taxonID	type	description	language	source
496E3FA0FAC9534CA2652A80715B7F91.taxon	description	Figs 1, 3, 4, 5, 6, Table 1 New English name: Klapanunggal Blind Cave Barb New Indonesian name: Wader Gua Buta Klapanunggal	en	Wibowo, Kunto, Willyanto, M. Iqbal, Dharmayanthi, Anik Budhi, Rahmadi, Cahyo, Lumbantobing, Daniel Natanael (2025): Barbodes klapanunggalensis, a new species of blind subterranean fish (Cypriniformes, Cyprinidae) from Klapanunggal karst area, West Java, Indonesia, with notes on its conservation. ZooKeys 1229: 43-59, DOI: 10.3897/zookeys.1229.135950
496E3FA0FAC9534CA2652A80715B7F91.taxon	diagnosis	Diagnosis. A species of Barbodes distinguished from all its congeners by the absence of eyes, as the eye is vestigially replaced by an orbital concavity being fully closed by an epidermal layer, while lacking the orbital rim. The species is also uniquely diagnosed by having relatively long pectoral and pelvic fins, with their adpressed tips each extending past the vertical through the insertion or origin of the next fin posterior to the tip, as to further pass posteriorly about two scales in anteroposterior; and by the presence of a relatively short and rounded axillary pelvic-fin scale, with tip not reaching posterior edge of pelvic-fin base. It is further distinguished from other Barbodes species in having by the following combination of characters: head length 32.9 – 35.3 % of SL; pre-pectoral fin length 32.6 – 33.6 % of SL; pre-pelvic fin length 54.0 – 59.6 % of SL; pectoral fin length 26.0 – 31.4 % of SL; pelvic fin length 21.5 – 24.4 % of SL; anal-fin base length 9.7 – 11.8 % of SL; caudal peduncle depth 13.2 – 18.2 % of SL; body without pigmentation (black dots, bars, stripes, blotches, and triangular markings all absent from lateral surface); all fins with translucent interradial membrane and light cream to brownish rays.	en	Wibowo, Kunto, Willyanto, M. Iqbal, Dharmayanthi, Anik Budhi, Rahmadi, Cahyo, Lumbantobing, Daniel Natanael (2025): Barbodes klapanunggalensis, a new species of blind subterranean fish (Cypriniformes, Cyprinidae) from Klapanunggal karst area, West Java, Indonesia, with notes on its conservation. ZooKeys 1229: 43-59, DOI: 10.3897/zookeys.1229.135950
496E3FA0FAC9534CA2652A80715B7F91.taxon	description	Description. Data for holotype presented first, followed by paratype data in parentheses (if different). Selected meristic and morphometric data given as percentages of SL in Table 1. General appearance shown in Figs 3 A, B, 4 A, B. Body deep, laterally moderately compressed along anterior portion, progressively more compressed posteriorly. Greatest body depth at vertical through dorsal-fin origin. Dorsal profile of head posterodorsally slanted overall, with slight concavity along supraorbital profile. Limit between head and trunk marked by slight convexity of anterior predorsal profile. Dorsal profile of anterior body slightly arched anteriorly, nearly flat posteriorly (entirely posterodorsally slanted in paratype). Snout slightly rounded. Mouth subterminal, marked by two pairs of maxillary barbels, anterior pair shorter than posterior pair. Eye absent, ocular vestige marked by orbital concavity completely covered by skin, orbital rim absent. A short flap on upper posterior edge of low membranous tube associated with anterior nostril. Cycloid scales covering body, not extending onto fin rays or membranes, except basally on dorsal, anal, and caudal fins. Pelvic-fin axillary scale short, with rounded tip (slightly projecting on right side of holotype), not reaching vertical through posterior edge of pelvic-fin base (Fig. 5 A, B). Dorsal fin with distal profile slightly concave, 4 (3) unbranched spinous rays, 8 ½ soft rays. First unbranched dorsal-fin ray very small; second ray about one-quarter length of third; last unbranched ray wide at base, progressively narrowing distally, with 10 (19) serrae along posterior edge of stiffened upper half. All soft dorsal-fin rays branched, first branched ray longest, slightly longer than fourth (third) unbranched ray. Pectoral fin slightly rounded, with 15 (14) rays; anteriormost ray unbranched, its adpressed tip extending past vertical through pelvic fin insertion. Pelvic fin with 10 (9) rays; anteriormost ray unbranched, its adpressed tip just reaching (extending past) anal fin origin; pelvic fin insertion about level with dorsal fin origin. Anal fin with 3 unbranched and 5 ½ branched rays; distal profile slightly concave. Caudal fin with 10 + 9 principal rays, uppermost and lowermost rays unbranched, symmetrically forked, tip of lobes pointed (rounded); upper caudal-fin lobe length 32.0 (34.8) % of SL; median caudal fin length 14.9 (18.7) % of SL; lower caudal-fin lobe length 31.8 (34.9) % of SL. Lateral-line scales complete 24 + 1 (22 + 1). Pre-dorsal scales 10. Scale rows above lateral line 4 ½, below lateral line at pelvic fin base 3 ½, below lateral line at anal fin base 3 ½. Scale rows in transverse line on caudal peduncle ½ 2 / 1 / 2 ½. Total gill rakers ca. 12, rakers short, length of longest raker on first gill arch about half length of adjacent gill filaments. Vertebrae 16 + 13 (14 + 14). Coloration when fresh (Figs 3 A, 4 A). Head and body silvery-white, slightly cream to brownish dorsally, becoming lighter ventrally; indistinct longitudinal pinkish band on mid-lateral surface of body from below dorsal fin to caudal-fin base. All fins with semitranslucent interradial membrane and light cream to brownish rays. Coloration in alcohol (Figs 3 B, 4 B). Head and body pale yellowish brown; body slightly darker dorsally, becoming lighter ventrally. All fins semi-translucent.	en	Wibowo, Kunto, Willyanto, M. Iqbal, Dharmayanthi, Anik Budhi, Rahmadi, Cahyo, Lumbantobing, Daniel Natanael (2025): Barbodes klapanunggalensis, a new species of blind subterranean fish (Cypriniformes, Cyprinidae) from Klapanunggal karst area, West Java, Indonesia, with notes on its conservation. ZooKeys 1229: 43-59, DOI: 10.3897/zookeys.1229.135950
496E3FA0FAC9534CA2652A80715B7F91.taxon	distribution	Distribution. Currently known from a subterranean creek in Cisodong 1 Cave, in the karst area at Klapanunggal, Bogor, West Java, Indonesia (Fig. 2). The creek drains into Cileungsi River, a tributary of the Bekasi River Drainage that is eventually emptied into the Jakarta Bay.	en	Wibowo, Kunto, Willyanto, M. Iqbal, Dharmayanthi, Anik Budhi, Rahmadi, Cahyo, Lumbantobing, Daniel Natanael (2025): Barbodes klapanunggalensis, a new species of blind subterranean fish (Cypriniformes, Cyprinidae) from Klapanunggal karst area, West Java, Indonesia, with notes on its conservation. ZooKeys 1229: 43-59, DOI: 10.3897/zookeys.1229.135950
496E3FA0FAC9534CA2652A80715B7F91.taxon	etymology	Etymology. The specific epithet klapanunggalensis is derived from the type locality, the Klapanunggal karst area, which includes the Cisodong 1 Cave, Nambo Village. The name reflects the unique habitat and geological significance of the Klapanunggal karst area, where the species is likely endemic.	en	Wibowo, Kunto, Willyanto, M. Iqbal, Dharmayanthi, Anik Budhi, Rahmadi, Cahyo, Lumbantobing, Daniel Natanael (2025): Barbodes klapanunggalensis, a new species of blind subterranean fish (Cypriniformes, Cyprinidae) from Klapanunggal karst area, West Java, Indonesia, with notes on its conservation. ZooKeys 1229: 43-59, DOI: 10.3897/zookeys.1229.135950
