taxonID	type	description	language	source
22ADAF935CB2508DBC890A645C4F5A6C.taxon	description	Figs 3, 4, 5, 6	en	Pham, Anh Van, Van Hoang, Chung, Tapley, Benjamin, Nguyen, Luan Thanh, Nguyen, Hanh Huu, La, Toi Van, Ziegler, Thomas, Rowley, Jodi J. L., Nguyen, Truong Quang, Le, Minh Duc (2025): Amolops cuongi (Amphibia, Anura, Ranidae), a new species from the Hoang Lien Range, Vietnam. ZooKeys 1256: 235-257, DOI: 10.3897/zookeys.1256.158846
22ADAF935CB2508DBC890A645C4F5A6C.taxon	diagnosis	Diagnosis. Amolops cuongi sp. nov. from the Hoang Lien Range is assigned to the A. mantzorum species group based on the absence of a dorsolateral fold and the absence of a circummarginal groove on the first finger (Fei et al. 2009, 2017). The new species is also supported as a member of the A. mantzorum group based on the molecular analyses (Fig. 2). Amolops cuongi sp. nov. is distinguishable from its congeners by a combination of the following morphological characteristics: (1) size small (SVL 33.9 – 36.9 mm in males; 37.9 – 44.4 mm in females); (2) head longer than wide; (3) vomerine teeth absent or weakly developed; (4) snout short (SE / SVL 0.15 – 0.17 in males; 0.14 – 0.16 in females); (5) tympanum small, round (TD / ED 0.24 – 0.37 in males; 0.26 – 0.35 in females); (6) the absence of a circummarginal groove on the first finger; (7) width of disc of finger III larger than tympanum; (8) supratympanic fold present; (9) dorsolateral fold absent; (10) webbing formula I 0 – 1 II 0 – 1 III 0 – 1 IV 1 – 0 V; (11) the presence of a band of small spinules and / or tubercles running from below nares, along upper lip, around lower half of eye, between tympanum and eye and rear axis of mandibles; (12) granular skin on flanks and ventral surfaces of body; (13) in life, dorsal body colouration of dark brown with diffuse-edged blotches of bluish grey, copper and yellowish green or pale green and copper; (14) ventral surface of throat, chest and belly pale cream with white dots; (15) males without vocal sacs; and (16) nuptial pad velvety without spines.	en	Pham, Anh Van, Van Hoang, Chung, Tapley, Benjamin, Nguyen, Luan Thanh, Nguyen, Hanh Huu, La, Toi Van, Ziegler, Thomas, Rowley, Jodi J. L., Nguyen, Truong Quang, Le, Minh Duc (2025): Amolops cuongi (Amphibia, Anura, Ranidae), a new species from the Hoang Lien Range, Vietnam. ZooKeys 1256: 235-257, DOI: 10.3897/zookeys.1256.158846
22ADAF935CB2508DBC890A645C4F5A6C.taxon	description	Description of holotype. Adult male; SVL 36.0 mm; head broad and flat (HD / SVL 0.14), longer than wide (HL 12.2 mm, HW 11.8 mm, HL / SVL 0.34, HW / SVL 0.33); snout round anteriorly in dorsal view (SE / SVL 0.15), projecting beyond lower jaw; snout length greater than eye diameter (SE 5.0 mm, ED 5.3 mm); nostril lateral, closer to eye the than to snout tip (SND 3.0 mm, END 2.3 mm); canthus rostralis distinct; loreal region slightly concave; eyes large (ED / HL 0.43, ED / SE 1.0); pupil horizontally oval; internarial distance as wide as interorbital distance (IND 4.2 mm, IOD 4.2 mm), larger than upper eyelid width (UEW 2.9 mm); tympanum distinct, round, small (TD 1.5 mm, TD / ED 0.28); vomerine teeth absent; tongue cordiform, notched posteriorly; body gracile. Fore limbs robust; relative finger lengths: I <II <IV <III; fingers without webbing; tips of fingers expanded into discs, II – IV with circummarginal grooves; tip of first finger smaller, without circummarginal groove; width of disc of finger III greater than the diameter of tympanum (TD 1.5 mm, FTD 2.5 mm, TD / FTD 0.60); subarticular tubercles at base of fingers II – IV round, others indistinct; inner metatarsal tubercle oval, elongate, indistinct; outer metatarsal tubercle absent, finger I with a large nuptial pad, elongate, ~ 2 / 3 the length of finger. Hind limbs long; tibia longer than thigh (FL 17.9 mm, TL 20.2 mm); relative toe length I <II <III <V <IV; tips of toes expanded into discs; width of disc of toe IV narrower than that of finger III (FTD 2.5 mm, HTD 1.6 mm); webbing formula I 0 – 1 II 0 – 1 III 0 – 1 IV 1 – 0 V; subarticular tubercles oval, formula 1, 1, 2, 3, 2; inner metatarsal tubercle elongate; outer metatarsal tubercle absent; tibiotarsal articulation reaches the nostril.	en	Pham, Anh Van, Van Hoang, Chung, Tapley, Benjamin, Nguyen, Luan Thanh, Nguyen, Hanh Huu, La, Toi Van, Ziegler, Thomas, Rowley, Jodi J. L., Nguyen, Truong Quang, Le, Minh Duc (2025): Amolops cuongi (Amphibia, Anura, Ranidae), a new species from the Hoang Lien Range, Vietnam. ZooKeys 1256: 235-257, DOI: 10.3897/zookeys.1256.158846
22ADAF935CB2508DBC890A645C4F5A6C.taxon	distribution	Distribution. Amolops cuongi sp. nov. is currently known only from the Hoang Lien Range in northern Vietnam, including Tam Duong, Lai Chau Province and Sa Pa and Bat Xat, Lao Cai Province (Fig. 1).	en	Pham, Anh Van, Van Hoang, Chung, Tapley, Benjamin, Nguyen, Luan Thanh, Nguyen, Hanh Huu, La, Toi Van, Ziegler, Thomas, Rowley, Jodi J. L., Nguyen, Truong Quang, Le, Minh Duc (2025): Amolops cuongi (Amphibia, Anura, Ranidae), a new species from the Hoang Lien Range, Vietnam. ZooKeys 1256: 235-257, DOI: 10.3897/zookeys.1256.158846
22ADAF935CB2508DBC890A645C4F5A6C.taxon	etymology	Etymology. The new species is named after Dr. Cuong The Pham, our colleague from Institute of Biology, Vietnam to honour his contributions to the herpetological research in Vietnam, particularly in the taxonomy of the anuran species complexes. For the common names we suggest Cuong’s Torrent Frog (English) and Ếch bám đá cư ờng (Vietnamese).	en	Pham, Anh Van, Van Hoang, Chung, Tapley, Benjamin, Nguyen, Luan Thanh, Nguyen, Hanh Huu, La, Toi Van, Ziegler, Thomas, Rowley, Jodi J. L., Nguyen, Truong Quang, Le, Minh Duc (2025): Amolops cuongi (Amphibia, Anura, Ranidae), a new species from the Hoang Lien Range, Vietnam. ZooKeys 1256: 235-257, DOI: 10.3897/zookeys.1256.158846
