identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F541C527FF93FF96A4709CD39933FA02.text	F541C527FF93FF96A4709CD39933FA02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia aporophylla Rchb. f.	<div><p>Oberonia aporophylla Rchb.f. — Fig. 1</p><p>Oberonia aporophylla Rchb.f. (1855) 223. — Syntypes: Cuming 2113 (BM000088320, K s.n., K s.n. Drawing from type at Vienna Reichenbach Herbarium #37820 W68779. Additional pocket labelled Cuming 2112 or Cuming 2113 with flower parts and leaf sample. Agrees with drawing of Cuming 2113 on herbarium sheet. Philippines in protologue; Bohol on K syntype.</p><p>Oberonia longirepens J.J.Wood in J.J. Wood et al. (2011) 434, f. 2.214. — Type: Clemens 50250 (holo SING: not found in 8/2018, iso K 73164: not found 4/2018), [Malaysia, Sabah,] Tenompok Orchid Garden, 1500 m, syn. nov.</p><p>Notes — Oberonia aporophylla and O. longirepens are conspecific. The overall habit of erect stems with short imbri- cate leaves and creeping rhizomes (Fig. 1a, c) in conjunction with multiple floral attributes, such as the narrow petals, the presence of distinct auricles on the lip, the bilobed epichile of the lip with serrated edges (Fig. 1b, d) clearly show that the two names refer to the same species. One of the most striking characters is the creeping rhizome, typically associated with the genus Hippeophyllum Schltr. While Reichenbach (1855) did not mention this character in the protologue, it is found on historical herbarium records such as Vanoverbergh 1919, P00263280 from Luzon, the Philippines, collected in flower in October 1912, is known from field observation in the Philippines (Ronny Boos, pers. comm.), and from horticultural specimens of Philippine provenance seen by the author (Fig. 1e–k). The probability of two species with identical vegetative growth and flowers, but with and without rhizomes, is extremely unlikely.</p><p>Wood et al. (2011) did not compare O. longirepens to any species occurring outside of Borneo, and only noted the distinct creeping habit and the salmon coloured flowers. The colour of the flower can vary extensively in Oberonia species. Oberonia aporophylla is known in light cream and yellow-green forms (Fig. 1e–f), with salmon rachis (Fig. 1f). The salmon colour of the flower is well within the expected range.</p><p>The illustration for the flower of O. longirepens might erroneously suggest different proportions of the floral elements. However, the measurements of the floral elements in Wood et al. (2011) are incongruent with the drawings, both in relative and absolute terms. The sepals as well as the petals are indicated as being 0.9 mm long, but petals are markedly smaller in the drawing compared to the sepals. Based on the 1 mm scale bar, the sepals are at least 1.0 mm long, while the petal is only 0.7 mm long. The lip is described as 0.9–1 mm long, while from top of auricles to longest projection of epichile it is 0.85 mm based on the 0.5 mm scale bar provided. From that follows, that the proportion of the sepals to lip is off by 15 % in the drawing. The implied live position of the lateral sepals is equally misleading (Fig. 1d). While the drawing shows them inclined towards the petal and median sepal, they are actually pointed away at an approximately 120° angle, as shown in the much more realistic drawing on the syntype sheet of O. aporophylla at K (Fig. 1b). Notice also the absence of disc and a fortiori the sac in both drawings. It is a further example that drawings are frequently highly inaccurate, even by well-respected authors in highly regarded publications.</p><p>Unfortunately, the type specimens of O. longirepens allegedly deposited at SING and K were not found during collection visits in 2018. The K on-line data portal does not have an image of the specimen, suggesting it was never deposited there. The fate of those type specimens is unknown. The collector is given as ‘Clemens’ in the protologue, while the K data portal gives J. Clemens &amp; M.S. Clemens . As the type sheets are missing, the collector could not be verified.</p><p>A biogeographic distribution spanning from the Philippines to Borneo is well within the dispersal abilities of plants with minute seeds. The seeds of O. aporophylla are approximately 150 µm long (Fig. 1l), typical for Oberonia spp. (Barthlott et al. 2014, Geiger 2014, unpubl. data, Geiger et al. 2020).</p></div>	https://treatment.plazi.org/id/F541C527FF93FF96A4709CD39933FA02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF90FF96A4709DA1989AFBB3.text	F541C527FF90FF96A4709DA1989AFBB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia bertoldii King & Pantl.	<div><p>Oberonia bertoldii King &amp; Pantl. — Fig. 2</p><p>Oberonia bertoldii King &amp; Pantl. (1897) 581 (‘ Bertholdi ’); Holttum (1953) 222 [spelling correction, ‘ Bertoldii ’]. — Syntypes: Scortechini 1525 (repository unknown), [Malay Peninsula,] Perak.</p><p>Oberonia gracillima Ridl. (1905) 189. — Syntypes: Rostados s.n. (K 000942998, SING 0047515), Tringanu [= Terengganu,Peninsular Malaysia] at Bundi, Malaya, syn. nov.</p><p>Oberonia laeta J.J.Sm. (1927) 41, pl. 4, I. — Syntypes: Backer 31514 (repository unknown), Java, Banjoemas, Kinderzee [Banyumas Regency], syn. nov.</p><p>Notes — The three names had been treated as synonyms already by Seidenfaden (1968) but are currently still listed as distinct by WCSP (2020), for which reason these taxa are here discussed in more detail. The oldest name, O. bertoldii was introduced as O. Bertoldi [sic], but requires a double -ii ending according to ICN (2018) Art. 60.8(b). That spelling was first used by Holttum (1953), though still with the customary capitalisation of the specific epithet honouring a person. Seidenfaden (1968) was the first author to use the correct and modern spelling of bertoldii .</p><p>Oberonia gracillima is here confirmed as a synonym of O. bertoldii after examination of the SING syntype. The shared characters include the habit, the broad triangular lip with short broadly bifid epichile, deeply serrated lateral petals, and broad, serrated bract. The collector is indicated as Rostado in the protologue but should be spelled Rostados according to Van Steenis-Kruseman (2017) .</p><p>The habit of all three names are identical for all intents and purposes. King &amp; Pantling (1897) noted the very short stem, and narrowly lanceolate acuminate leaves, which perfectly describes the syntype of O. gracillima (Fig. 2d) and illustration of O. laeta (Fig. 2c).</p></div>	https://treatment.plazi.org/id/F541C527FF90FF96A4709DA1989AFBB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF90FF97A470997B9ED3FF0C.text	F541C527FF90FF97A470997B9ED3FF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia bifida Schltr.	<div><p>Oberonia bifida Schltr. — Fig. 3</p><p>Oberonia bifida Schltr. in K.Schum. &amp; Lauterb. (1905) 109. — Type: Schlechter 14081 (holo B, lost, single exemplar), on trees in the forests of the Bismarck Mountains, ~ 1000 m.</p><p>Oberonia celebica Schltr. (1911b) 23. — Syntype: Schlechter 20474 (B, lost),[Sulawesi, Minahassa,] on trees in the forest of the Gunong Masrang, 1200 m, syn. nov.</p><p>Oberonia fissiglossa N. Hallé (1977) 274, carte 73, f. 115. — Type: MacKee 26809 (holo P00081700;flowers originally in alcohol, dry mounted on herbarium sheet),[New Caledonia,] Ponerihouen, Mt Auoupini, 900–1000 m, syn. nov.</p><p>Notes — Oberonia bifida is a fairly common species in the Central and Western Pacific Islands. It is characterised by its caulescent habit with long falcate leaves and orange flowers, acuminate bract, narrow petals, and lip with auricles, small but deep sac, darker orange disc, moderate constriction in mesochile, and bifid epichile.All those characters are shared between the correct name O. bifida, and two synonyms: O. celebica and O. fissiglossa . Schlechter (1911b) described his O. celebica with yellow-brown (= orange) flowers, and did not differentiate it from any other species. Hallé (1977) compared his O. fissiglossa only to O. equitans (G.Forst.) Mutel, and noted the brown buds. The illustrations either from the protologues or by the respective author of the three species show no differences (Fig. 3a–c).</p><p>The SEM images (Fig. 3d–i) reveal some variability in the degree of the constriction of the mesochile and the erose tips of the epichile lobes. The specimen in Fig. 3d–e is somewhat larger than the others, which may be attributable to it having been grown in cultivation. There is natural variability in size, and the less than 10 % difference seems well within natural variability.</p></div>	https://treatment.plazi.org/id/F541C527FF90FF97A470997B9ED3FF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF91FF92A4709C1D9A45FEED.text	F541C527FF91FF92A4709C1D9A45FEED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia brunoniana Wight	<div><p>Oberonia brunoniana Wight — Fig. 4</p><p>Oberonia brunoniana Wight (1851) 3, pl. 1622. — Lectotype (designated by Geiger 2019): Wight s.n. (lecto K 000387708), India, Coimbatore,Iyamally Hills, Mount Agamullu. Wight 2914 (paralecto K 000387707), Mount Paulghautcherry.</p><p>Oberonia wallichii Hooker (1888) 681. — Syntype: De Silva s.n. (repository unknown), [India,] Silhet, syn. nov.</p><p>Oberonia saintberchmansii Kad.V.George &amp; J.Mathew in Kad.V. George et al. (2019) 110, f. 1 (‘ saint-berchmansii ’). — Type: K. V. George &amp; S. Antony 0126 (holo RHK;iso MSSRF), [India,] Kerala, Idukki District,way to Nedukandam, Cardomom Hills, 3rd mile,alt. 1310 m, Jan. 2014, syn. nov .</p><p>Notes — The repository of the type for Oberonia wallichii is unknown. Specimens in K, K 000387713 Wallich 1948, and K 000387690 s. coll. are not from the type locality nor by the indicated collector, thus they do not seem to be types. A specimen, Lobb s.n. K s.n., from Khasiya Hills identified by Lindley as O. lindleyana (= O. brunoniana) was identified by Hooker as O. wallichii, hence, constitutes an ideotype (Fernald 1939). The associated drawing of the flower on the ideotype shows a typical O. brunoniana morphology.</p><p>This species has been overlooked and confused. For instance, Ansari &amp; Balakrishnan (1990) in their monograph of Indian Oberonia, did not mention O. wallichii at all. The species was distinguished by Hooker (1888) by the smaller flower of 1/ 20 inch = 1.25 mm, which would be on the smaller side of any known Oberonia species, and would be extremely small for species in Ansari &amp; Balakrishnan’s (1990) Section III. Those small-flowered specimens from Khasia in NE India were contrasted to the more typical larger-flowered ‘peninsular’ O. brunoniana s.str., i.e., those of SW India. It is interesting to note that the alleged smaller flower size has been repeated by reference to Hooker (1888), but has not been independently confirmed. The flower size seems to vary along a north-south gradient. The dimensions for O. saintberchmansii from Karnataka (N9°, see below) are 3.5 mm, the flower from Myosore (N12°) shown in Fig. 4d measures 2.4 mm, while specimens from Mumbai (N19°) were indicated as &lt;2 mm in size (Santapau &amp; Kapadia 1960), and Hooker’s (1888) indication of 1.25 mm from Khasia (N24°).Accordingly, a latitudinal size gradient becomes evident. Without either a clear geographic break, or a distinctly bimodal size distribution, once specimens are sampled across the geographic range, no taxonomic separation seems justifiable for now. The observed differences appear to be ecophenotypical variation.</p><p>In the absence of any distinguishing characters, and the contradictory identifications of the same plant with various synonyms of O. brunoniana, O. wallichii is a further synonym of O. brunoniana .</p><p>Oberonia saintberchmansii is a clear synonym of O. brunoniana . The shared characters include the lip with broad lateral lobes, two relatively small epichile lobes, a well-defined smooth disc, narrow petals, overall fan-shaped vegetative habit of the plant. Oberonia brunoniana is well-known from Kerala (e.g., Ansari &amp; Balakrishnan 1990), the type locality of O. saintberchmansii . There is no variant character of any kind. George et al. (2019) compared their new species only to O. falconeri, a species outside of Section III of Ansari &amp; Balakrishnan (1990) to which O. brunoniana and allied species belong. In fact, George et al. (2019) did not even cite Ansari &amp; Balakrishnan (1990), a key work on Oberonia of India. Schuiteman (pers. comm.) has indicated the synonymy previously. Additional synonyms of O. brunoniana were recently discussed by Geiger (2019).</p></div>	https://treatment.plazi.org/id/F541C527FF91FF92A4709C1D9A45FEED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF94FF92A73F9C3F9A1EFC46.text	F541C527FF94FF92A73F9C3F9A1EFC46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia costeriana J. J. Sm.	<div><p>Oberonia costeriana J.J.Sm. — Fig. 5</p><p>Oberonia costeriana J.J.Sm. (1905) 244. — Syntypes: Blume s.n. (L 0061737), [Indonesia, Java] Mount Gede, at the water fall, parasitic; De Monchy s.n. (repository unknown), [Indonesia, Java,] Krawang; Junghuhn s.n. (L 0063834); J.J. Smith s.n. (repository unknown), [Indonesia,] Pekalongan, Djolotigo; J.J. Smith s.n. (repository unknown), [Indonesia,] Sumatra; unknown collector s.n. (repository unknown), Indonesia, Java, Buitenzorg [= Bogor], Bobodjong.</p><p>Oberonia vulcanica Schltr. (1911b) 24; (1934) pl. 18, f. 96. — Syntype: Schlechter 20509 (B, lost), [Indonesia,] on trees in the forest of the Gunong Mahawo ( Minahassa), Sulawesi, 1200 m, syn. nov.</p><p>Notes — Oberonia costeriana has several synonyms (Geiger 2019), such as O. elmeri Ames (Fig. 5b), O. kinabaluensis Ames &amp; C.Schweinf. (Fig. 5d), O. obesa Ames (Fig. 5c), and O. gigantea Fukuy. Here one further taxon, O. vulcanica, is identified as conspecific with O. costeriana (Fig. 5e). The shared characters include the trapezoidal lateral lobes with downward pointed tips, the bifid epichile, the rhombic petals and the acuminate acute floral bract with erose margins, and the dirty orange colour (fox-brown in Schlechter 1911b: 24). The description of the habit also matches O. costeriana (acaulescent with falcate-linear leaves).</p></div>	https://treatment.plazi.org/id/F541C527FF94FF92A73F9C3F9A1EFC46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF94FF92A73F9EE89F2DFCBC.text	F541C527FF94FF92A73F9EE89F2DFCBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia ensiformis (Sm.) Lindl.	<div><p>Oberonia ensiformis (Sm.) Lindl. — Fig. 6</p><p>Oberonia ensiformis (Sm.) Lindl. (1859) 4. — Malaxis ensiformis Sm.(1812) Malaxis #14, [p. 180]. — Lectotype (first step designation by Ansari &amp; Balakrishnan (1990:27): Hamilton-Buchanan s.n. (LINN-HS 1396.11.1 –2), Nepal, Narainhetty. Buchanan s.n. (questionable isolecto BM000088250), Nepal.</p><p>Oberonia vesiculifera Aver. in Aver. et al. (2019) 151, f. 1292. — Type: AL 965 /1 (holo LE 01048682), 300–530 m, [Vietnam,] Nam No Npa [inferred from title]; AL 920 (para LE 01048684), 400–600 m, [Vietnam,] Nam No Npa [inferred from title], syn. nov.</p><p>Notes — The identity of O. ensiformis is rather uncontroversial, despite the fact that the protologue did not contain any illustrations and that images of the type are not available. The written account of the species by Smith (1812) describing it having sword-shaped leaves with basal abscission scar and dull orange flowers with ovate sepals and petals and a fourlobed lip only matches one species occurring in India, namely what is commonly referred to O. ensiformis (e.g., Ansari &amp; Balakrishnan 1990). Despite the long chresonymy for the species, there are surprisingly few herbarium specimens, with not a single fluid preserved sample in any institutional repository visited or contacted.</p><p>Ansari &amp; Balakrishnan’s (1990: 27) reference to “Type: Hammilton-Buchanan s.n. ... (LINN)” constitutes a first-step lectotypification. Averyanov (2013) cited LINN as holotype, most likely not aware of the two sheets.</p><p>Averyanov et al. (2019) distinguished O. vesiculifera from O. ensiformis solely on the basis of vesicles or calli above the constriction in the middle of the lip. The presence of those calli has been documented in O. ensiformis, for which reason O. vesiculifera is an obvious synonym of O. ensiformis . It is unclear what Averyanov considered ‘ ensiformis ’. He did not refer any of his previously illustrated O. ensiformis with calli (e.g., CPC 1887 Averyanov 2013: f. 76b,c) to his new taxon.</p><p>The reporting and illustration of those calli is rather variable. They can be recognized in the photograph of Ansari &amp; Balakrishnan (1990: pl. 2b) from India, Raskoti (2009: 191, text-fig.) from Nepal, and Averyanov (2013: f. 76c) from Vietnam. Seidenfaden &amp; Smitinand (1959), Seidenfaden (1968, 1978), Abraham &amp; Vatsala (1981), Pradhan (1979), Ho (1993), Chen et al. (1999), and Lucksom (2007) show them clearly on their drawings. However, usually only a single callus is evident. The otherwise exquisite colour drawing in Hooker (1858: f. 5), though misidentified as O. acaulis, and King &amp; Pantling (1898: pl. 9) also alluded to them. The drawings in Ansari &amp; Balakrishnan (1990) and Averyanov (2013: f. 77h) do not show a trace of those calli, but the written description in both referred to pairs of minute calli. Given the numerous demonstrated inaccuracies in drawings of Oberonia flowers (Geiger 2019) I consider those inconsistencies are problems with drawings rather than representing additional species. SEM images always show the dual calli. The claim that the species with dual calli is microendemic to the type locality in Vietnam is demonstrably false.</p><p>Misra (2004) considered the specimens from Orissa to be distinct at the species level because of slightly different distribution of the calli on the lip. Such a statement cannot be found in the subsequent edition (Misra 2014), which suggests that he reconsidered his earlier statement and regarded the observed differences to lie within intraspecific variability. Given the rather slight nature of differences, the details of callus morphology are taxonomically insignificant.</p></div>	https://treatment.plazi.org/id/F541C527FF94FF92A73F9EE89F2DFCBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF94FF92A4709E6C9ED3F9A1.text	F541C527FF94FF92A4709E6C9ED3F9A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia griffithiana Lindl.	<div><p>Oberonia griffithiana Lindl. — Fig. 7</p><p>Oberonia griffithiana Lindl. (1838) t. 1779/t8B. — Syntypes: Griffith s.n. ( Griffith 355 K 00097422), Burma [= Myanmar], Moulmein [= Mawlamyine].</p><p>Oberonia khuongii Aver.&amp; V.C.Nguyen in Aver. et al. (2019) 185, 188, f. 8. — Type: Van Canh Nguyen AL 437 (holo LE 01042183),[Vietnam,] Dak Nong province, Dak Song district, ~ 700 m, syn. nov.</p><p>Notes — Averyanov et al. (2019: 185) referred to their figure 8 as a ‘digital epitype’. As the figure is part of the protologue and original material, it cannot be an epitype per ICN (2018) Art. 9.9 and glossary.</p><p>Oberonia khuongii is identical to O. griffithiana . Shared characters include the caulescent habit, the narrow petals with irregular edges, the lip with multiply incised lateral lobes, the bifid epichile, and the granular surface of the lip. The species was mainly compared to O. cavaleriei Finet with terete leaves. Oberonia griffithiana was mentioned in the remarks, but no discriminating characters were given. Averyanov et al. (2019: f. 8) illustrated some interesting colour morphs. The red dots on the leaves are taxonomically not meaningful. A similar case is found in O. punctata J.J.Sm. (with dots) and O. subanajamensis J.J.Sm. and O. caprina Gilli without (Geiger 2019).</p><p>It is interesting to note that the authors of O. khuongii were well aware of the identity of the specimen as O. griffithiana through personal communication by the present author, but introduced the new name anyway without further discussion or justification.</p></div>	https://treatment.plazi.org/id/F541C527FF94FF92A4709E6C9ED3F9A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF94FF9CA4709B8B992BF7C2.text	F541C527FF94FF9CA4709B8B992BF7C2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia heliophila Rchb. f.	<div><p>Oberonia heliophila Rchb.f. — Fig. 8, 9</p><p>Oberonia heliophila Rchb.f. (1878) 56. — Lectotype (designated by Kores 1989): U.S. Exploring Expedition s.n. (lecto W37726), Fiji, Vanua Levu, Mountains of Mathuata Province. Paralectotypes: U.S. Exploring Expedition s.n. (W Reichenbach 37727 W68809), Samoa, Ovalau; U.S. Exploring Expedition s.n. (US 37773) Samoa; Seemann 587 (AMES 00101970, BM000088475, K 00079501,K-L s.n., P [?: see notes], W 37711), [Fiji,] Viti Levu, Tailevu Province, Namara; Graeffe s.n. (repository unknown), Upolu, Samoa; Whitmee s.n. (BM 000082108, K 00079502), [Samoa,] Sawai’i.</p><p>Oberonia asperula J.J.Sm. (1908) 30. — Syntypes: Versteeg 1322 (BO, L0061733), [Indonesia,] S. New Guinea (L type label), Noord River, at foot of Nepenthes Hill (protologue), on trees in Metroxylon swamps (Smith 1909), syn. nov.</p><p>Oberonia rivularis Schltr. in K.Schum. &amp; Lauterb. (1905) 115. — Syntypes: Schlechter 13801 (B, lost,K 00079467 [no flowers]),[Papua New Guinea,] at the riverboard of the Nuru mid-section, ~ 200 m, syn. nov.</p><p>Oberonia (Sect. Scytoxiphium) inversiflora J.J.Sm. (1912) 487 (‘ inversiflorum ’). — Type: Lorentz Exp. 1909, Rachmat live plant number 431 (syn BO Bogor cult.). Exp. s.n. (possible syn L 0061748), s. coll. (possible syn BO 0071552,L 0727884,U.1459111,U.1459112), without locality ‘Cult in Hort. Bog.’signed by J.J. Smith, Dutch New Guinea, hills near Alkmaar, syn. nov.</p><p>Oberonia hosokawai Fukuy. in Hosok. (1941) 290 ( ‘ hosokawae ’). — Syntypes: Hosokawa 9599 (syn AMES 67290 /Harvard Barcode 114495, L0061745,MICH 1115255, TAI 118740,US 2036596), Micronesia,Ponape, Mount Asama-san, 500–800 m (protologue), US label 500– 200 m, syn.nov .</p><p>Notes — The types of O. heliophila were treated in detail by Kores (1989). His cited ‘isolectotype’ AMES 3034 is not recorded in the AMES herbarium catalogue. AMES 3034 was cited for multiple localities, which suggest that multiple specimens from several different gatherings were mounted on one herbarium sheet. Given the large size of the plant, an alternate interpretation is that AMES 3034 does not refer to a specimen registration number, but an accession number of an entire collection. A specimen of Seemann 587 at P according to Kores (1989) could not be found in the P collection or catalogue.</p><p>Oberonia asperula is a synonym of O. heliophila . The shared characters (Fig. 8e) include the very large plant size in the genus, the acaulescent habit and long acuminate, fleshy leaves. The flower with the thick trapezoidal lip, the narrow oblong petals, the broad, reflexed and pubescent sepals, the pubescent pedicelled ovary, the broad strongly double incised bracts, the very short column, and the green coloration of the flower.</p><p>Oberonia rivularis is a clear synonym of O. heliophila after examining syntype material of both taxa. The large size of the plant, the flowers in well-separated whorls, the green lip with tan remaining tepals, the broad ovate-acuminate lip with serrated margins all support the synonymy (Fig. 8c). Schlechter in Schumann &amp; Lauterbach (1905) noted that O. rivularis is the smallest of the section Scytoxiphium Schltr. in New Guinea, a statement copied by O’Byrne (1994). It is unclear what may have led Schlechter to such a statement because in terms of vegetative size, it is amongst the largest species.</p><p>The possible syntypes of O. inversiflora in L are difficult to as- sess. Only one specimen identified by J.J. Smith as O. inversiflora (L 1513897) can positively be excluded, because of a collection number mismatch (C 278 on sheet, Rachmat 431 in protologue). The other specimens can neither be confirmed nor excluded. The 1912 and 1913 Soeda Sublimaat [= Soda sublimate = mercury chloride] dates are pest treatment dates, and not collection dates.</p><p>Oberonia inversiflora shares multiple characters with O. heliophila (Fig. 8d). It is an unusually large plant for the genus with leaves to 40 cm long and has a very compressed vegetative axis. The flowers share the deeply incised bract, the very hairy pedicelled ovary, the few hairs on the back of the sepals, the straw to pale brown sepals, the thick, light green lip with small triangular side lobes and broad ovate apical lobes. There are no characters that could support a separate status. The illustration of O. inversiflora by Smith (Fig. 8d) shows somewhat wider petals. Those fall within the range of variability found by examination of multiple flowers by SEM (Fig. 9), accordingly, are taxonomically meaningless. Oberonia heliophila is known from Papua New Guinea. Accordingly, the two species are synonyms.</p><p>The spelling of the species introduced as O. hosokawae [sic] has to be corrected. The species was named in honour of the collector of the plant Takahide Hosokawa, a man. In accordance with ICN (2018) Art. 60.8a, the spelling of the name is here corrected from O. hosokawae to O. hosokawai . There are an additional eight species epithets that need to be corrected as well: Carpesium hosokawae Kitam. ( Asteraceae), Glochidion hosokawae Fosberg ( Phyllanthaceae), Pandanus hosokawae Kaneh. ( Pandanaceae), Psychotria hosokawae Fosberg ( Rubiaceae), Eria hosokawae A.D.Hawkes &amp; A.H.Heller, Microtatorchis hosokawae Fukuy. (now Taeniophyllum hosokawae (Fukuy.) L.O.Williams), Habenaria hosokawae Fukuy., Moerenhoutia hosokawae (Fukuy.) Tuyama (all Orchidaceae). There is one properly formed name, Cirsium hosokawai Kitam. ( Asteraceae). The type repository for O. hosokawai was not specified in the protologue, for which reasons all specimens of the type gathering are syntypes.</p><p>Oberonia hosokawai is a synonym of O. heliophila . The shared characters include the very large size of the plant with slightly falcate leaves in an acaulescent fan, the broad floral bract with serrated margin, the narrow acuminate petals, the thickened lip with broad ovate basiochile, short and thick column, hairs on fruit (MICH 1115255: Fig. 8b). Further supporting evidence includes that Hawkes (1952) placed the species in the section Scytoxyphium, to which O. heliophila belongs.</p></div>	https://treatment.plazi.org/id/F541C527FF94FF9CA4709B8B992BF7C2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF9AFF9EA47099D29AB2FE07.text	F541C527FF9AFF9EA47099D29AB2FE07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia lycopodioides (J. Koenig) Ormerod	<div><p>Oberonia lycopodioides (J.Koenig) Ormerod</p><p>Oberonia lycopodioides (J.Koenig) Ormerod in Seidenf. (1995) 21. — Epidendrum lycopodioides J. Koenig (1791) 21. Lectotype (designated by Averyanov 2013): Koenig s.n., 5 Sept. 1779 (lecto K: not seen in 2018, not in K database; isolecotype LIV), [Peninsular Malaysia,] Malacca.</p><p>Notes — Oberonia ancipita used by Náves in Blanco (1880: 230) with ‘type’ Cuming 2115 (BM 000088321, K s.n., L.1513842, MO 125587, Philippines; Bohol on one portion of K sheet) has been identified as a synonym of O. lycopodioides (e.g., Bunpha et al. 2019, WCSP 2020). Náves referred to ‘ Malaxis ancip . Rchb.f. in Walp l. c. 214 ex Vidal ms Philipp, fide’. The entry by Náves was not intended to be an introduction of a new name as he clearly attributed the name ancipita to Lindley, and also cited ‘ Malaxis ancip . Rchb in Walp. l. c. 214’ (= Reichenbach 1861: 214). The confusion might have started with Reichenbach (1855: 223) inflecting Lindley’s species epithet anceps in his Latin text: ‘aff. O. ancipiti; quae in affinibus anciptibus Lindl.; In O. ancipiti (e.g., Cuming 2115)’. The ancip... spelling is only found in Reichenbach (1855: in Bonplandia) and not in Reichenbach (1861). It suggests that Náves’ reference to ‘Rchb.f. in Walp.’ is a further error and should have referred to Bonplandia. The mention of Cuming 2115 by Náves, copied from Reichenbach (1855) and not found in Reichenbach (1861), should be regarded as a voucher for the presence of the species in the Philippines, and not as a type designation. Oberonia ancipita is an orthographic variant of Oberonia anceps by Náves without any nomenclatural consequences. Vidal (1886: 266) referred to O. ancipita, Lindl. ? ( Malaxis anceps Reichb.f.) so wondered whether ancipita was a Lindley name, not a Náves name. It is further evidence that Náves never intended to introduce a new name, and that it was not perceived as a new name by his contemporaries. It is not clear when the orthographic variant was given the status of a distinct taxon; Merrill (1924: 307), Kress et al. (2003) and Bunpha et al. (2019) are the only known sources that use the name, in all cases as a synonym of O. anceps or O. lycopodioides . A contributing factor for the confusion could be the rarity of the work. A copy could be examined in the P library. Cuming 2115 has been confirmed to be O. lycopodioides by the present author. Náves introduced known orthographic spelling variants such as Phalaenopsis lueddemannii for P. lueddemanniana Rchb.f. Accordingly, treating Oberonia ancipita as an orthographic variant has a precedent with other ‘names’ in Náves (in Blanco 1880).</p></div>	https://treatment.plazi.org/id/F541C527FF9AFF9EA47099D29AB2FE07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF98FF9EA73F9F24993BFB54.text	F541C527FF98FF9EA73F9F24993BFB54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia obcordata Lindl.	<div><p>Oberonia obcordata Lindl. — Fig. 10</p><p>Oberonia obcordata Lindl.(1859) 7. — Syntypes: Griffith s.n. (K 000974238), [India,] Darjeeling; Gr iffith s.n. (K 000974231), [India, Meghalaya,] Khasia Myrung; Hooker f. &amp; Thomson 112 (K 000974237), [India,] Sikkim, 2000 ft.</p><p>Oberonia menghaiensis S.C. Chen (1982) 190, f. 1.1–3. — Syntypes: C.W. Wang 76269 (AMES 50102/Harvard Barcode 00101993, PE 1641349/ barcode 27299 (see Lin &amp; Yang 2015: text-fig. p. 335), PE 1641348/barcode 273000), China, Yunna, Menghai, Hei-lung-tan, 1800 m, syn. nov.</p><p>Notes — Chen (1982) referred to the gathering as a ‘holotype’, but did not specify a repository, for which reason any specimen from that gathering is a syntype. The specimen has been identified on the PE type sheets as O. obcordata in 1973. The shared characters include the caulescent habit with falcate leaves widest in the lower third of the leaf’s length, the ovate petals with erose margins (variable in the species from more club-shaped to broad-base oblong), the incised lip with downturned lateral lobes. The lip is not shown bilobed in the main figure, but the insert (Fig. 10e insert: arrows) shows at least some flowers with the more typical indentation of the apical lobe. The indentation is variable in the species as shown on the type sheets of O. obcordata (Fig. 10a, b). In conclusion, O. menghaiensis is a synonym of O. obcordata .</p></div>	https://treatment.plazi.org/id/F541C527FF98FF9EA73F9F24993BFB54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF98FF9EA73F99D69F69FAC2.text	F541C527FF98FF9EA73F99D69F69FAC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia pachyphylla King & Pantl.	<div><p>Oberonia pachyphylla King &amp; Pantl. — Fig. 11</p><p>Oberonia pachyphylla King &amp; Pantl.(1898) 5,pl. 4. — Lectotype (designated by Seidenfaden 1968: 102, inadvertent lectotype designation by reference to CAL!, type): Pantling 429 (CAL), [India,] near the base of the Sikkim-Himalaya, at Salgurra near Siliguri, 900 ft.</p><p>Oberonia janae Aver. in Aver. et al. (2015) 171, f. 12C, 14. — Type: Leong– Skornickova et al. JLS-2678 (holo SING [not found in Oct. 2018]; iso LE, VNMN), Vietnam, Binh Thuan province, Bac Binh district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.338394&amp;materialsCitation.latitude=11.493501" title="Search Plazi for locations around (long 108.338394/lat 11.493501)">Phan Son</a> com- mune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.338394&amp;materialsCitation.latitude=11.493501" title="Search Plazi for locations around (long 108.338394/lat 11.493501)">Sa Mai mountain</a>, N11°29'36.6" E108°20'18.2", 824 m, syn. nov.</p><p>Notes — Oberonia janae is a synonym of O. pachyphylla . Averyanov et al. (2015) compared the two species and noted as differentiating characters the more deeply incised serrations on the lateral lobes of the lip and the papillae/ hairs on the back of the sepals. The depth of the serration appears to be variable based on various illustrations in the literature (Seidenfaden 1968: f. 69, see Fig. 11e–g, Pradhan 1979: text-fig.; Ansari &amp; Balakrishnan 1990: f. 33, see Fig. 11g). The backside of the sepals has never been illustrated, yet Averyanov et al. (2015) did not cite voucher material for O. pachyphylla s.str. Ansari &amp; Balakrishnan (1990) described the sepals as gland dotted and illustrated it with small circles over the surface. That description was based on the lectotype, a pressed specimen. Proper glands are unknown on the flowers of Oberonia, but pressed hairs may appear like glands.Accordingly, this alleged differences has not been properly documented. Additionally, the cited difference in the erose margin of the bract is in error, as the illustration of King &amp; Pantling (1898: pl. 4, see Fig. 11c) showed that serrations clearly in the protologue of O. pachyphylla . The shared characters include the habit (Fig. 11a, h), the fleshy leaves, the subsessile flowers, the bract with erose margin of the bract (Fig. 11c, l), the overall shape of the lip with incised lateral lobes (Fig. 11b, d–g, i–k). The distribution of O. pachyphylla s.str. encompasses Vietnam and neighbouring countries (e.g., Averyanov 2013), making a microendemic cryptic species even more unlikely.</p><p>The case of O. equitans and its 10 synonyms confirmed by molecular phylogeny and SEM investigations of the flowers shows the extensive phenotypic plasticity in Oberonia spp. (Geiger et al. 2020). The variability of lateral processes is equally well documented in other species such as O. insectifera Hook.f. (Geiger 2019), as well as in O. pachyphylla discussed above.Accordingly, the variety named in Averyanov et al. (2015) does not justify species-level recognition; it falls well within intraspecific variability.</p><p>Oberonia seidenfadenii (H.J.Su) Ormerod appears to be another likely synonym of O. pachyphylla . As I have not yet been able to track the protologue in the Journal of the Experimental Forest of National Taiwan University, I cannot address it with sufficient confidence.</p></div>	https://treatment.plazi.org/id/F541C527FF98FF9EA73F99D69F69FAC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF98FF9FA470986C9A49FE06.text	F541C527FF98FF9FA470986C9A49FE06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia pachystachya Lindl.	<div><p>Oberonia pachystachya Lindl. — Fig. 12</p><p>Oberonia pachystachya Lindl. (1859) 6. — Syntypes: Catlett 37 (K s.n.; illustration of type lot in W35392). S.[outh] Sea Islands.</p><p>Oberonia trigonoglossa Schltr.(1911a) 166; (1923) pl. 65, f. 231. — Syntype: Schlechter 18840 (B, lost), [Papua New Guinea,] forests of the Bismarck mountains, 1600 m, syn. nov.</p><p>Notes — Lindley’s description is rather rudimentary; however, the illustration on the type in Lindley herbarium at K clearly shows the same species (Fig. 12). The erect habit with multiple rather short leaves on the stem, and rather short inflorescence agrees between O. trigonoglossa and O. pachystachya . Additional common characters include the acuminate acute bract (Fig. 12c, e), the ovate-oval petals with erose margin (Fig. 12b, f), and the inverted triangular lip (Fig. 12b, d), described as oval pointed by Lindley.</p></div>	https://treatment.plazi.org/id/F541C527FF98FF9FA470986C9A49FE06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF99FF99A73F9F289F5AFEF3.text	F541C527FF99FF99A73F9F289F5AFEF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia rhizomatosa J. J. Sm.	<div><p>Oberonia rhizomatosa J.J.Sm. — Fig. 13, 14</p><p>Oberonia rhizomatosa J.J.Sm. (early 1911a) 2; (1911b) 531,pl. 79A. — Syntype: Gjellerup 156 (BO), German New Guinea, Bougainville Mountains.</p><p>Oberonia repens Schltr. (11 Oct. 1911 a) 161; (1923) pl. 62, f. 221. — Syntype: Schlechter 20045 (B, lost), [Papua New Guinea,] in the forests of the Torricelli mountains, 600 m, syn. nov.</p><p>Oberonia torana J.J.Sm. (1915) 216, pl. 70, f. 122. — Syntypes: Gjellerup 777 (BO), Indonesia, Irian Jaya, Berkombor at Tor river, 25 m; Gjellerup 912 (BO, L 0061802), Indonesia, Irian Jaya, Gauthier mountains on N slope, 300 m, syn. nov.</p><p>Oberonia ponapensis Tuyama (1940) 275, pl. 2, f. i. — Type: Hatusima 11102 (holo TI 12969; iso TI 12968), Caroline Islands [Federate States of Micronesia],Ponape [= Pohnpei],at peak of Mount Troton, 600 m, syn.nov.</p><p>Hippeophyllum microphyllum S.C. Chen (2003) 180, f. 1 (non Oberonia microphylla (Blume) Lindl.). — Oberonia chenii Ormerod (2004) 378. — Type: van Royen &amp; Sleumer 7662 (holo K 000942975; iso L.1512780), Indonesia, Irian Jaya, Vogelkop Peninsula, Ilje River valley, path from Banfot to Sudjak, 840 m, syn. nov.</p><p>Notes — Oberonia rhizomatosa and O. repens were both described in 1911. Schlechter’s O. repens is precisely dated to October 11, 1911. The description of O. rhizomatosa was on page 2 of the volume published in 1911, so most likely in the first half of the year. Accordingly, Smith’s name is taken as the correct name for this species. The fact that Schlechter referred to O. rhizomatosa in his description of O. repens further supports the earlier publication date for O. rhizomatosa .</p><p>The illustrations in Smith (1911b) were given with fractional size indications, which were used as basis to scale the floral elements to same relative size. The 1 mm scale bar is derived from the absolute size indications in the Latin protologue, where the longer scale bar is based on the lip, while the shorter is based on the measurement for the sepal. The difference between the two scale bars is over 40 %. It is a further example of how imprecise measurements and scales are in descriptions of Oberonia . Accordingly, such measurements should be viewed with much caution, and minor size differences in the literature are most likely due to measurement errors.</p><p>Additionally, the shape of the lip differs between the drawing of the entire flower and in the illustration of the parts. For the entire flower, the lateral lobes are more narrow than the epichile lobes, while those relations are reversed in the isolated lip. The lateral lobes are shown without auricles in the entire flower, but with distinct auricles in the isolated lip.</p><p>Oberonia repens is a synonym of O. rhizomatosa . The shared characters include the uncommon creeping habit, narrow leaves and the overall shape of the lip. Schlechter (1911a: translated from original German) noted the similarity to O. rhizomatosa and indicated that ‘the labellum and petals in the here described species seem to be quite different’ without elaborating on any specifics.</p><p>The description of O. torana was based solely on Gjellerup 777, so should be given preferential standing; however, the repository of that gathering is uncertain as the specimen was not figured by Schuiteman &amp; De Vogel (2006). Smith (1915) wondered whether O. torana might be the same as O. repens, but did not compare his O. torana to his earlier O. rhizomatosa . The descriptions and flower drawings leave no doubt about Smith’s conclusion. The creeping rhizome, the moderately caulescent habit with long, thin, falcate leaves also seen in Gjellerup 912 specimens, are in prefect agreement with O. rhizomatosa .</p><p>Oberonia ponapensis shares the rather uncommon creeping habit, moderately caulescent growth with narrow leaves, and the flower with lobed petals and the lip with upward-turned, trapezoidal lateral lobes and two lobes of the epichile. The habit of the two growths mounted on the holotype sheet (Fig. 13h) show how smaller plants are little caulescent similar to Smith’s drawing of O. rhizomatosa, while larger plants develop a more caulescent habit. Notice the horizontal rhizome at the base of the larger two growths.</p><p>The epichile lobes were described by Tuyama (1940: 275) as ‘falcatos triangulos acutos producta’, which is different from the rounded lobes seen in the SEM images shown here. While Tuyama did not illustrate the flower, a specimen from Ponape identified as O. ponapensis and with identical habit could be examined (Fig. 13i). The imperfectly rehydrated flower shows lateral lobes wider than the epichile lobes, and the epichile lobes appear to be more curved with pointed tip similar to Tuyama’s description. Those differences, though, can easily be attributed to artefacts due to drying and rehydration of the flower, including the necessary unfolding of overlapping floral elements. The right petals with erose margin appears to have a rather narrow shape. The latter is due to the top margin being folded over, and is remarkably similar to J.J. Smith’s (1911b, see Fig. 13a) illustration of O. rhizomatosa . The white rectangular inclusions are calcium oxalate raphide bundles (Geiger unpubl. data).</p><p>Oberonia ponapensis was compared to O. brunnea Schltr., O. linearis Schltr., O. radicans Schltr. with similar lateral lobes but much more pointed lobes of the epichile, and most importantly without creeping rhizomes.</p><p>Hippeophyllum micranthum = O. chenii is another synonym of O. rhizomatosa . It shares the uncommon creeping habit, the serrated lateral petals and the quadrilobed lip with O. rhizomatosa . The flowers were described as dark red in colour, as in O. repens by Schlechter (1911a), and 0.8 mm wide. Chen (2003) did not compare his species to any others, and did not include a single reference in his description. A perusal of Schlechter’s (1911a) well-known work from the same region would have quickly identified his plant to at least one of the already described names.</p></div>	https://treatment.plazi.org/id/F541C527FF99FF99A73F9F289F5AFEF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
F541C527FF9FFF9AA4709C3B998AFDE3.text	F541C527FF9FFF9AA4709C3B998AFDE3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oberonia spathipetala J. J. Sm.	<div><p>Oberonia spathipetala J.J.Sm. — Fig. 15</p><p>Oberonia spathipetala J.J.Sm. (1908) 30. — Syntype: Branderhorst 191 (BO), [Indonesia, Irian Jaya,] S coast, N of the Kampoeng Gelieb .</p><p>Oberonia odontopetala Schltr.(1911a) 159;(1923) pl.61,f. 217. — Syntypes: Schlechter 19571 (B, lost; BO, L 0061767,AMES 34056/Harvard barcode 00102012), on trees in the forest around Dschischungari, Papua New Guinea, 800 m, syn. nov.</p><p>Oberonia pectinata Schltr. (1911a) 159; (1923) pl. 61, f. 218. — Syntype: Schlechter 18540 (B, lost),forests of the Bismarck mountains, Papua New Guinea, 1100 m, syn. nov.</p><p>Notes — Smith (1909) used ‘n. sp.’ in the treatment of O. spathipetala, obviously a lapsus calami, that cannot be interpreted as an introduction of a new, homonymous name by the same author.</p><p>Oberonia odontopetala and O. pectinata share the strongly toothed lateral petals, the T-shaped lip with expanded epichile, and have identical habits. The margin of the median sepal rang- es from entire (Fig. 15d – e) to apically toothed (Fig. 15f– g, i) with every intermediate state seen. Similarly, the degree of toothing of the lateral petals is variable between specimens based on additional 16 herbarium specimens examined in BM, CANB, E, K, NSW, SING, and WU. The two illustrations in Schlechter (1923) represent points along a continuous gradient.</p></div>	https://treatment.plazi.org/id/F541C527FF9FFF9AA4709C3B998AFDE3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Geiger, D. L.	Geiger, D. L. (2020): Studies in Oberonia 8 (Orchidaceae: Malaxideae). Additional 24 new synonyms, a corrected spelling, and other nomenclatural matters. Blumea 65 (3): 188-203, DOI: 10.3767/blumea.2020.65.03.02, URL: https://doi.org/10.3767/blumea.2020.65.03.02
