus calyx 4–5 mm long, petals 5– 6 mm long, petals white with purple veins, and an inflorescence with 4–8 sessile or subsessile flowers in B. thymoides). In addition, while B. thymoides occurs in serpentinized stony fields of the Mediterranean phytogeographical region, the new species is found in fissures in tuffaceous volcanic rocks in the IranoTuranian phytogeographical region. Also, due to overgrazing, the habitat of this speciesis is under threat, it has been recommended to be placed in the EN B1a category (IUCN 2017). Its distribution in Türkiye was given in Figure 16.</p><p>Specimens examined:— Bolanthus thymoides Hub.-Mor. Türkiye. Bilecik: Bozüyük, 740 m, 07 Jun 1975, Çetik (KNYA, No: 1239); Muğla Sandras mountain, NW of Çiçekbaba hill, Serçegediği site, 1850 m, 14 July 1979, Özhatay (ISTO, No: 23928) ; Burdur: Altınyayla, Dirmil pass, Esenli plateau, 1610 m, 06 Jun 1996, Özhatay &amp; Özhatay, Duman ¨ (ISTE, No: 72219) ; Dirmil, hills above Dirmil, 1750–1800 m, 08 Jul 1993, Bekat &amp; Leblebici (EGE, No: 18864) ; Altınyayla, Dirmil pass, 605 m, 20 Jun 2002, Dönmez 10922 &amp; Dönmez (HUB) ; Çamlıyayla, Dirmil pass, 1650 m, 30 Jun 1996, Aytaç 7405 (GAZI) ; YeŞilova, Salda town, EŞeler mountain, 2000 m, 18 Aug 2014, Koç 1848 &amp; Hamzaoğlu (Bozok Univ. Herb.) ; Antalya: Elmalı, Çağlıkara, 1540 m, 28 Jun 1975, Çetik (KNYA, No: 1238) .</p></div>	https://treatment.plazi.org/id/E5727F10FF80FFBB14A3FABC56E488D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF80FFBB14A3F8FC51678DF8.text	E5727F10FF80FFBB14A3F8FC51678DF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus huber-morathii	<div><p>Rediscovering Bolanthus huber-morathii</p><p>This species, firstly described by Simon (1977), was not reported by Davis et al. (1988) in Flora of Turkey and The Aegean Islands. In Checklist 4, only the type address information is given, but ıts description, distribution, and habitat features are not included (Özhatay et al. 2009). We here recorded for the first time in Turkey Bolanthus huber-morathii . In addition, samples were collected outside the type address for the first time in Kütahya province.</p><p>Specimens exemined:— TÜRKİYE. C2 Denizli: The South of Kartal Lake above, flats close to the shed, serpantine areas, 1960 m, 17 July 2016, Koç 2343 (BOZOK Üniv. Herb.).</p><p>Additional specimens examined (paratypes):— Bolanthus thymoides Hub.-Mor. Türkiye. Bilecik: Bozüyük, 740 m a.s.l., 7 Jun 1975, Çetik (KNYA, no. 1239) ; Burdur: Altınyayla, Dirmil pass, Esenli plateau, 1610 m a.s.l., 6 Jun 1996, Özhatay &amp; Duman (ISTE, no. 72219) ; Burdur: Dirmil, hills above Dirmil, 1750–1800 m a.s.l., 8 Jul 1993, Bekat &amp; Leblebici (EGE, no. 18864) ; Burdur: Altınyayla, Dirmil pass, 605 m a.s.l., 20 Jun 2002, Dönmez 10922 &amp; Dönmez (HUB) ; Burdur: Çamlıyayla, Dirmil pass, 1650 m a.s.l., 30 Jun 1996, Aytaç 7405 (GAZI) ; Burdur: YeŞilova, Salda town, EŞeler Mountain, 2000 m a.s.l., 18 Aug 2014, Koç 1848 &amp; Hamzaoğlu (Bozok Univ. Herb.) ; Antalya: Elmalı, Çağlıkara, 1540 m a.s.l., 28 Jun 1975, Çetik (KNYA, no. 1238) .</p></div>	https://treatment.plazi.org/id/E5727F10FF80FFBB14A3F8FC51678DF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
E5727F10FF80FFB614A3FFA857A28E6C.text	E5727F10FF80FFB614A3FFA857A28E6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolanthus stenopetalus Hartvig & Strid, Bot. Jahrb. Syst.	<div><p>Bolanthus stenopetalus Hartvig &amp; Strid, Bot. Jahrb. Syst. 108: 318 (1987).</p><p>Type: — [ Turkey C2 Muğla]. Sandras Dağ, W. Side of the summit area, 2100–2200 m, snowbed meadows and rocky slopes, serpentine, lat. 37˚04’ N, long. 28˚50’ E, 7 July 1984, Hartvig, Seçmen &amp; Strid 23375 (holotype C; isotypes: B, E! EGE, G, G00226570 [photo!]) .</p><p>Description (Figs. 17–19): — Perennial herbs. Stems forming lax cushions, 5–50-piece, 8–15 cm, prostrate, base 0.7–1 mm in diam, entirely covered with glandular hairs. Stems, 3–6-noded, internodes 5–15 mm, glandular-hairy. Leaves linear-oblanceolate, 3–5 × 0.5–0.7 mm, glandular-hairy, 3—nerved, greenish, not adpressed to stem, apex acute; sheath membranous, 0.2–0.3 mm, glandular-hairy. Inflorescence in terminal and lateral, glandular-hairy, 3–15-flowered, flowers pedicellate. Bracts linear-lanceolate, 4–5.5 × 0.5–0.7 mm, as long as inflorescence; edge membranous with ciliae 0.1–0.15 mm long; apex acute. Pedicels 2–4 mm long, glandular-hairy. Calyx tubular, glandular hairy, 3–3.5 × 0.9–1.1 mm; teeth narrowed lanceolate, 1–1.3 mm long, edge ciliate, apex acute. Petals oblanceolate, 4–5 × 0.6–1 mm, 1.5 times longer than calyx, limb pink, petal veins dark pink; apex truncate; base cuneate. Stamens 10, filaments 3–4 mm. Styles 2, 3–4 mm long. Capsule oblong, 2.5–3 × 0.9–1.1 mm, 2–4-seeded, usually as long as calyx. Seeds comma-shaped, side and dorsal face with tuberculate, granulate, 1–1.1 × 0.8–0.9 mm, brown.</p><p>Phenology: July–August, 1340–2400 m, serpantine and rocky slopes.</p><p>General Distribution: Endemic. East Mediterranean element.</p><p>Distribution for Türkiye: Aegean region.</p><p>Notes:— The taxon was firstly defined with the name Bolanthus stenopetalus by Hartvig and Strid in 1987 (Hartvig &amp; Strid, 1987). The type adress of taxon is Muğla Sandras mountain and also known that there is a taxon Bolanthus frankenioides in this area (Huber-Morath 1967). The type specimen of taxon and samples collected in field studies on Muğla Sandras mountain have been examined in detail. The article is published by Hartvig and Strid and detailed descriptions have been gathered. The articles and detailed descriptions published by Hartvig and Strid were combined and it was determined that they were ciliated and long-leaved, having short pedicellate flowers, petals pink and oblanceolate, ovary with 8–10 ovules which were given as a difference by the researchers, and that their characters were already features of the Bolanthus frankenioides var. fasciculatus taxon. Fort this reason, it was decided to select the species B. stenopetalus as synonym to the B. frankenioides var. fasciculatus . Its distribution in Türkiye was given in Figure 19.</p><p>Specimens exemined:— TÜRKİYE. C2 Muğla: Köyceğiz, Sandras Mountain, 1900–2000 m, 18 July 1968, Pamukçuoğlu &amp; Quezel (HUB-03414) ; Köyceğiz, Yayla Village, above Gökçeova Lake, Sandras Mountain, 2030 m, 05 August 2015, Koç 2196 &amp; Hamzaoğlu (Bozok Üniv. Herb.) ; Muğla: Köyceğiz Yayla (Ağla) Village, Sandras Mountain, subalphin meadows, serpentine, 1950 m, 15 August 2014, Koç 1840 &amp; Hamzaoğlu (Bozok Üniv. Herb.) .</p></div>	https://treatment.plazi.org/id/E5727F10FF80FFB614A3FFA857A28E6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Yildizbaş, Avni;Koç, Murat	Yildizbaş, Avni, Koç, Murat (2025): The taxonomic revision of genus Bolanthus (Caryophyllaceae) in Türkiye. Phytotaxa 696 (1): 28-54, DOI: 10.11646/phytotaxa.696.1.2, URL: https://doi.org/10.11646/phytotaxa.696.1.2
