taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
CF49EC2A480257119CF9275091312ABE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313008	https://doi.org/10.3897/zse.101.137663.figure6	Figure 6. Haploniscus apaticus sp. nov. female paratype, SKB Hap 18 (A, C); male holotype SKB Hap 08 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm.	Figure 6. Haploniscus apaticus sp. nov. female paratype, SKB Hap 18 (A, C); male holotype SKB Hap 08 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
CF49EC2A480257119CF9275091312ABE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313009	https://doi.org/10.3897/zse.101.137663.figure7	Figure 7. Haploniscus apaticus sp. nov. male holotype, SKB Hap 08. A. Antenna II; B. Pereopod I; C. Pereopod II; D. Pereopod VI; E. Antenna I. Scale bar: 0.4 mm.	Figure 7. Haploniscus apaticus sp. nov. male holotype, SKB Hap 08. A. Antenna II; B. Pereopod I; C. Pereopod II; D. Pereopod VI; E. Antenna I. Scale bar: 0.4 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
CF49EC2A480257119CF9275091312ABE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313010	https://doi.org/10.3897/zse.101.137663.figure8	Figure 8. Haploniscus apaticus sp. nov. male holotype, SKB Hap 08. A. maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. Maxilla I; F. Maxillipeds, detail of distomedial margins of endites; G. Right mandible, detail of incisor and molar process; H. Left mandible, detail of incisor, lacinia mobilis, and molar process. Scale bars: 0.1 mm.	Figure 8. Haploniscus apaticus sp. nov. male holotype, SKB Hap 08. A. maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. Maxilla I; F. Maxillipeds, detail of distomedial margins of endites; G. Right mandible, detail of incisor and molar process; H. Left mandible, detail of incisor, lacinia mobilis, and molar process. Scale bars: 0.1 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
CF49EC2A480257119CF9275091312ABE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313011	https://doi.org/10.3897/zse.101.137663.figure9	Figure 9. Haploniscus apaticus sp. nov. male holotype, SKB Hap 08. A. Pleopod I; B. Pleopod II, ventral view; C. Pleopod II, dorsal view; D. Pleopod III; E. Pleopod IV; F. Pleopod V. Scale bar: 0.25 mm.	Figure 9. Haploniscus apaticus sp. nov. male holotype, SKB Hap 08. A. Pleopod I; B. Pleopod II, ventral view; C. Pleopod II, dorsal view; D. Pleopod III; E. Pleopod IV; F. Pleopod V. Scale bar: 0.25 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
CF49EC2A480257119CF9275091312ABE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313028	https://doi.org/10.3897/zse.101.137663.figure26	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
CF49EC2A480257119CF9275091312ABE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313029	https://doi.org/10.3897/zse.101.137663.figure27	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
BC04A7E2765F554B8A48FEB397DB12D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313003	https://doi.org/10.3897/zse.101.137663.figure1	Figure 1. Morphological plasticity of adult males within the Haploniscus belyaevi species complex from Knauber et al. (2022). Dorsal habitus CLSM scans of H. hades sp. nov. (A), H. belyaevi (B), and H. erebus sp. nov. (C). Scale bar: 1 mm.	Figure 1. Morphological plasticity of adult males within the Haploniscus belyaevi species complex from Knauber et al. (2022). Dorsal habitus CLSM scans of H. hades sp. nov. (A), H. belyaevi (B), and H. erebus sp. nov. (C). Scale bar: 1 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
BC04A7E2765F554B8A48FEB397DB12D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313004	https://doi.org/10.3897/zse.101.137663.figure2	Figure 2. Haploniscus belyaevi Birstein, 1963 female paraneotype SKB Hap 24 (A, C); male neotype, SKB Hap 46 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm.	Figure 2. Haploniscus belyaevi Birstein, 1963 female paraneotype SKB Hap 24 (A, C); male neotype, SKB Hap 46 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
BC04A7E2765F554B8A48FEB397DB12D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313005	https://doi.org/10.3897/zse.101.137663.figure3	Figure 3. Haploniscus belyaevi Birstein, 1963 male neotype, SKB Hap 46. A. Antenna II; B. Antenna I; C. Pereopod I; D. Pereopod II; E. Pereopod VI. Scale bars: 0.4 mm.	Figure 3. Haploniscus belyaevi Birstein, 1963 male neotype, SKB Hap 46. A. Antenna II; B. Antenna I; C. Pereopod I; D. Pereopod II; E. Pereopod VI. Scale bars: 0.4 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
BC04A7E2765F554B8A48FEB397DB12D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313006	https://doi.org/10.3897/zse.101.137663.figure4	Figure 4. Haploniscus belyaevi Birstein, 1963 male neotype, SKB Hap 46. A. Maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. Maxilla I; F. Maxillipeds, detail of distomedial margins of endites; G. Left mandible, detail of incisor, lacinia mobilis, and molar process; H. Right mandible, detail of incisor and molar process. Scale bars: 0.1 mm.	Figure 4. Haploniscus belyaevi Birstein, 1963 male neotype, SKB Hap 46. A. Maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. Maxilla I; F. Maxillipeds, detail of distomedial margins of endites; G. Left mandible, detail of incisor, lacinia mobilis, and molar process; H. Right mandible, detail of incisor and molar process. Scale bars: 0.1 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
BC04A7E2765F554B8A48FEB397DB12D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313007	https://doi.org/10.3897/zse.101.137663.figure5	Figure 5. Haploniscus belyaevi Birstein, 1963 male neotype, SKB Hap 46. A. Pleopod I; B. Pleopod II, ventral view; C. Pleopod II, dorsal view; D. Pleopod III; E. Pleopod IV; F. Pleopod V. Scale bar: 0.25 mm.	Figure 5. Haploniscus belyaevi Birstein, 1963 male neotype, SKB Hap 46. A. Pleopod I; B. Pleopod II, ventral view; C. Pleopod II, dorsal view; D. Pleopod III; E. Pleopod IV; F. Pleopod V. Scale bar: 0.25 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
BC04A7E2765F554B8A48FEB397DB12D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313028	https://doi.org/10.3897/zse.101.137663.figure26	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
BC04A7E2765F554B8A48FEB397DB12D1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313029	https://doi.org/10.3897/zse.101.137663.figure27	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
13D95A2E853659AC950D7E6D636BAF09.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313003	https://doi.org/10.3897/zse.101.137663.figure1	Figure 1. Morphological plasticity of adult males within the Haploniscus belyaevi species complex from Knauber et al. (2022). Dorsal habitus CLSM scans of H. hades sp. nov. (A), H. belyaevi (B), and H. erebus sp. nov. (C). Scale bar: 1 mm.	Figure 1. Morphological plasticity of adult males within the Haploniscus belyaevi species complex from Knauber et al. (2022). Dorsal habitus CLSM scans of H. hades sp. nov. (A), H. belyaevi (B), and H. erebus sp. nov. (C). Scale bar: 1 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
13D95A2E853659AC950D7E6D636BAF09.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313012	https://doi.org/10.3897/zse.101.137663.figure10	Figure 10. Haploniscus erebus sp. nov. female paratype, SKB Hap 50 (A, C); male holotype SKB Hap 54 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm.	Figure 10. Haploniscus erebus sp. nov. female paratype, SKB Hap 50 (A, C); male holotype SKB Hap 54 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
13D95A2E853659AC950D7E6D636BAF09.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313013	https://doi.org/10.3897/zse.101.137663.figure11	Figure 11. Haploniscus erebus sp. nov. male holotype, SKB Hap 54. A. Antenna II; B. Antenna I; C. Pereopod I; D. Pereopod II; E. Pereopod VI. Scale bars: 0.4 mm.	Figure 11. Haploniscus erebus sp. nov. male holotype, SKB Hap 54. A. Antenna II; B. Antenna I; C. Pereopod I; D. Pereopod II; E. Pereopod VI. Scale bars: 0.4 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
13D95A2E853659AC950D7E6D636BAF09.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313014	https://doi.org/10.3897/zse.101.137663.figure12	Figure 12. Haploniscus erebus sp. nov. male holotype, SKB Hap 54. A. Maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. Maxilla I; F. Maxilliped, right, detail of distomedial margin of endite; G. Maxilliped, left, detail of distomedial margin of endite; H. Right mandible, detail of incisor and molar process; I. Left mandible, detail of incisor, lacinia mobilis, and molar process. Scale bars: 0.1 mm.	Figure 12. Haploniscus erebus sp. nov. male holotype, SKB Hap 54. A. Maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. Maxilla I; F. Maxilliped, right, detail of distomedial margin of endite; G. Maxilliped, left, detail of distomedial margin of endite; H. Right mandible, detail of incisor and molar process; I. Left mandible, detail of incisor, lacinia mobilis, and molar process. Scale bars: 0.1 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
13D95A2E853659AC950D7E6D636BAF09.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313015	https://doi.org/10.3897/zse.101.137663.figure13	Figure 13. Haploniscus erebus sp. nov. male holotype, SKB Hap 54. A. Pleopod I; B. Pleopod II, ventral view; C. Pleopod II, dorsal view; D. Pleopods III; E. Pleopod IV; F. Pleopod V. Scale bar: 0.25 mm.	Figure 13. Haploniscus erebus sp. nov. male holotype, SKB Hap 54. A. Pleopod I; B. Pleopod II, ventral view; C. Pleopod II, dorsal view; D. Pleopods III; E. Pleopod IV; F. Pleopod V. Scale bar: 0.25 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
13D95A2E853659AC950D7E6D636BAF09.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313028	https://doi.org/10.3897/zse.101.137663.figure26	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
13D95A2E853659AC950D7E6D636BAF09.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313029	https://doi.org/10.3897/zse.101.137663.figure27	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
C0C4A9BB10B053E2B91F5B5522A6CC44.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313003	https://doi.org/10.3897/zse.101.137663.figure1	Figure 1. Morphological plasticity of adult males within the Haploniscus belyaevi species complex from Knauber et al. (2022). Dorsal habitus CLSM scans of H. hades sp. nov. (A), H. belyaevi (B), and H. erebus sp. nov. (C). Scale bar: 1 mm.	Figure 1. Morphological plasticity of adult males within the Haploniscus belyaevi species complex from Knauber et al. (2022). Dorsal habitus CLSM scans of H. hades sp. nov. (A), H. belyaevi (B), and H. erebus sp. nov. (C). Scale bar: 1 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
C0C4A9BB10B053E2B91F5B5522A6CC44.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313016	https://doi.org/10.3897/zse.101.137663.figure14	Figure 14. Haploniscus hades sp. nov. female paratype, KBII Hap 213 (A, C); male holotype KBII Hap 137 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm.	Figure 14. Haploniscus hades sp. nov. female paratype, KBII Hap 213 (A, C); male holotype KBII Hap 137 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
C0C4A9BB10B053E2B91F5B5522A6CC44.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313017	https://doi.org/10.3897/zse.101.137663.figure15	Figure 15. Haploniscus hades sp. nov. male holotype, KBII Hap 137. A. Antenna II; B. Antenna I; C. Pereopod I; D. Pereopod II; E. Pereopod VI. Scale bar: 0.4 mm.	Figure 15. Haploniscus hades sp. nov. male holotype, KBII Hap 137. A. Antenna II; B. Antenna I; C. Pereopod I; D. Pereopod II; E. Pereopod VI. Scale bar: 0.4 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
C0C4A9BB10B053E2B91F5B5522A6CC44.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313018	https://doi.org/10.3897/zse.101.137663.figure16	Figure 16. Haploniscus hades sp. nov. male holotype, KBII Hap 137. A. Maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. Maxilla I; F. Maxillipeds, detail of distomedial margins of endites; G. Right mandible, detail of incisor and molar process; H. Left mandible, detail of incisor, lacinia mobilis, and molar process. Scale bars: 0.1 mm.	Figure 16. Haploniscus hades sp. nov. male holotype, KBII Hap 137. A. Maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. Maxilla I; F. Maxillipeds, detail of distomedial margins of endites; G. Right mandible, detail of incisor and molar process; H. Left mandible, detail of incisor, lacinia mobilis, and molar process. Scale bars: 0.1 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
C0C4A9BB10B053E2B91F5B5522A6CC44.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313019	https://doi.org/10.3897/zse.101.137663.figure17	Figure 17. Haploniscus hades sp. nov. male holotype, KBII Hap 137. A. Pleopod I; B. Pleopod II, dorsal view; C. Pleopod II, ventral view; Pleopod III; D. Pleopod IV; E. Pleopod V. Scale bar: 0.25 mm.	Figure 17. Haploniscus hades sp. nov. male holotype, KBII Hap 137. A. Pleopod I; B. Pleopod II, dorsal view; C. Pleopod II, ventral view; Pleopod III; D. Pleopod IV; E. Pleopod V. Scale bar: 0.25 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
C0C4A9BB10B053E2B91F5B5522A6CC44.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313028	https://doi.org/10.3897/zse.101.137663.figure26	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
C0C4A9BB10B053E2B91F5B5522A6CC44.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313029	https://doi.org/10.3897/zse.101.137663.figure27	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
A19368E9BC2F5FA4BDDF0F0F45D53813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313020	https://doi.org/10.3897/zse.101.137663.figure18	Figure 18. Haploniscus kerberos sp. nov. female paratype, KBII Hap 123 (A, C); male holotype KBII Hap 136 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm.	Figure 18. Haploniscus kerberos sp. nov. female paratype, KBII Hap 123 (A, C); male holotype KBII Hap 136 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
A19368E9BC2F5FA4BDDF0F0F45D53813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313021	https://doi.org/10.3897/zse.101.137663.figure19	Figure 19. Haploniscus kerberos sp. nov. male holotype, KBII Hap 136. A. Antenna II; B. Antenna I; C. Pereopod VI; D. Pereopod I. Scale bar: 0.4 mm.	Figure 19. Haploniscus kerberos sp. nov. male holotype, KBII Hap 136. A. Antenna II; B. Antenna I; C. Pereopod VI; D. Pereopod I. Scale bar: 0.4 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
A19368E9BC2F5FA4BDDF0F0F45D53813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313022	https://doi.org/10.3897/zse.101.137663.figure20	Figure 20. Haploniscus kerberos sp. nov. male holotype, KBII Hap 136. A. Maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. Maxilla I; F. Left maxilliped, detail of distomedial margin of endite; G. Right maxilliped, detail of distomedial margin of endite; H. Right mandible, detail of incisor and molar process; I. Left mandible, detail of incisor, lacinia mobilis, and molar process. Scale bars: 0.1 mm.	Figure 20. Haploniscus kerberos sp. nov. male holotype, KBII Hap 136. A. Maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. Maxilla I; F. Left maxilliped, detail of distomedial margin of endite; G. Right maxilliped, detail of distomedial margin of endite; H. Right mandible, detail of incisor and molar process; I. Left mandible, detail of incisor, lacinia mobilis, and molar process. Scale bars: 0.1 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
A19368E9BC2F5FA4BDDF0F0F45D53813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313023	https://doi.org/10.3897/zse.101.137663.figure21	Figure 21. Haploniscus kerberos sp. nov. male holotype, KBII Hap 136. A. Pleopod II, dorsal view; B. Pleopod II, ventral view; C. Pleopod III; D. Pleopod IV; E. Pleopod V. Scale bar: 0.25 mm.	Figure 21. Haploniscus kerberos sp. nov. male holotype, KBII Hap 136. A. Pleopod II, dorsal view; B. Pleopod II, ventral view; C. Pleopod III; D. Pleopod IV; E. Pleopod V. Scale bar: 0.25 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
A19368E9BC2F5FA4BDDF0F0F45D53813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313028	https://doi.org/10.3897/zse.101.137663.figure26	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
A19368E9BC2F5FA4BDDF0F0F45D53813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313029	https://doi.org/10.3897/zse.101.137663.figure27	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
F15FB4C76CF051AA9A301C37681E021D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313024	https://doi.org/10.3897/zse.101.137663.figure22	Figure 22. Haploniscus nyx sp. nov. female paratype, SKB Hap 21 (A, C); male holotype SKB Hap 34 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm. Due to pronounced body curvature, the habitus drawing of the female paratype was prepared by two separate drawings, stitching them together along the illustrated line.	Figure 22. Haploniscus nyx sp. nov. female paratype, SKB Hap 21 (A, C); male holotype SKB Hap 34 (B, D). A, B. Habitus, dorsal view; C, D. Head, lateral view. Scale bars: 0.5 mm. Due to pronounced body curvature, the habitus drawing of the female paratype was prepared by two separate drawings, stitching them together along the illustrated line.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
F15FB4C76CF051AA9A301C37681E021D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313025	https://doi.org/10.3897/zse.101.137663.figure23	Figure 23. Haploniscus nyx sp. nov. male holotype, SKB Hap 34. A. Antenna II; B. Antenna I; C. Pereopod I; D. Pereopod II; E. Pereopod VI. Scale bars: 0.4 mm.	Figure 23. Haploniscus nyx sp. nov. male holotype, SKB Hap 34. A. Antenna II; B. Antenna I; C. Pereopod I; D. Pereopod II; E. Pereopod VI. Scale bars: 0.4 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
F15FB4C76CF051AA9A301C37681E021D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313026	https://doi.org/10.3897/zse.101.137663.figure24	Figure 24. Haploniscus nyx sp. nov. male holotype, SKB Hap 34. A. Maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. maxilla I; F. Maxillipeds, detail of distomedial margins of endites; G. Right mandible, detail of incisor and molar process; H. Left mandible, detail of incisor, lacinia mobilis, and molar process. Scale bars: 0.1 mm.	Figure 24. Haploniscus nyx sp. nov. male holotype, SKB Hap 34. A. Maxillipeds; B. Right mandible; C. Left mandible; D. Maxilla II; E. maxilla I; F. Maxillipeds, detail of distomedial margins of endites; G. Right mandible, detail of incisor and molar process; H. Left mandible, detail of incisor, lacinia mobilis, and molar process. Scale bars: 0.1 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
F15FB4C76CF051AA9A301C37681E021D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313027	https://doi.org/10.3897/zse.101.137663.figure25	Figure 25. Haploniscus nyx sp. nov. male holotype, SKB Hap 34. A. Pleopod I; B. Pleopod II, ventral view; C. Pleopod II, dorsal view; D. Pleopods III; E. Pleopod IV; F. Pleopod V. Scale bar: 0.25 mm.	Figure 25. Haploniscus nyx sp. nov. male holotype, SKB Hap 34. A. Pleopod I; B. Pleopod II, ventral view; C. Pleopod II, dorsal view; D. Pleopods III; E. Pleopod IV; F. Pleopod V. Scale bar: 0.25 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
F15FB4C76CF051AA9A301C37681E021D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313028	https://doi.org/10.3897/zse.101.137663.figure26	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	Figure 26. Comparison of pleotelson shapes amongst adult male and female members of the Haploniscus belyaevi species complex (CLSM). H. erebus sp. nov. female (A) and male (B) from Knauber et al. (2022); H. kerberos sp. nov. female (C) and male (D); H. hades sp. nov. female (E) and male (F); H. belyaevi female (G) and male (H); H. apaticus sp. nov. female (I) and male (J); H. nyx sp. nov. female (K) and male (L). Scale bars: 0.5 mm.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
F15FB4C76CF051AA9A301C37681E021D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1313029	https://doi.org/10.3897/zse.101.137663.figure27	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	Figure 27. Distribution of haploniscid species of the belyaevi - complex in the greater Kuril-Kamchatka Trench and Sea of Okhotsk area of the Northwest Pacific. Stars indicate each species type locality.	2025-04-22	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben		Zenodo	biologists	Knauber, Henry;Schell, Tilman;Brandt, Angelika;Riehl, Torben			
